Nest-Site Selection by Black Sparrowhawks Accipiter melanoleucus: Implications for Managing Exotic Pulpwood and Sawlog Forests in South Africa

This study provides timber growers with silvicultural guidelines for establishing and maintaining nest-tree habitat for native black sparrowhawks (Accipiter melanoleucus) in commercial planted forests in South Africa. In this country, exotic eucalypts and pines are planted principally for pulpwood and sawlog production. Nineteen nests were sampled in indigenous forests and 58 nests in exotic forests. Although mean nest heights differed between indigenous and exotic trees, in all trees, nests were positioned, on average, at 64% of tree height. Black sparrowhawks nested near stand edges, probably seeking a compromise between nesting adjacent to open hunting habitat and selecting an insulated tree from within the forest. Black sparrowhawks nested in tall trees ( X- = 18-33 m for different tree species classes) with a large diameter (>60 cm). Unfortunately, the South African pulpwood and sawlog industry employ short rotations (<16 years) and high tree densities (>700 trees/ha) that do not allow the trees to attain the characteristics suitable for black sparrowhawk nesting sites. Eucalypt and pine nest stands must be of 25 x 25 m minimum size and incorporate 10 trees at minimum heights of 21 and 18 m and diameters of 37 and 35 cm, respectively. If such nest-tree stands are set aside as islands in a sea of commercial forests, and black sparrowhawks and other forest raptors nest in them, timber growers will improve the tree-nesting raptor diversity of planted forests. If, however, these raptors prey upon species of conservation importance, the management recommendations could be reversed to limit the potential for predation.     

Assessing the Cost of an Invasive Forest Pathogen: A Case Study with Oak Wilt

Economic assessment of damage caused by invasive alien species provides useful information to consider when determining whether management programs should be established, modified, or discontinued. We estimate the baseline economic damage from an invasive alien pathogen, Ceratocystis fagacearum, a fungus that causes oak wilt, which is a significant disease of oaks (Quercus spp.) in the central United States. We focus on Anoka County, Minnesota, a 1,156 km² mostly urban county in the Minneapolis-Saint Paul metropolitan region. We develop a landscape-level model of oak wilt spread that accounts for underground and overland pathogen transmission. We predict the economic damage of tree mortality from oak wilt spread in the absence of management during the period 2007–2016. Our metric of economic damage is removal cost, which is one component of the total economic loss from tree mortality. We estimate that Anoka County has 5.92 million oak trees and 885 active oak wilt pockets covering 5.47 km² in 2007. The likelihood that landowners remove infected oaks varies by land use and ranges from 86% on developed land to 57% on forest land. Over the next decade, depending on the rates of oak wilt pocket establishment and expansion, 76–266 thousand trees will be infected with discounted removal cost of $18–60 million. Although our predictions of removal costs are substantial, they are lower bounds on the total economic loss from tree mortality because we do not estimate economic losses from reduced services and increased hazards. Our predictions suggest that there are significant economic benefits, in terms of damage reduction, from preventing new pocket establishment or slowing the radial growth of existing pockets.     

Integration of Satellite Imagery and Forest Inventory in Mapping Dominant and Associated Species at a Regional Scale

To achieve the overall objective of restoring natural environment and sustainable resource usability, each forest management practice effect needs to be predicted using a simulation model. Previous simulation efforts were typically confined to public land. Comprehensive forest management practices entail incorporating interactions between public and private land. To make inclusion of private land into management planning feasible at the regional scale, this study uses a new method of combining Forest Inventory and Analysis (FIA) data with remotely sensed forest group data to retrieve detailed species composition and age information for the Missouri Ozark Highlands. Remote sensed forest group and land form data inferred from topography were integrated to produce distinct combinations (ecotypes). Forest types and size classes were assigned to ecotypes based on their proportions in the FIA data. Then tree species and tree age determined from FIA subplots stratified by forest type and size class were assigned to pixels for the entire study area. The resulting species composition map can improve simulation model performance in that it has spatially explicit and continuous information of dominant and associated species, and tree ages that are unavailable from either satellite imagery or forest inventory data. In addition, the resulting species map revealed that public land and private land in Ozark Highlands differ in species composition and stand size. Shortleaf pine is a co-dominant species in public land, whereas it becomes a minor species in private land. Public forest is older than private forest. Both public and private forests have deviated from historical forest condition in terms of species composition. Based on possible reasons causing the deviation discussed in this study, corresponding management avenues that can assist in restoring natural environment were recommended.     

