Comparison of Agroforests and Protected Forests in the East Usambara Mountains,Tanzania

Comparative studies on plant species richness, endemism, floristic composition, and structure between protected and unprotected forests are few in the Eastern Arc Mountains, one of the most biodiverse ecosystems in Africa. This study from one mountain range, the East Usambaras, examines floristic and structural tree data from 41–0.5 ha plots in four types of Eastern Arc forest: active agroforests, recently abandoned agroforests, mature secondary forest, and natural forest. Active agroforests had significantly lower tree species richness, endemic species richness, and stand density compared to natural and mature secondary forest. Recently abandoned agroforests contained a higher tree species richness, density, and tree height than active agroforests. Active and abandoned agroforests were dominated by an invasive tree, Maesopsis eminii. This tree species makes up a large percentage of the stems in active agroforests (26%), recently abandoned agroforests (32%), and in the canopy of mature secondary forests ∼ 30 years post logging (30%). Through time the increasing dominance of this non-native tree in active agroforests is a concern when considering the role of agroforests in a landscape scale conservation strategy.     

Effects of Logged and Unlogged Forest Patches on Avifaunal Diversity

In the Hyrcanian forests of northern Iran, reduced-impact silviculture systems, (single-tree and group-tree selection) were applied over a large area, which generated different local habitat structures. The aim of this study was to assess the differences between treated and untreated areas of forest and their effect on avian richness, abundance and diversity (R.A.D). Birds were surveyed during the breeding season in 2009 by 100-point counts, equally distributed in the treated and untreated area. Avian R.A.D was significantly different and higher in the untreated area. Generally, forestry practices cause noticeable changes in canopy percentage, tree composition, snags and shrub number. Treated forest habitats in the area of study had a much more developed understory, fewer snags and fewer large diameter trees. The results highlighted the importance of forest maturity and showed that preventing silvicultural disturbances may not be the best solution for conserving and enhancing biodiversity. Rather, methods such as selective cutting seem an appropriate and sustainable way of forest management. It is suggested that forests should be managed to conserve structural elements which create favorable habitat for bird species, preventing future species losses due to logging practices.     

Postfire regeneration in Pinus pinea L. and Pinus pinaster aiton in Andalucia (Spain)

The objective of this study was to examine postfire regeneration of tree, shrub, and dwarf shrub species, in relation to levels of damage in four planted pine forests (Pinus pinea, Pinus pinaster) in Andalusia. A prefire vegetation map was used for detailing species composition, vertical structure, and density and another for detailing the extent and intensity of fire damage. Between 3 and 7 years after the fires, an inventory was made of the vegetation in each area, using the step-point method. The information thus obtained was used to determine the amount of cover in the dwarf/shrub and tree layers, the frequency of species in each of the layers, floristic richness, and diversity (Shannon index). The botanical composition of the dwarf and shrub layer was analyzed using TWINSPAN. Variables were poorly correlated with level of fire damage, which suggests that the forests in this study followed the autosuccession model. Because of the artificial origin or seminatural condition, regeneration of the dominant tree species is poor, and it seems unlikely that forests will recover to their prefire state. Therefore action is recommended to restore these ecosystems.     

Postfire Regeneration in <Emphasis Type="Italic">Pinus pinea</Emphasis> L. and <Emphasis Type="Italic">Pinus pinaster</Emphasis> Aiton in Andalucia (Spain)

The objective of this study was to examine postfire regeneration of tree, shrub, and dwarf shrub species, in relation to levels of damage in four planted pine forests (Pinus pinea, Pinus pinaster) in Andalusia. A prefire vegetation map was used for detailing species composition, vertical structure, and density and another for detailing the extent and intensity of fire damage. Between 3 and 7 years after the fires, an inventory was made of the vegetation in each area, using the step-point method. The information thus obtained was used to determine the amount of cover in the dwarf/shrub and tree layers, the frequency of species in each of the layers, floristic richness, and diversity (Shannon index). The botanical composition of the dwarf and shrub layer was analyzed using TWINSPAN. Variables were poorly correlated with level of fire damage, which suggests that the forests in this study followed the autosuccession model. Because of the artificial origin or seminatural condition, regeneration of the dominant tree species is poor, and it seems unlikely that forests will recover to their prefire state. Therefore action is recommended to restore these ecosystems.     

