A Paleozoological Perspective on White-Tailed Deer (<Emphasis Type="Italic">Odocoileus virginianus texana</Emphasis>) Population Density and Body Size in Central Texas

Archaeological and paleontological datasets are used in conservation to add time-depth to ecology. In central Texas, several top carnivores including prehistoric Native American hunters have been extirpated or have had their historic ranges restricted, which has resulted in pest-level white-tailed deer (Odocoileus virginianus texana) populations in some areas. Differences in body size of deer between prehistory and modernity are expected, given that a lack of predation likely has increased intraspecific competition for forage among deer, resulting in smaller body size today. In fact, modern deer from settings without harvest pressure are significantly smaller than those from harvested areas and from prehistoric deer. From a natural history perspective, this research highlights potential evolutionary causes and effects of top-predator removal on deer populations and related components of biological communities in central Texas.     

Seasonal food use by white-tailed deer at Valley Forge National Historical Park,Pennsylvania, USA

Food habits of white-tailed deer (Odocoileus virginianus) were examined from January to November 1984 via fecal-pellet analysis at Valley Forge National Historical Park (VFNHP), which represents an island habitat for deer surrounded by extensive urbanization, in southeastern Pennsylvania. In addition, use of fields by deer was compared to food habits. Herbaceous vegetation (forbs, leaves of woody plants, and conifer needles) was the predominant food type in all seasons except fall. Acorns and graminoids (grasses and sedges) were important food resources in fall and spring, respectively. Use of woody browse (twigs) was similar among seasons. Field use was relatively high during fall, winter without snow cover (<20 cm), and spring when food resources in fields were readily available. In contrast, use of fields was lowest in summer when preferred woodland foods were available and in winter with snow cover when food in fields was not readily accessible. Patterns of food-type use by deer at VFNHP indicate the year-round importance of nonwoody foods and field habitats to deer populations on public lands such as national parks in the northeastern United States.     

Changes in interacting species with disturbance

Human-influenced changes in the diversity and abundance of native wildlife in a southern boreal forest area, which became a national park in 1975, are used to develop working hypotheses for predicting and subsequently measuring the effects of disturbance or restoration programs on groups of interacting species. Changes from presettlement conditions began with early 1900 hunting, which eliminated woodland caribou (Rangifer tarandus) and elk (Cervus elaphus), and reduced moose (Alces alces) to the low numbers which still persist. Increases in white-tailed deer (Odocoileus virginianus), as these other cervid species became less abundant or absent, provided enough alternative food to sustain the system's carnivores until plant succession on previously burned or logged areas also caused deer to decline. With increased competition for reduced food, carnivore species also became less abundant or absent and overexploited some prey populations. The abilities of interacting species to maintain dynamically stable populations or persist varied with their different capacities to compensate for increased exploitation or competition. These relationships suggested a possible solution to the problem of predicting the stability of populations in disturbed systems. For the 1976–1985 period, a hypothesis that the increased protection of wildlife from exploitation in a national park would restore a more diverse, abundant, and productive fauna had to be rejected.     

Using Satellite Imagery to Assess Large-Scale Habitat Characteristics of Adirondack Park, New York, USA

Satellite imagery is a useful tool for large-scale habitat analysis; however, its limitations need to be tested. We tested these limitations by varying the methods of a habitat evaluation for white-tailed deer (Odocoileus virginianus) in the Adirondack Park, New York, USA, utilizing harvest data to create and validate the assessment models. We used two classified images, one with a large minimum mapping unit but high accuracy and one with no minimum mapping unit but slightly lower accuracy, to test the sensitivity of the evaluation to these differences. We tested the utility of two methods of assessment, habitat suitability index modeling, and pattern recognition modeling. We varied the scale at which the models were applied by using five separate sizes of analysis windows. Results showed that the presence of a large minimum mapping unit eliminates important details of the habitat. Window size is relatively unimportant if the data are averaged to a large resolution (i.e., township), but if the data are used at the smaller resolution, then the window size is an important consideration. In the Adirondacks, the proportion of hardwood and softwood in an area is most important to the spatial dynamics of deer populations. The low occurrence of open area in all parts of the park either limits the effect of this cover type on the population or limits our ability to detect the effect. The arrangement and interspersion of cover types were not significant to deer populations.     