Thinning Pine Plantations to Reestablish Oak Openings Species in Northwestern Ohio

Globally the area in forest plantations is rising by 2% annually, increasing the importance of plantations for production of human goods and services and for ecological functions such as carbon storage and biodiversity conservation. Specifically in the Great Lakes states and provinces of Midwestern North America, thousands of hectares of pine plantations were established in the early and mid-1900s to revegetate abandoned agricultural fields that had replaced mixed-species forests and oak–prairie ecosystems. Plantation establishment also was intended to bolster the timber base. Management priorities have shifted, with many resource managers currently seeking to manage existing plantations for promoting mixed-species ecosystems. The purpose of this study was to assess plant succession and the reestablishment of oak savanna and prairie species after thinning 14 plantations of Pinus resinosa and strobus in northwestern Ohio, USA. Thinning reduced tree basal area by an average of 75%. Plant communities were sampled on 0.05-ha plots one and 3 years after thinning and compared to 10 unthinned control plantations. By 3 years after thinning, thinned plots contained 2–3 times more species and 14 times more plant cover than control plots. The species composition of colonizing plants was most strongly correlated with residual pine basal area and soil variables related to drainage (e.g., sand concentration, available water capacity). Although plant composition was dominated by widespread colonizers such as Erechtites hieraciifolia, the coefficient of conservatism (indicative of species of more intact, undisturbed communities) significantly increased on thinned plots from year 1 to 3. This finding, coupled with the presence of four rare, state-listed Ohio species whose eight plot occurrences all were on thinned plots, suggests that plant composition is moving towards species typifying more high-quality savanna and prairie habitats.     

Effects of recreational use on forested sites

Impact of recreational activities on soil and vegetation was evaluated in eight forested camping and picnic areas in southern Rhode Island. Forest vegetation consists of mixed-oak and white pine stands. Soils are of granitic glacial till or outwash origin and textures range from loamy sand to find sandy loam. Recreational use resulted in significant compaction of soils as indexed by soil penetration resistance and bulk density. Evidence indicates that compaction influences bulk densities to a depth of about 12.7 cm. Rates of water infiltration are less on recreation areas. Soil water accretion and depletion during the growing season are less rapid on recreation sites than on control sites. Differences are attributed to reduced infiltration, percolation, and rooting activity. Much of the ground surface on recreation areas is devoid of vegetation. The surface consists primarily of bare mineral soil, rock, or litter. The plants most commonly present are grasses. Native ground cover vegetation including tree seedlings, ericaceous shrubs and herbs has been eliminated or greatly reduced by trampling. Damage to tree trunks is common in recreation areas. White pine radial growth and scarlet oak height growth were significantly less on recreation sites. Scarlet oak appears intolerant to heavy recreation use.     

Transpiration and Root Development of Urban Trees in Structural Soil Stormwater Reservoirs

Stormwater management that relies on ecosystem processes, such as tree canopy interception and rhizosphere biology, can be difficult to achieve in built environments because urban land is costly and urban soil inhospitable to vegetation. Yet such systems offer a potentially valuable tool for achieving both sustainable urban forests and stormwater management. We evaluated tree water uptake and root distribution in a novel stormwater mitigation facility that integrates trees directly into detention reservoirs under pavement. The system relies on structural soils: highly porous engineered mixes designed to support tree root growth and pavement. To evaluate tree performance under the peculiar conditions of such a stormwater detention reservoir (i.e., periodically inundated), we grew green ash (Fraxinus pennsylvanica Marsh.) and swamp white oak (Quercus bicolor Willd.) in either CUSoil or a Carolina Stalite-based mix subjected to three simulated below-system infiltration rates for two growing seasons. Infiltration rate affected both transpiration and rooting depth. In a factorial experiment with ash, rooting depth always increased with infiltration rate for Stalite, but this relation was less consistent for CUSoil. Slow-drainage rates reduced transpiration and restricted rooting depth for both species and soils, and trunk growth was restricted for oak, which grew the most in moderate infiltration. Transpiration rates under slow infiltration were 55% (oak) and 70% (ash) of the most rapidly transpiring treatment (moderate for oak and rapid for ash). We conclude this system is feasible and provides another tool to address runoff that integrates the function of urban green spaces with other urban needs.     