Assessment of frequent cutting as a plant-community management technique in power-line corridors

Repeated cutting of vegetation at or near ground level in power-line corridors is a common practice for inhibiting tree growth and regeneration. However, few data exist on long-term community responses. In this study, we sampled 20 northern Kentucky power-line corridors and compared their seedling and sapling communities to the edges and interiors of adjacent undisturbed forests. Mean seedling and sapling density in corridors was roughly twice that of adjacent undisturbed forest interiors, suggesting that repeated cutting is not a viable method of inhibiting tree regeneration. Corridor communities were dominated byRobinia pseudoacacia (black locust) andFraxinus americana (white ash), but ordinations indicated strong similaritties among communities in corridors and adjacent forests. Many of the tree species found in adjacent forests, with the exception of a few shade-tolerant species, had highest seedling and sapling densities in corridors. Stump or root sprouting by many species appears to regenerate forests quickly after cutting. However, disturbed soil and detritus accumulations caused by management crews and their equipment may also create a large variety of microsites for seedling establishment. Because repeated cutting selects for dominance by species with highest sprout growth rates, it should not be used as the sole management technique. It may instead be used to alter the vigor, stature, and stored reserves of trees so that herbicides or other methods of tree control can be used more efficiently.     

Woody Vegetation Composition and Structure in Peri-urban Chongming Island, China

Chongming, the world’s largest alluvial island, is located within the municipality of Shanghai, China. Recent projects have now linked peri-urban Chongming to Shanghai’s urban core and as a result will soon undergo substantial changes from urbanization. We quantitatively analyzed the structure and composition of woody vegetation across subtropical, peri-urban Chongming as a basis for sustainable management of these rapidly urbanizing subtropical ecosystems elsewhere. We used 178 permanent, random plots to statistically and spatially analyze woody plant composition and tree structure across the 1,041 km² of Chongming. A total of 2,251 woody plants were measured comprising 42 species in 37 genera. We statistically and geospatially analyzed field data according to land uses and modeled air pollution removal by trees. Average tree diameter at breast height, total height, and crown widths on transportation land uses were greater than other land uses. These same values were lowest on forest land use and greater tree cover was associated with areas of increased anthropogenic activity. Less than 20 % of the woody vegetation was exotic and a species richness index was significantly different between land uses due to legacy effects. Composition of agriculture and forest land uses were similar to residential and transportation. Tree cover across Chongming was also estimated to annually remove 1,400 tons of air pollutants. We propose that this integrated and quantitative method can be used in other subtropical, peri-urban areas in developing countries to establish baseline trends for future sustainability objectives and to monitor the effects of urbanization and climate change.     

Effects of anthropogenic fire history on savanna vegetation in northeastern Namibia

Anthropogenic fires in Africa are an ancient form of environmental disturbance, which probably have shaped the savanna vegetation more than any other human induced disturbance. Despite anthropogenic fires having played a significant role in savanna management by herders, previous ecological research did not incorporate the traditional knowledge of anthropogenic fire history. This paper integrates ecological data and anthropogenic fire history, as reconstructed by herders, to assess landscape and regional level vegetation change in northeastern Namibia. We investigated effects of fire frequency (i.e. <5, 5-10 and >10 years) to understand changes in vegetation cover, life form species richness and savanna conditions (defined as a ratio of shrub cover to herbaceous cover). Additionally, we analysed trends in the vegetation variables between different fire histories at the landscape and regional scales. Shrub cover was negatively correlated to herbaceous cover and herbaceous species richness. The findings showed that bush cover homogenisation at landscape and regional scales may suggest that the problem of bush encroachment was widespread. Frequent fires reduced shrub cover temporarily and promoted herbaceous cover. The effects on tree cover were less dramatic. The response to fire history was scale-independent for shrub, herbaceous and tree cover, but scale-dependent for the richness of grass and tree life forms. Fire history, and not grazing pressure, improved savanna conditions. The findings emphasise the need to assess effects of anthropogenic fires on vegetation change before introducing new fire management policies in savanna ecosystems of northeastern Namibia.     

