The Holocene Dynamics of Vegetation and Ecological Conditions in the Polar Urals

Palynological, paleocarpological, and paleoentomological analyses of frozen peat deposits near Lake Pereval'noe, the Polar Urals, were performed to reveal the main stages of change in the pattern of vegetation over the period from the beginning of warming after the last Pleistocene glaciation to the late Holocene. Nine to four thousand years ago, the study region (at the present-day upper boundary of open larch forests) was covered with taiga forests, as the climate there was significantly warmer. These were larch–birch forests with an admixture of spruce and, later, spruce forests with larch and birch.     

Holocene dynamics of vegetation and ecological conditions in the southern Yamal Peninsula according to the results of comprehensive analysis of a relict peat bog deposit

A comprehensive analysis of samples from a frozen peat deposit in the upper reaches of the Khadytayakha River by palynological, carpological, entomological, and radiocarbon methods has been performed to reconstruct changes in the regional and local vegetation and climatic conditions during the Holocene. The results show that this peat deposit was formed from 8000 to 5000 years BP. During that period, the climate in the southern Yamal Peninsula was considerably warmer, and the present-day subarctic shrub tundra zone was occupied by plant communities of spruce, birch, and larch forests.     

Arthropods and birds in the forest–tundra of Western Siberia and the edge effect

Over 2500 species of invertebrate macrofauna and over 177 species of avifauna live in the forest–tundra of the Lower Ob region and adjacent areas of the northern taiga and shrub tundra. About half of the invertebrate species of the study area are northern—mostly hypo- and hemiarctic—tundra species; the rest are boreal and arctomontane-boreal Holarctic and trans-Palearctic species. The anthophilic and aerobiont insect assemblages include many boreal forest forms. A total of 153 species have been observed nesting at the taiga–foresttundra boundary; 81 species have been observed nesting at the tundra–forest-tundra boundary. The avifauna of the forest–tundra becomes enriched with taiga species in floodplain forests and with tundra species on the flat interfluve, in areas of tundra and tundra-like wetlands. The forest–tundra of the Lower Ob region has no avifauna and entomofauna of its own and contains a mixture of taiga and tundra faunas. It has been found that the ecotone effect is not pronounced in the taxonomic composition of birds and insects in the forest–tundra, where only some particular groups (such as amphibiont insects) have increased abundance.     

The holocene dynamics of vegetation and environmental conditions on the eastern slope of the Northern Urals

Consideration is given to changes in the vegetation and climatic conditions on the eastern slope of the Northern Urals in the second half of the Holocene as reconstructed on the basis of integrated palynological, botanical, paleocarpological, and radiocarbon analysis of material from a peat bog section in the floodplain of the Loz’va River. The results show that the northern taiga zone of the study region in the period between approximately 5000 and 700 years BP was occupied by forests of southern taiga facies, as the climate was significantly warmer than it is today.     

Pollution-induced successions in forests of the Kola Peninsula

Trends of changes in the successional status of northern taiga forests exposed to technogenic pollution in the Kola Peninsula are discussed. Main studies of digression in spruce and pine forests are characterized on the basis of the dynamics of species composition of phytocenoses and chemical composition of atmospheric fallout, soils, and dominant plants.     

Holocene Dynamics of Vegetation and Ecological Conditions in the Center of the East European Plain

Changes in the vegetation and fire regimes in the central East European Plain during the second half of the Holocene have been reconstructed based on the results of paleobotanical analysis and radiocarbon dating of material from a section of peat deposit in the Mordovia State Nature Reserve. It has been shown that birch–pine forests were widespread in the region between 7000 and 5000 yr BP, with the frequency of fires in that period being high (the fire return interval ranged from 10–20 to 100 years). Beginning from 5000 yr BP and to the early 20th century, broadleaf forests were dominant, with the fire return interval increasing to 300–500 years or longer.     

Mineral composition of assimilative organs of conifers after reduction of atmospheric pollution in the Kola Peninsula

An analysis has been made of changes in the mineral composition of conifer needles in northern taiga forests after reduction of emissions from the Severonickel Copper-Nickel Smelter Complex in the city of Monchegorsk, the most powerful source of air pollution in Northern Europe. The data obtained in permanent monitoring plots in 1991, 1993, 2000, and 2007 have been compared. The results provide evidence for reduction in the concentrations of heavy metals (Cu, Ni, Fe) and sulfur in the needles of spruce and pine trees in areas with different levels of air pollution. However, mineral nutrition of conifers has not been optimized. The plants are insufficiently supplied with nutrient elements such as P, Mg, Mn, and Zn. The contents of these and other elements essential for the normal functioning of needles (in particular, Ca, K, and P) continue to decrease, especially in pollution-induced sparse spruce forests. The most significant alterations of nutrient regime have been revealed in the Siberian spruce.     