The Effects of Population Density on Juvenile Growth Rate in White-Tailed Deer

Animal body size is driven by habitat quality, food availability, and nutrition. Adult size can relate to birth weight, to length of the ontogenetic growth period, and/or to the rate of growth. Data requirements are high for studying these growth mechanisms, but large datasets exist for some game species. In North America, large harvest datasets exist for white-tailed deer (Odocoileus virginianus), but such data are collected under a variety of conditions and are generally dismissed for ecological research beyond local population and habitat management. We contend that such data are useful for studying the ecology of white-tailed deer growth and body size when analyzed at ordinal scale. In this paper, we test the response of growth rate to food availability by fitting a logarithmic equation that estimates growth rate only to harvest data from Fort Hood, Texas, and track changes in growth rate over time. Results of this ordinal scale model are compared to previously published models that include additional parameters, such as birth weight and adult weight. It is shown that body size responds to food availability by variation in growth rate. Models that estimate multiple parameters may not work with harvest data because they are prone to error, which renders estimates from complex models too variable to detect interannual changes in growth rate that this ordinal scale model captures. This model can be applied to harvest data, from which inferences about factors that influence animal growth and body size (e.g., habitat quality and nutritional availability) can be drawn.     

Diversity, Seasonality, and Context of Mammalian Roadkills in the Southern Great Plains

Thousands of mammals are killed annually from vehicle collisions, making the issue an important one for conservation biologists and environmental managers. We recorded all readily identifiable kills on or immediately adjacent to roads in the southern Great Plains from March 2004–March 2007. We also recorded distance traveled, whether a road was paved or divided, the number of lanes, and prevailing habitat. Surveys were opportunistic and were conducted by car during conditions of good visibility. Over our 239 surveys and >16,500 km traveled, we recorded 1412 roadkills from 18 different mammal species (size ranged from Sciurus squirrels to the white-tailed deer, Odocolieus virginianus). The overall kill rate was 8.50 / 100 km. Four species were prone to collisions: the Virginia opossum (Didelphis virginiana), nine-banded armadillo (Dasypus novemcinctus), striped skunk (Mephitis mephitis), and northern raccoon (Procyon lotor). Together they accounted for approximately 85% (1198) of all roadkills. Mortality rate differed significantly between 2- and 4-lane roads (8.39 versus 7.79 / 100 km). Kill rates were significantly higher on paved versus unpaved roads (8.60 versus 3.65 / 100 km), but did not depend on whether a road was divided. Roadkills were higher in spring than in fall (1.5×), winter (1.4×), or summer (1.3×). The spring peak (in kills / 100 km) was driven chiefly by the armadillo (2.76 in spring/summer versus 0.73 in autumn/winter) and opossum (2.65 versus 1.47). By contrast, seasonality was dampened by a late winter/early spring peak in skunk mortalities, for which 41% occurred in the 6-week period of mid-February through March. The raccoon did not exhibit a strong seasonal pattern. Our data are consistent with dispersal patterns of these species. Our results underscore the high rate of highway mortality in the southern plains, as well as differences in seasonality and road type that contribute to mortality. Conservation and management efforts should focus on creating underpasses or using other means to reduce roadkill rates.     

The Impact of Sika Deer on Vegetation in Japan: Setting Management Priorities on a National Scale

Irreversible shifts in ecosystems caused by large herbivores are becoming widespread around the world. We analyzed data derived from the 2009–2010 Sika Deer Impact Survey, which assessed the geographical distribution of deer impacts on vegetation through a questionnaire, on a scale of 5-km grid-cells. Our aim was to identify areas facing irreversible ecosystem shifts caused by deer overpopulation and in need of management prioritization. Our results demonstrated that the areas with heavy impacts on vegetation were widely distributed across Japan from north to south and from the coastal to the alpine areas. Grid-cells with heavy impacts are especially expanding in the southwestern part of the Pacific side of Japan. The intensity of deer impacts was explained by four factors: (1) the number of 5-km grid-cells with sika deer in neighboring 5 km-grid-cells in 1978 and 2003, (2) the year sika deer were first recorded in a grid-cell, (3) the number of months in which maximum snow depth exceeded 50 cm, and (4) the proportion of urban areas in a particular grid-cell. Based on our model, areas with long-persistent deer populations, short snow periods, and fewer urban areas were predicted to be the most vulnerable to deer impact. Although many areas matching these criteria already have heavy deer impact, there are some areas that remain only slightly impacted. These areas may need to be designated as having high management priority because of the possibility of a rapid intensification of deer impact.     