Seed bank composition of open and overgrown calcareous grassland soils--a case study from Southern Belgium

The success of calcareous grassland recreation following abandonment depends to a large extent on the composition of the soil seed bank. We studied the species richness and composition of the seed bank along a chronosequence from well-developed calcareous grassland to scrub and forest vegetation, which had developed on calcareous grassland over the last 225 years. The seed bank density was highest in the calcareous grassland soils (930 seeds/m(2)). However, the seed bank was mainly composed of rather common species of nutrient poor grassland, which were poorly represented in the surface vegetation. There were no significant differences in soil seed bank density and species composition between the calcareous grassland and scrub vegetation that was less than 15 years old, largely because several grassland species had persisted in the scrub vegetation and were therefore able to replenish the soil seed bank. In contrast seed density and species richness declined significantly after 40 years of grassland abandonment. Indeed, forest soils had the lowest seed densities (214 seeds/m(2)) with only a few grassland species represented. This reflects the lack of grassland species in the field layer of the forest, and, therefore, the lack of seed production and seed bank replenishment. It is clear that recreation of calcareous grassland on long abandoned sites cannot rely on germination of target species from the seed bank alone. Even in the calcareous grassland soil, seeds of target species are not abundant as the majority produces transient seeds, which decay rapidly if they do not germinate immediately. Successful grassland recreation on such sites therefore may require seeds of target species to be introduced artificially (e.g. as seed mixtures, green hay, etc.). Alternatively, reinstatement of traditional practices, such as grazing or mowing, will increase the natural dispersal potential of these species, allowing population reestablishment in the long term.     

A Multi-scale Spatial Analysis of Native and Exotic Plant Species Richness Within a Mixed-Disturbance Oak Savanna Landscape

Impacts of human land use pose an increasing threat to global biodiversity. Resource managers must respond rapidly to this threat by assessing existing natural areas and prioritizing conservation actions across multiple spatial scales. Plant species richness is a useful measure of biodiversity but typically can only be evaluated on small portions of a given landscape. Modeling relationships between spatial heterogeneity and species richness may allow conservation planners to make predictions of species richness patterns within unsampled areas. We utilized a combination of field data, remotely sensed data, and landscape pattern metrics to develop models of native and exotic plant species richness at two spatial extents (60- and 120-m windows) and at four ecological levels for northwestern Ohio’s Oak Openings region. Multiple regression models explained 37–77 % of the variation in plant species richness. These models consistently explained more variation in exotic richness than in native richness. Exotic richness was better explained at the 120-m extent while native richness was better explained at the 60-m extent. Land cover composition of the surrounding landscape was an important component of all models. We found that percentage of human-modified land cover (negatively correlated with native richness and positively correlated with exotic richness) was a particularly useful predictor of plant species richness and that human-caused disturbances exert a strong influence on species richness patterns within a mixed-disturbance oak savanna landscape. Our results emphasize the importance of using a multi-scale approach to examine the complex relationships between spatial heterogeneity and plant species richness.     

Effects of Coffee Management Intensity on Composition, Structure, and Regeneration Status of Ethiopian Moist Evergreen Afromontane Forests