First data on the Middle to Late Holocene dynamics of vegetation in the Upper Kama region

A detailed palynological record and the results of radiocarbon dating of sediments from the Dedyukhinskii floodplain massif in the vicinity of Lake Chashkinskoe (the Upper Kama region; 59°23′ N, 56°34.5′ E) have been used to reconstruct basic stages in the Middle to Late Holocene dynamics of vegetation. The results show that in the Atlantic period broadleaf tree species played a secondary role in forest formations of taiga and subtaiga types. Broadleaf–conifer forests became dominant in the Subboreal period, with fir widely spreading in the forests during its second half. During the Subatlantic period, forest formations acquired their recent taiga character.     

Climate changes and overstory recruitment of larch at the northern timberline in the Yamal Peninsula

Studies on overstory recruitment of the Siberian larch (Larix sibirica Ledeb.) in open larch forests depending on the time of seedling establishment (1880–1980) have been performed in the northern timberline ecotone on the Yamal Peninsula. Two periods of active larch establishment have been revealed: from 1900 to 1935 and from 1950 to 1970. A comparison of these data with the results of instrumental observations on temperature and precipitation has shown that the overstory recruitment of larch in valleys of northern rivers is largely dependent on July air temperatures in the first years of tree life, when seedlings grow 10–20 cm high.     

Structure and formation of conifer stands in the upper timberline ecotone on the North Chuya Ridge,Central Altai

The structure of Siberian stone pine (Pinus sibirica Du Tour.) and Siberian larch (Larix sibirica Ledeb.) stands and specific features of their formation have been studies in the forest-tundra ecotone on the North Chuya Ridge (2235–2475 m a.s.l.). Changes in the structure of these stands along the transition from the upper boundary of closed forests to the high-mountain tundra have proved to have an ambiguous pattern. Both tree species form mixed clusters of similar-aged trees in the lower part of the ecotone but grow singly, in scatters, in its upper part. The formation of conifer stands (tree clusters) in the lower part of the ecotone, on the slopes of the Aktru River valley, began during climate warming in the second half of the 19th century. The expansion of confers to its upper part took place markedly later, in the early 20th century (Siberian larch) or even in the 1930s (Siberian stone pine).     

The Holocene Evolution of Vegetation in the Southeast of the Volga Upland

The Holocene evolution of vegetation in the high-plain territory of the northwestern Volga Upland has been shown to have certain specific features. Based on analysis of spore–pollen assemblages from peat deposits, the onset of bog formation has been dated to the late Boreal period (about 8500 yr BP); the pine–birch phase in the development of forests, to the early Atlantic period (8000?6000 yr BP); the appearance and peak development of the zonal broadleaf species complex, to the late Atlantic period (6000?4500 yr BP); and early anthropogenic changes in the vegetation, to about 3000 yr BP.     

Spatiotemporal characteristics of wildfire frequency and relative area burned in larch-dominated forests of Central Siberia

Wildfire frequency, relative area burned, and fire return intervals (FRI) have been studied in larchdominated forests along the transect from the southern (45° N) to the northern (73° N) distribution limits of larch stands based on analysis of satellite imagery (NOAA/AVHRR, Terra/MODIS; 1996–2015) and collection of tree cross cuts with fire scars. A significant increasing trend in fire extent (R² = 0.50, p < 0.05) has been revealed. Histograms of fire extent and frequency are bimodal in the southern and middle taiga (with peaks in spring–summer and late summer–autumn periods) but become unimodal toward the north (>55° N). The length of FRI increases from 80 years at 62° N to ~200 years at the Arctic Circle and reaches ~300 years near the northern limit of larch stands, showing a significant inverse correlation with the length of fire season (r =–0.69). In turn, the length of fire season, area burned and FRI are closely correlated with latitudinal variation in solar irradiance (r = 0.97, 0.81, and –0.95, respectively).     

Holocene dynamics of forest ecosystems on the upper plateau of the Volga Upland

Main stages in the development of forest ecosystems on the upper plateau of the Volga Upland in the Holocene have been reconstructed by analyzing palynological assemblages from peat deposits. Dominance of the forest landscape in the region began in the Boreal time. Pine and, to a lesser extent, birch were the main forest-forming species, whereas the role of spruce was never significant. Broad-leaved species appeared in forests in the early Atlantic time. The formation of floodplain and upland bogs began at the end of the Boreal time and in the first half of the Atlantic time, respectively. Significant anthropogenic changes in the vegetation began in the Subboreal time.     

Indicatory aspect of the geographic analysis of plant communities with respect to floristic composition on mountain slopes differing in exposure (the Polar Urals)

Latitudinal geographic groups of vascular plant, moss, and lichen species included in tundra communities and lower vegetation layers of larch forest and open forest have been analyzed with respect to their distribution according to different altitudinal levels and slopes differing in exposure. Prevailing plant groups have been identified. The results were compared with corresponding data on the composition of plant communities in the neighboring flatland area (Kharp Research Station).     