Testing the Use of a Land Cover Map for Habitat Ranking in Boreal Forests

Habitat loss and modification is one of the major threats to biodiversity and the preservation of conservation values. We use the term conservation value to mean the benefit of nature or habitats for species. The importance of identifying and preserving conservation values has increased with the decline in biodiversity and the adoption of more stringent environmental legislation. In this study, conservation values were considered in the context of land-use planning and the rapidly increasing demand for more accurate methods of predicting and identifying these values. We used a k-nearest neighbor interpreted satellite (Landsat TM) image classified in 61 classes to assess sites with potential conservation values at the regional and landscape planning scale. Classification was made at the National Land Survey of Finland for main tree species, timber volume, land-use type, and soil on the basis of spectral reflectance in satellite image together with broad numerical reference data. We used the number and rarity of vascular plant species observed in the field as indicators for potential conservation values. We assumed that significant differences in the species richness, rarity, or composition of flora among the classes interpreted in the satellite image would also mean a difference in conservation values among these classes. We found significant differences in species richness among the original satellite image classes. Many of the classes examined could be distinguished by the number of plant species. Species composition also differed correspondingly. Rare species were most abundant in old spruce forests (>200 m³/ha), raising the position of such forests in the ranking of categories according to conservation values. The original satellite image classification was correct for 70% of the sites studied. We concluded that interpreted satellite data can serve as a useful source for evaluating habitat categories on the basis of plant species richness and rarity. Recategorization of original satellite image classification into such new conservation value categories is challenging because of the variation in species composition among the new categories. However, it does not represent a major problem for the purposes of early-stage land-use planning. Benefits of interpreted satellite image recategorization as a rapid conservation value assessment tool for land-use planners would be great.     

ENVIRONMENTAL AUDITING: An Approach for Characterizing Tropospheric Ozone Risk to Forests

/ The risk tropospheric ozone poses to forests in the United States is dependent on the variation in ozone exposure across the distribution of the forests in question and the various environmental and climate factors predominant in the region. All these factors have a spatial nature, and consequently an approach to characterization of ozone risk is presented that places ozone exposure-response functions for species as seedlings and model-simulated tree and stand responses in a spatial context using a geographical information systems (GIS). The GIS is used to aggregate factors considered important in a risk characterization, including: (1) estimated ozone exposures over forested regions, (2) measures of ozone effects on species' and stand growth, and (3) spatially distributed environmental, genetic, and exposure influences on species' response to ozone. The GIS-based risk characterization provides an estimation of the extent and magnitude of the potential ozone impact on forests. A preliminary risk characterization demonstrating this approach considered only the eastern United States and only the limited empirical data quantifying the effect of ozone exposures on forest tree species as seedlings. The area-weighted response of the annual seedling biomass loss formed the basis for a sensitivity ranking: sensitive-aspen and black cherry (14%-33% biomass loss over 50% of their distribution); moderately sensitive-tulip popular, loblolly pine, eastern white pine, and sugar maple (5%-13% biomass loss); insensitive-Virginia pine and red maple (0%-1% loss). In the future, the GIS-based risk characterization will include process-based model simulations of the three- to 5-year growth response of individual species as large trees with relevant environmental interactions and model simulated response of mixed stands. The interactive nature of GIS provides a tool to explore consequences of the range of climate conditions across a species' distribution, forest management practices, changing ozone precursors, regulatory control strategies, and other factors influencing the spatial distribution of ozone over time as more information becomes available.KEY WORDS: Ecological risk assessment; GIS; Ozone; Risk characterization; Forests; Trees     

Effects of Sea-Level Rise and Anthropogenic Development on Priority Bird Species Habitats in Coastal Georgia, USA