The effect of arabica coffee management intensity on composition, structure, and regeneration of moist evergreen Afromontane forests was studied in three traditional coffee-management systems of southwest Ethiopia: semiplantation coffee, semiforest coffee, and forest coffee. Vegetation and environmental data were collected in 84 plots from forests varying in intensity of coffee management. After controlling for environmental variation (altitude, aspect, slope, soil nutrient availability, and soil depth), differences in woody species composition, forest structure, and regeneration potential among management systems were compared using one way analysis of variance. The study showed that intensification of forest coffee cultivation to maximize coffee production negatively affects diversity and structure of Ethiopian moist evergreen Afromontane forests. Intensification of coffee productivity starts with the conversion of forest coffee to semiforest coffee, which has significant negative effects on tree seedling abundance. Further intensification leads to the conversion of semiforest to semiplantation coffee, causing significant diversity losses and the collapse of forest structure (decrease of stem density, basal area, crown closure, crown cover, and dominant tree height). Our study underlines the need for shade certification schemes to include variables other than canopy cover and that the loss of species diversity in intensively managed coffee systems may jeopardize the sustainability of coffee production itself through the decrease of ecosystem resilience and disruption of ecosystem services related to coffee yield, such as pollination and pest control.     

Maximizing Benefits from Riparian Revegetation Efforts: Local- and Landscape-Level Determinants of Avian Response

With limited financial resources available for habitat restoration, information that ensures and/or accelerates success is needed to economize effort and maximize benefit. In the Central Valley of California USA, riparian habitat has been lost or degraded, contributing to the decline of riparian-associated birds and other wildlife. Active restoration of riparian plant communities in this region has been demonstrated to increase local population sizes and species diversity of landbirds. To evaluate factors related to variation in the rate at which bird abundance increased after restoration, we examined bird abundance as a function of local (restoration design elements) and landscape (proportion of riparian vegetation in the landscape and riparian patch density) metrics at 17 restoration projects within five project areas along the Sacramento River. We developed a priori model sets for seven species of birds and used an information theoretic approach to identify factors associated with the rate at which bird abundance increased after restoration. For six of seven species investigated, the model with the most support contained a variable for the amount of riparian forest in the surrounding landscape. Three of seven bird species were positively correlated with the number of tree species planted and three of seven were positively correlated with the planting densities of particular tree species. Our results indicate that restoration success can be enhanced by selecting sites near existing riparian habitat and planting multiple tree species. Hence, given limited resources, efforts to restore riparian habitat for birds should focus on landscape-scale site selection in areas with high proportions of existing riparian vegetation.     

The Importance of Temporal and Spatial Vegetation Structure Information in Biotope Mapping Schemes: A Case Study in Helsingborg,Sweden

Temporal and spatial vegetation structure has impact on biodiversity qualities. Yet, current schemes of biotope mapping do only to a limited extend incorporate these factors in the mapping. The purpose of this study is to evaluate the application of a modified biotope mapping scheme that includes temporal and spatial vegetation structure. A refined scheme was developed based on a biotope classification, and applied to a green structure system in Helsingborg city in southern Sweden. It includes four parameters of vegetation structure: continuity of forest cover, age of dominant trees, horizontal structure, and vertical structure. The major green structure sites were determined by interpretation of panchromatic aerial photographs assisted with a field survey. A set of biotope maps was constructed on the basis of each level of modified classification. An evaluation of the scheme included two aspects in particular: comparison of species richness between long-continuity and short-continuity forests based on identification of woodland continuity using ancient woodland indicators (AWI) species and related historical documents, and spatial distribution of animals in the green space in relation to vegetation structure. The results indicate that (1) the relationship between forest continuity: according to verification of historical documents, the richness of AWI species was higher in long-continuity forests; Simpson’s diversity was significantly different between long- and short-continuity forests; the total species richness and Shannon’s diversity were much higher in long-continuity forests shown a very significant difference. (2) The spatial vegetation structure and age of stands influence the richness and abundance of the avian fauna and rabbits, and distance to the nearest tree and shrub was a strong determinant of presence for these animal groups. It is concluded that continuity of forest cover, age of dominant trees, horizontal and vertical structures of vegetation should now be included in urban biotope classifications.     