Agricultural area impacts within a natural area: Cades cove,a case history

Agricultural management in Cades Cove, an historic district in Great Smoky Mountains National Park, has affected natural resources both within the district and in the adjoining natural areas. Aquatic impacts of haying and cattle grazing included increases in water temperatures, turbidity, nutrient loading, and bacterial counts and decreases in benthic macroinvertebrate density and fish biomass. Wildlife populations, including groundhogs, wild turkeys, and white-tailed deer, have increased in the open fields and around the periphery of the historic district. Intensive deer foraging has removed deciduous seedlings and saplings from woodlots, lowering species diversity and favoring coniferous reproduction. Cades Cove has limestone habitats unique in the park, and both deer browse and cattle grazing may have disturbed populations of rare plant species. Effects on water quality are detectable at a campground 15 stream km from the agricultural area, and the effects of deer foraging extend about 1 km beyond the open fields.Since historic landscape preservation is presently a goal of the park, managing for open vistas in Cades Cove will require some sort of continuing disturbance. Conversion of cattle pastures to hayfields would reduce aquatic impacts but the deer herd might increase as a result of reduced competition for forage. Retarding old field succession would increase populations of native plant species dependent on sunlight, but would require government-funded mowing. Other options are discussed. Completely eliminating the effects of the historic district on adjoining areas may be impossible, at least under present economic constraints.     

Introduced Species and Management of a <Emphasis Type="Italic">Nothofagus/Austrocedrus</Emphasis> Forest

Isla Victoria (Nahuel Huapi National Park, Argentina), a large island dominated by native Nothofagus and Austrocedrus forest, has old plantations of many introduced tree species, some of which are famed invaders of native ecosystems elsewhere. There are also large populations of introduced deer and shrubs that may interact in a complex way with the introduced trees, as well as a recently arrived population of wild boar. Long-standing concern that the introduced trees will invade and transform native forest may be unwarranted, as there is little evidence of progressive invasion, even close to the plantations, despite over 50 years of opportunity. Introduced and native shrubs allow scattered introduced trees to achieve substantial size in abandoned pastures, but in almost all areas neither the trees nor the shrubs appear to be spreading beyond these sites. These shrub communities may be stable rather than successional, but the technology for restoring them to native forest is uncertain and probably currently impractical. Any attempt to remove the exotic tree seedlings and saplings from native forest would probably create the very conditions that would favor colonization by exotic plants rather than native trees, while simply clear-cutting the plantations would be unlikely to lead to regeneration of Nothofagus or Austrocedrus. The key to maintaining native forest is preventing catastrophic fire, as several introduced trees and shrubs would be favored over native dominant trees in recolonization. Deer undoubtedly interact with both native and introduced trees and shrubs, but their net effect on native forest is not yet clear, and specific management of deer beyond the current hunting by staff is unwarranted, at least if preventing tree invasion is the goal. The steep terrain and shallow soil make the recently arrived boar a grave threat to the native forest. Eradication is probably feasible and should be attempted quickly.     

Identification of Swamp Deer (<Emphasis Type="Italic">Cervus duvauceli duvauceli</Emphasis> Cuvier) Potential Habitat in Jhilmil Jheel Conservation Reserve,Uttarakhand, India Using Multi-Criteria Analysis