Dynamics of snowmelt water composition in conifer forests exposed to airborne industrial pollution

An analysis is made of the spatial variability of snowmelt water composition (within and between biogeocenoses), with regard to its long-term dynamics, in pine and spruce forests exposed to airborne industrial pollution from the Europe’s largest Severonikel Copper–Nickel Smelter Complex. Snowmelt waters from under the tree canopy, compared to those from intercrown areas, contain higher concentrations of chemical elements due to their washing and leaching from tree crowns. This is especially true of spruce forests, since the crowns of spruce trees have a high sorption capacity. Distinct trends in the long-term dynamics of snowmelt water composition, related to reduction of industrial emissions, are observed in background forest areas and defoliating forests but not in the vicinity of pollution sources. It is shown that the main factors determining these dynamics in forests of the Kola Peninsula are edificator tree species, airborne industrial pollution, and, possibly, an increase in the number of days with above-zero temperatures in the period of snow accumulation, which facilitates washing and leaching of chemical compounds from tree crowns.     

Vegetation-permafrost relationships in the zone of sporadic permafrost distribution in the Kamchatka Peninsula

Studies on vegetation and permafrost table depth in the zone of sporadic permafrost distribution in the Uksichan River valley (the central Kamchatka Peninsula) have provided evidence that these components of biogeocenosis are interrelated and develop coordinately. In open larch forests with green forest mosses dominating in the ground vegetation layer, the permafrost table lies approximately 60 cm below the soil surface. When the ground vegetation layer is dominated by sphagnums, the permafrost table rises to 40–20 cm. In areas with a dwarf shrub-lichen ground layer, the soil thaw depth increases. A hypothesis is proposed that cyclic successional replacement of plant communities may take place in open larch forests on permafrost soils, including four consecutive stages with dominance of green mosses, sphagnums, lichens, and dwarf shrubs. In areas disturbed by fires, pioneer moss or herbaceous communities develop in the ground layer.     

Diet structure of the Siberian roe deer in Yakutia

Daily food requirements and seasonal aspects of the diet have been studied in the Siberian roe deer (Capreolus pygargus Pall.) at the northernmost limits of its range in northeastern Asia, under specific sharply continental climatic conditions of the East Siberian light conifer (larch) taiga forests. Seasonal changes in the composition of the diet and the consumption of different food items are discussed.     

Expansion of Evergreen Conifers to the Larch-Dominated Zone and Climatic Trends

The expansion of so-called evergreen conifers (EGCs), including Siberian stone pine, spruce, and fir, along the transect oriented from the boundary of the larch-dominated zone (LDZ; mixed forests of the Yenisei Ridge) to its center has been studied. The normalized dispersal coefficient calculated as K _i = (n _i – N _i )/(n _i + N _i ), where n _i and N _i are the relative numbers of the ith species in the undergrowth and the upper layer, respectively, serves as an indicator of the expansion. It has been found that the K _i values for EGCs (and birch) are higher than the K _i of larch even in the zone absolutely dominated by larch, where the relative numbers of EGCs in the upper layer is less than 1%. The EGC undergrowth has mainly been formed during the past 20–30 years, which is correlated with the trend of summer temperatures The spread of EGCs in the LDZ depends on the frequency of forest fires. The decrease in the time intervals between fires in the 20th century to 65 years (versus 100 years in the 19th century) may have prevented the expansion of competing species in the LDZ. The results obtained indicate that EGCs and birch penetrate into the zone traditionally dominated by larch, which is related to climatic changes during the past three decades. At the same time, tree stand density is increasing in the forest-tundra ecotone, and larch is spreading further into the tundra zone.__________Translated from Ekologiya, No. 3, 2005, pp. 186–193.Original Russian Text Copyright © 2005 by Kharuk, Dvinskaya, Ranson, Im.     

Dynamics of vegetation in the tundra-taiga ecotone on the Kola Peninsula depending on climate fluctuations

The dynamics of northern forest boundary and the state of vegetation in the tundra-taiga ecotone in the northwestern Kola Peninsula have been evaluated using Landsat satellite images obtained in 1986 and 2005, the years encompassing the period of recent climate warming. Values of the normalized difference vegetation index (NDVI) in these years have been calculated to construct a differential index image reflecting changes in the vegetation. Its analysis has revealed an increase in the density of open forests and in the thickness of dwarf shrub vegetation, which has expanded to the lichen tundra.     

Spiders in mountain habitats of the Giant Mountains

Ground-living spiders were studied using modified pitfall traps during several years in four characteristic habitats in Giant Mountains (Krkono?e Mts.), the High Sudetes, Czech Republic: alpine tundra, subalpine mire, tall-herb stand at the bottom of a glacial corrie, and decaying mountain spruce forest. Ecological and zoogeographical aspects of spider communities were analysed. The spider communities of alpine tundra, subalpine mire, and glacial corrie exhibited long-term stability, whereas the community of decaying mountain forest changed during observations. Small linyphiid spiders, dominating in mature forest, were gradually replaced by larger Iycosid and gnaphosid species. Zoogeographic characterization of mountain habitats was made based on species exhibiting disjunctive area. In contrast to plants, for spiders of boreal origin alpine tundra is the most important habitat for survival, followed by screes, mires, spruce forests, and corries.