Associations between forest characteristics and socio-economic development: a case study from Portugal

The integration of socio-economic and environmental objectives is a major challenge in developing strategies for sustainable landscapes. We investigated associations between socio-economic variables, landscape metrics and measures of forest condition in the context of Portugal. The main goals of the study were to 1) investigate relationships between forest conditions and measures of socio-economic development at national and regional scales, 2) test the hypothesis that a systematic variation in forest landscape metrics occurs according to the stage of socio-economic development and, 3) assess the extent to which landscape metrics can inform strategies to enhance forest sustainability. A ranking approach and statistical techniques such as Principal Component Analysis were used to achieve these objectives. Relationships between socio-economic characteristics, landscape metrics and measures of forest condition were only significant in the regional analysis of municipalities in Northern Portugal. Landscape metrics for different tree species displayed significant variations across socio-economic groups of municipalities and these differences were consistent with changes in characteristics suggested by the forest transition model. The use of metrics also helped inform place-specific strategies to improve forest management, though it was also apparent that further work was required to better incorporate differences in forest functions into sustainability planning.     

Chemical properties of soils in relation to forest composition in moist temperate valley slopes of Garhwal Himalaya, India

The present study was undertaken in moist temperate forest of Mandal-Chopta area in the Garhwal region of Uttarakhand, India. The aim of the present study was to assess the chemical properties of soils in relation to the forest structure and composition. Twelve forest types according to the altitude, slope aspect, and species composition were selected for the study. Chemical properties of the soil, i.e., total nitrogen (N), available phosphorus (P), available potassium (K), organic carbon (C), soil organic matter (SOM), pH and C:N ratios were analyzed for three different depths viz., (i) ??Upper?? (0?C10?cm), (ii) ??Middle?? (11?C30?cm), and (iii) ??Lower?? (31?C60?cm) in all the selected forest types. Phytosociological and diversity parameters viz. total basal cover (Gha^?1), stem density (Nha^?1), tree species richness (SR), Simpson concentration of dominance and Shannon?CWiener diversity index ( $ \overline{\rm H} $ ) were also calculated for each forest type. Values of N, P, K, C, SOM, C:N ratio and pH ranged from 0.17 to 0.45?%, 2.73 to 20.17?ppm, 40.67 to 261.17?ppm, 2.29 to 4.31?%, 3.95 to 7.43?%, 8.12 to 14.49 and 5.47 to 6.67, respectively. N showed a positive relationship with C and K. P was positively correlated with C and negatively correlated with altitude. P was higher in the soil of lower horizons of all the forest types. K was found to be positively correlated with altitude, C and N. pH of all the forest types was slightly acidic. C was comparatively higher in mixed broad-leaved forest types. No relationship between altitude and C was observed, which may be due to different composition of forest types along the altitudinal gradient and their differential decomposition rates. C content decreased with the depth of the soil. C showed positive correlation with N, P, and K. N showed a positive correlation with Gha^?1. K showed a negative correlation with SR and $ \overline{\rm H} $ . P showed no relationship with the phytosociological parameters. This study also provides the comparisons between the results of chemical analysis of the present study with numerous other previous studies in the temperate Himalayan region of the Uttarakhand. The values of soil chemical properties of the present study in most of the forest types are on the higher end than the values previously recorded for the other similar forests of the region. The possible reason being luxuriant vegetation and undisturbed nature of these forest types, which is evident from higher values of diversity and other phytosociological parameters.     

A Water Yield‐Oriented Practical Approach for Multifunctional Forest Management and its Application in Dryland Regions of China

Mountainous forest areas are vitally important for water supply in dryland regions which suffer from high erosion risk and severe water shortage. Massive afforestation, mainly for erosion control, may reduce the water yield and threaten local water supply security. Moreover, many over‐dense forests due to a strict logging ban policy have produced remarkably negative impacts for both forests (e.g., low timber quality, restricted natural regeneration, and high stand instability) and water yield. To satisfy the rapidly increasing demands on water supply and other services, a practical approach for managing forest stands in a multifunctional way, which particularly addresses water yielding, is urgently required. For this purpose, we integrated the existing knowledge and experience, designed an “ideal” stand structure to represent multifunctional forest (MFF) and determined its key parameters (a ground coverage of >0.7, a canopy density around 0.7, and an H/DBH ratio (tree height [m] to the diameter at breast height [cm]) of <0.7). Moreover, a decision process for MFF stand management was recommended as: (1) investigating the site quality; (2) identifying the site‐specific main forest functions; (3) quantifying the stand structure; (4) diagnosing the stand structure by comparing with the “ideal” one; and (5) arranging the functions/structure‐oriented management measures. In this way, the water‐yielding function can be improved and meanwhile other forest functions can be promoted.     