The present study aims to identify the potential habitat for swamp deer (Cervus duvauceli duvauceli Cuvier) in Jhilmil Jheel Conservation Reserve in the Uttarakhand province of India using multi-criteria analysis. The study area represents one of the last remnant habitats of the flagship species, the swamp deer in Uttarakhand, which is considered as vulnerable. The study showed that only 6.08% of the study area (225 km²) was highly suitable to suitable for the swamp deer. An area of 135.52 km² (60.23%) turned out to be moderately suitable. Within the officially designated Conservation Reserve (area 37.84 km²), 10.91% (4.13 km²) area was found highly suitable to suitable, while 74.19% (28.07 km²) happens to be moderately suitable. Only 14 km² area, which was found as suitable habitat for swamp deer falls short of the space required by a population of 134 animals. The problem could be mitigated if the agricultural land (2.47 km²) adjacent to the Jhilmil Jheel is brought under the Reserve management. This would provide additional area to meet the fodder requirement. The study brings out a particularly grim situation with limited options for conservation and management of the swamp deer in the Indo-Gangetic plains. It also emphasizes the role of geospatial techniques in quick appraisal of habitat attributes and identification of potential sites for protected areas.     

Model-Based Assessment of Aspen Responses to Elk Herbivory in Rocky Mountain National Park,USA

In Rocky Mountain National Park (RMNP), aspen (Populus tremuloides Michx.) has been observed to be declining on elk (Cervus elaphus nelsoni) winter range for many decades. To support elk management decisions, the SAVANNA ecosystem model was adapted to explore interactions between elk herbivory and aspen dynamics. The simulated probability of successful vegetative regeneration for senescent aspen stands declines sharply when elk densities reach levels of 3–5 elk/km², depending on model assumptions for the seasonal duration of elk foraging activities. For aspen stands with a substantial component of younger trees, the simulated regeneration probability declines more continuously with increasing elk density, dropping below 50% from densities at 8–14 elk/km².At the landscape scale, simulated aspen regeneration probability under a scenario of extensive seasonal use was little affected by elk population level, when this level was above 300–600 elk (25%–50% current population) over the ca. 107 km² winter range. This was because elk distribution was highly aggregated, so that a high density of elk occupied certain areas, even at low population levels overall. At approximately current elk population levels (1000–1200 elk), only 35%–45% of senescent aspen stands are simulated as having at least a 90% probability of regeneration, nearly all of them located on the periphery of the winter range. Successful management for aspen persistence on core winter range will likely require some combination of elk population reduction, management of elk distribution, and fencing to protect aspen suckers from elk browsing.     

Environmental and Strategic Uncertainty in Common Property Management: The Case of Scottish Red Deer

The range of red deer populations in the Scottish Highlands can cover several different landholdings (estates), many of which derive their income primarily from the private hunting (stalking) of stags. The deer belong to nobody and their seasonal movement does not respect the boundaries of individual estates, but a 'rule of capture' does apply as, once shot, the deer become the property of the estate owner. This paper argues that deer populations would best be managed as a common property resource. Indeed, for this reason, the Deer Commission for Scotland has encouraged owners to form Deer Management Groups. These groups are now able to utilize a computer program that models the relationship between grazing quality and population dynamics. However, drawing upon examples from the Western Highlands, the paper finds that common property management must overcome incentives to behave strategically that arise from the characteristics of flow and storage of the resource. The problem is aggravated by the degradation of the habitat that has occurred over centuries. This often means that the resource is below its optimum in terms of stag numbers and quality. Scientific models are an important contribution, but only reduce the environmental uncertainty. Their full potential can only be realized if all landowners are obliged to actively engage in common property management.     

Decision-making for fire use among farmers in savannas: an exploratory study in the Distrito Federal, central Brazil

Today, fire in the Braziliancerrado(savanna) is predominantly caused by farmers. They use fire during the dry season to promote fresh grass regrowth in pastures for their cattle and to clear areas of planting ready for cultivation in the wet season. This study researched fire use by farmers in thecerradoof the Distrito Federal, central Brazil, in order to investigate what influences their fire management decision-making. The results show that the two main factors limiting the choice of feasible options for farmers to meet their objectives are income and time. However, perceptions also play a major role in the decision-making process, and for many farmers this alone determines whether fire is used. Various fire practices involved with timing, frequency and size, as well as precautions taken before and during the burn, and the factors influencing these decisions are presented. This study shows that it is vital to understand factors determining fire use if effective fire management plans and policies are to be developed.1998 Academic Press     

The Impacts of Human Visitation on Mussel Bed Communities Along the California Coast: Are Regulatory Marine Reserves Effective in Protecting These Communities?

Rocky intertidal habitats frequently are used by humans for recreational, educational, and subsistence-harvesting purposes, with intertidal populations damaged by visitation activities such as extraction, trampling, and handling. California Marine Managed Areas, particularly regulatory marine reserves (MRs), were established to provide legal protection and enhancement of coastal resources and include prohibitions on harvesting intertidal populations. However, the effectiveness of MRs is unclear as enforcement of no-take laws is weak and no regulations protect intertidal species from other detrimental visitor impacts such as trampling. The goal of this study was two-fold: (1) to determine impacts from human visitation on California mussel populations (Mytilus californianus) and mussel bed community diversity; and (2) to investigate the effectiveness of regulatory MRs in reducing visitor impacts on these populations. Surveys of mussel populations and bed-associated diversity were compared: (1) at sites subjected to either high or low levels of human use, and (2) at sites either unprotected or with regulatory protection banning collecting. At sites subjected to higher levels of human visitation, mussel populations were significantly lower than low-use sites. Comparisons of mussel populations inside and outside of regulatory MRs revealed no consistent pattern suggesting that California no-take regulatory reserves may have limited effectiveness in protecting mussel communities. In areas where many people visit intertidal habitats for purposes other than collecting, many organisms will be affected by trampling, turning of rocks, and handling. In these cases, effective protection of rocky intertidal communities requires an approach that goes beyond the singular focus on collecting to reduce the full suite of impacts.     

Northern New England's threatened soft-shell clam populations

Commercial exhaustion of soft-shell clam,Mya arenaria, had become the fate of many potentially productive clamming areas along the coasts of Massachusetts, New Hampshire, and southern Maine. Investigations of a particularly severe case of soft-shell clam stock depletion in Hampton-Seabrook estuary, New Hampshire, indicated that by 1976 human diggers had removed approximately 87% of the adult clam resource that had existed in 1971. Meanwhile, circumstantial evidence strongly implicated the green crab,Carcinus maenas, as responsible for successive stock recruitment failures, these crabs having consumed most of the young seed clams before they could grow to harvestable size. In 1976, this apparent population imbalance may have been at least partly redressed by a prodigious spatfall, resulting in a massive and widespread reseeding of many flats along the northern New England coast. Spat, or seed clam, densities of up to 1,700 individuals per square foot, over areas of several acres, have been observed. Those clamming areas where aggressive predator control programs had been instituted exhibited the most favorable response in terms of seed clam survival and growth. Evidence also indicated that crowding in thickly seeded portions of the clam flats inhibited shell growth.     

Wildlife Contact Rates at Artificial Feeding Sites in Texas

Given the popularity of feeding white-tailed deer (Odocoileus virginianus) in Texas and the increasing amount of corn that is distributed, more information is needed on the impacts of this activity on non-target wildlife. Our objectives were to report visitation, intra- and interspecific contact, and contact rates of wildlife at artificial feeding sites in Texas. Our study was conducted at three sites in Kleberg and Nueces counties, Texas. We trapped animals from February to April and August to September, 2009 and marked animals with passive integrated transponder (PIT) tags. At each site and season, we placed one feeder system containing a PIT tag reader within 600 m of trap locations. Readers detected PIT tags from a distance of 25 cm. We determined a contact event to occur when two different PIT tags were detected by feeder systems within 5 s. We recorded 62,719 passes by raccoons (Procyon lotor), 103,512 passes by collared peccaries (Pecari tajacu), 2,923 passes by feral swine (Sus scrofa), 1,336 passes by fox squirrels (Sciurus niger), and no passes by opossums (Didelphis virginiana) at feeder systems. For site–season combinations in which contact events occurred, we found intraspecific contact rates (contacts per day) for raccoons, collared peccaries, and feral swine to be 0.81–124.77, 0.69–38.08, and 0.0–0.66, respectively. Throughout our study we distributed ~2,625 kg of whole kernel corn, which resulted in 6,351 contact events between marked wildlife (2.4 contacts per kg of corn). If 136 million kg of corn is distributed in Texas annually, we would expect >5.2 billion unnatural contact events between wildlife would result from this activity each year in Texas. Consequently, we do not believe that it is wise for natural resource managers to maintain artificial feeding sites for white-tailed deer or other wildlife due to pathogen transmission risks.     