Integration of Satellite Imagery and Forest Inventory in Mapping Dominant and Associated Species at a Regional Scale

To achieve the overall objective of restoring natural environment and sustainable resource usability, each forest management practice effect needs to be predicted using a simulation model. Previous simulation efforts were typically confined to public land. Comprehensive forest management practices entail incorporating interactions between public and private land. To make inclusion of private land into management planning feasible at the regional scale, this study uses a new method of combining Forest Inventory and Analysis (FIA) data with remotely sensed forest group data to retrieve detailed species composition and age information for the Missouri Ozark Highlands. Remote sensed forest group and land form data inferred from topography were integrated to produce distinct combinations (ecotypes). Forest types and size classes were assigned to ecotypes based on their proportions in the FIA data. Then tree species and tree age determined from FIA subplots stratified by forest type and size class were assigned to pixels for the entire study area. The resulting species composition map can improve simulation model performance in that it has spatially explicit and continuous information of dominant and associated species, and tree ages that are unavailable from either satellite imagery or forest inventory data. In addition, the resulting species map revealed that public land and private land in Ozark Highlands differ in species composition and stand size. Shortleaf pine is a co-dominant species in public land, whereas it becomes a minor species in private land. Public forest is older than private forest. Both public and private forests have deviated from historical forest condition in terms of species composition. Based on possible reasons causing the deviation discussed in this study, corresponding management avenues that can assist in restoring natural environment were recommended.     

CHANNEL RESPONSE FROM SHRUB DOMINATED RIPARIAN COMMUNITIES AND ASSOCIATED EFFECTS ON SALMONTD HABITAT1

ABSTRACT: We surveyed first‐to third‐order streams (channel widths from 1.4 to 10 m) in the southeastern slopes of the Cascade Range of Washington and found two distinct endpoints of riparian vegetation. Where the forest overstory is dominated by park‐like Ponderosa pine (Pinus ponderosa), channels are commonly bordered with a dense scrub‐shrub vegetation community. Where fire suppression and/or lack of active riparian zone management have resulted in dense encroachment of fir forests that create closed forest canopies over the channel, scrub‐shrub vegetation communities are virtually absent near the channel. Other factors being equal, distinct differences in channel morphology exist in streams flowing thru each riparian community. The scrub‐shrub channels have more box‐like cross‐sections, lower width‐to‐depth ratios, more pools, more undercut banks, more common sand‐dominated substrates, and similar amounts of woody debris (despite lower tree density). Temperature comparisons of forest and scrub‐shrub sections of two streams indicate that summer water temperatures are slightly lower in the scrub‐shrub streams. We surmise that these morphology and temperature effects are driven by differences in root density and canopy conditions that alter dynamic channel processes between each riparian community. We suspect that the scrub‐shrub community was more common in the landscape prior to the 20th century and may have been the dominant native riparian community for these stream types. We therefore suggest that managing these streams for dense riparian conifer does not mimic natural conditions, nor does it provide superior in‐stream habitat.     

Large Woody Debris Input and Its Influence on Channel Structure in Agricultural Lands of Southeast Brazil

Riparian forests are important for the structure and functioning of stream ecosystems, providing structural components such as large woody debris (LWD). Changes in these forests will cause modifications in the LWD input to streams, affecting their structure. In order to assess the influence of riparian forests changes in LWD supply, 15 catchments (third and fourth order) with riparian forests at different conservation levels were selected for sampling. In each catchment we quantified the abundance, volume and diameter of LWD in stream channels; the number, area and volume of pools formed by LWD and basal area and tree diameter of riparian forest. We found that riparian forests were at a secondary successional stage with predominantly young trees (diameter at breast height <10 cm) in all studied streams. Results showed that basal area and diameter of riparian forest differed between the stream groups (forested and non-forested), but tree density did not differ between groups. Differences were also observed in LWD abundance, volume, frequency of LWD pools with subunits and area and volume of LWD pools. LWD diameter, LWD that form pools diameter and frequency of LWD pools without subunits did not differ between stream groups. Regression analyses showed that LWD abundance and volume, and frequency of LWD pools (with and without subunits) were positively related with the proportion of riparian forest. LWD diameter was not correlated to riparian tree diameter. The frequency of LWD pools was correlated to the abundance and volume of LWD, but characteristics of these pools (area and volume) were not correlated to the diameter of LWD that formed the pools. These results show that alterations in riparian forest cause modifications in the LWD abundance and volume in the stream channel, affecting mainly the structural complexity of these ecosystems (reduction in the number and structural characteristics of LWD pools). Our results also demonstrate that riparian forest conservation actions must consider not only its extension, but also successional stage to guarantee the quantity and quality of LWD necessary to enable the structuring of stream channels.     

Wood carbon content of tree species in Eastern China: interspecific variability and the importance of the volatile fraction

Data on the mass density and carbon content of tree organs, and in particular stem wood, are essential for accurate assessments of forest carbon sequestration. However most available data, including that for East Asia, has neglected the volatile C fraction. Wood samples were collected and assayed for C content from 14 native tree species in Jilin Province, NE China. C content showed statistically significant variation among species, ranging from 48.4% to 51.0%. The volatile C fraction was non-negligible, averaging 2.2%, and showed high variation among species. As found in prior studies, wood C content was appreciably higher in conifer than hardwood (angiosperm) species (50.8+/-0.1% vs. 49.5+/-0.2%, respectively). Wood carbon density (gC/cm(3)) showed very high inter-specific variation, due mainly to differences in wood specific gravity. Our analyses, in conjunction with recently published data from North America, indicate a global mean value of 47.5+/-0.5% wood C content exclusive of volatile C; the widely used 50% figure corresponds more closely to total wood C inclusive of the volatile fraction. Failure to include volatile C or to use species- or higher-taxon-specific C content values in forest C assessments is likely to introduce biases on the order approximately 4-6%. In addition, the stocks and flows of the volatile C fraction in wood are in themselves an important and sorely neglected aspect of forest C processes likely to be strongly impacted by harvests and other management practices.     

Introduced species and management of a Nothofagus/Austrocedrus forest

Isla Victoria (Nahuel Huapi National Park, Argentina), a large island dominated by native Nothofagus and Austrocedrus forest, has old plantations of many introduced tree species, some of which are famed invaders of native ecosystems elsewhere. There are also large populations of introduced deer and shrubs that may interact in a complex way with the introduced trees, as well as a recently arrived population of wild boar. Long-standing concern that the introduced trees will invade and transform native forest may be unwarranted, as there is little evidence of progressive invasion, even close to the plantations, despite over 50 years of opportunity. Introduced and native shrubs allow scattered introduced trees to achieve substantial size in abandoned pastures, but in almost all areas neither the trees nor the shrubs appear to be spreading beyond these sites. These shrub communities may be stable rather than successional, but the technology for restoring them to native forest is uncertain and probably currently impractical. Any attempt to remove the exotic tree seedlings and saplings from native forest would probably create the very conditions that would favor colonization by exotic plants rather than native trees, while simply clear-cutting the plantations would be unlikely to lead to regeneration of Nothofagus or Austrocedrus. The key to maintaining native forest is preventing catastrophic fire, as several introduced trees and shrubs would be favored over native dominant trees in recolonization. Deer undoubtedly interact with both native and introduced trees and shrubs, but their net effect on native forest is not yet clear, and specific management of deer beyond the current hunting by staff is unwarranted, at least if preventing tree invasion is the goal. The steep terrain and shallow soil make the recently arrived boar a grave threat to the native forest. Eradication is probably feasible and should be attempted quickly.