Spatial analysis of habitat selection by Sitka black-tailed deer in Southeast Alaska,USA

We used a vector-based geographic information system (GIS) to examine habitat selection by radiocollared Sitka black-tailed deer (Odocoileus hemionus sitkensis) in logged forests of southeast Alaska. Our main objective was to explain deer habitat selection relative to old-growth/clear-cut edges and edge habitats at two different spatial scales. Deer home ranges contained higher percentages of recent clear-cuts (50–69%) than the study area (37%;P<0.01) and had higher old-growth/clear-cut edge densities than expected by chance (P<0.01). Deer relocation points were closer to old-growth/clear-cut edges (average=135 m) than random points located within each deer's relocation area (average=168 m;P=0.05). Likewise, deer relocations were closer to old-growth/clear-cut edges than points randomly located within old-growth stands or recent clear-cuts (P<0.01). As the size of clear-cuts increased, both deer relocation density and the proportion of a clear-cut occupied by deer home ranges decreased. Because old growth is important deer habitat and clear-cuts can produce deer forage for only 20–30 years after logging in southeast Alaska, deer management plans such as preserving entire watersheds and maintaining mixes of old growth and recent clear-cut have been proposed. Our data suggest that deer need a diversity of habitats near each other within their home ranges.     

Is legislation a barrier to the sustainable management of game species? A case study of wild deer in Britain

Wild game management for hunting in Western society has become increasingly complex as stakeholders have multiplied and as ‘sustainability’ influences the contemporary debate. This paper questions whether the current legal framework for game management, which has evolved from early European civilisations to focus on ‘hunting rights’, is relevant to regulate the contemporary environmental, social and economic dimensions of wild game and their management. Employing a narrative analysis to focus on deer, the study identifies key legislative tenets and highlights the pertinence of historical laws to contemporary conflicts. The analysis suggests that current legislation is increasingly divergent from contemporary trends and has created inertia with respect to sustainable deer management. The paper offers four options to redress this: state intervention; voluntary collaboration; financial incentives; and establishing a legal responsibility for management. It is concluded that significant innovation is required in one or more of these four areas to facilitate the contemporary sustainable management of wild deer in Britain.     

How has a shortage of census and geological information impeded the regularization of artisanal and small‐scale mining?

There is growing consensus that a combination of laissez‐faire policies, ad hoc regulation and debilitating support services has perpetuated socio‐economic and environmental deterioration in the artisanal and small‐scale mining (ASM) industry. However, a lack of anthropological and geological information on ASM prevents many governments both from improving the policy environment of the industry, and from providing more robust extension services to its operators. This article aims to examine more precisely how a deficiency of baseline census and geological data has inhibited industry formalization and undermined many of the measures implemented to address pressing problems at ASM sites. Specifically, it is argued that insufficient knowledge of artisanal mining populations — including their demographic structure — and of areas suitable for ASM activities affects the ability of a government to regularize, as well as to improve, the organization of this largely informal sector of industry. Case studies of Ghana and Zimbabwe are used to illustrate how the undertaking of low‐budget projects in areas of geological prospecting and population analysis could improve the efficiency of ASM assistance.     

Possible Input of Nitrogen of Visitors’ Origin on a Protected Peatland

The Ozegahara peatland, in the Nikko-Oze National Park in Japan, is ecologically significant because of its oligotrophic environment; it is one of the most strictly preserved areas in the country. The isotope ratio of nitrogen (¹⁵N/¹⁴N) and carbon (¹³C/¹²C) and C/N ratio of peat moss (Sphagnum spp.) and sundew (Drosera rotundifolia) in the peatland were analyzed. The correlation of the isotope ratio with some parameters (sundew population density, number of trapped insects, water level, bog myrtle coverage, and visitor density) was investigated. The nitrogen isotope ratio of sundew showed the most significant covariation with visitor density, where sundew from lunch areas or along busy walkways showed a higher nitrogen isotope ratio. The nitrogen isotope ratio of peat moss covaried, not only with route traffic frequency but also with water level, bog myrtle coverage, and number of trapped insects by sundew, indicating that factors other than the visitor level influence the local nitrogen cycle. This study suggests that the nitrogen imported into the peatland by visitors is a principal factor to be monitored for the maintenance of the natural environment.