We modeled changes in area of five habitats, tidal-freshwater forest, salt marsh, maritime shrub-scrub (shrub), maritime broadleaf forest (oak) and maritime narrowleaf (pine) forest, in coastal Georgia, USA, to evaluate how simultaneous habitat loss due to predicted changes in sea level rise (SLR) and urban development will affect priority bird species of the south Atlantic coastal plain by 2100. Development rates, based on regional growth plans, were modeled at 1% and 2.5% annual urban growth, while SLR rates, based on the Intergovernmental Panel on Climate Change’s A1B mean and maximum scenarios, were modeled at 52 cm and 82 cm, respectively. SLR most greatly affected the shrub habitat with predicted losses of 35–43%. Salt marsh and tidal forest also were predicted to lose considerable area to SLR (20–45 and 23–35%, respectively), whereas oak and pine forests had lesser impact from SLR, 18–22% and 11–15%, respectively. Urban development resulted in losses of considerable pine (48–49%) and oak (53–55%) habitat with lesser loss of shrub habitat (21–24%). Under maximum SLR and urban growth, shrub habitat may lose up to 59–64% compared to as much as 62–65% pine forest and 74–75% oak forest. Conservation efforts should focus on protection of shrub habitat because of its small area relative to other terrestrial habitats and use by Painted Buntings (Passerina ciris), a Partners In Flight (PIF) extremely high priority species. Tidal forests also deserve protection because they are a likely refuge for forest species, such as Northern Parula and Acadian Flycatcher, with the decline of oak and pine forests due to urban development.     

Impingement losses of white perch at Hudson River power plants: Magnitude and biological significance

We performed a quantitative assessment of the impact of impingement at power plants on the Hudson River white perch population We estimated that impingement reduces the abundance of each white perch year class by at least 10% and probably by 15–20% or more after 2–3 years of vulnerability to power plants We attempted to detect effects of impingement on average year-class abundance of white perch from a time series of abundance indices derived from impingement data We found, however, that neither impingement collection rates observed at Hudson River power plants nor beach seine data provide a reliable index of year-class strength in white perch. Even if a reliable index were developed, natural fluctuations in year-class strength are great enough that a short-term monitoring program would be inadequate for detecting even a large reduction in average year-class strength. We performed a multipopulation analysis using simple food chain and food web models The results suggest that any long-term decline in white perch abundance caused by impingement should be accompanied by an increase in the abundance of one or more competing fish species and by an increase in the biomass of adult white perch relative to young-of-the-year.We conclude that 1) at present, assessments of population-level impact of impingement should focus on short-term effects, 2) research is needed to develop a reliable index of year-class strength for use in long-term monitoring programs, 3) identification and quantification of natural environmental factors influencing year-class strength are needed to improve our ability to predict and detect changes in abundance, and 4) it would be useful in designing monitoring programs to focus on detecting patterns of change among populations and age groups rather than solely on declines in abundance of individual populationsResearch sponsored by the Office of Nuclear Regulatory Research, US Nuclear Regulatory Commission under Interagency Agreement No. 40-550-75 with the US Department of Energy under Contract W-7405-eng-26 with Union Carbide Corporation. Publication No. 2030, Environmental Sciences Division, ORNL.     

Integrating forage,wildlife, water,and fish projections with timber projections at the regional level: A case study in southern United States

The impact of timber management and land-use change on forage production, turkey and deer abundance, red-cockaded woodpecker colonies, water yield, and trout abundance was projected as part of a policy study focusing on the southern United States. The multiresource modeling framework used in this study linked extant timber management and land-area policy models with newly developed models for forage, wildlife, fish, and water. Resource production was integrated through a commonly defined land base that could be geographically partitioned according to individual resource needs. Resources were responsive to changes in land use, particularly human-related, and timber management, particularly the harvest of older stands, and the conversion to planted pine.     

Using a GIS model to assess terrestrial salamander response to alternative forest management plans.

A GIS model predicting the spatial distribution of terrestrial salamander abundance based on topography and forest age was developed using parameters derived from the literature. The model was tested by sampling salamander abundance across the full range of site conditions used in the model. A regression of the predictions of our GIS model against these sample data showed that the model has a modest but significant ability to predict both salamander abundance and mass per unit area. The model was used to assess the impacts of alternative management plans for the Hoosier National Forest (Indiana, USA) on salamanders. These plans differed in the spatial delineation of management areas where timber harvest was permitted, and the intensity of timber harvest within those management areas. The spatial pattern of forest openings produced by alternative forest management scenarios based on these plans was projected over 150 years using a timber-harvest simulator (HARVEST). We generated a predictive map of salamander abundance for each scenario over time, and summarized each map by calculating mean salamander abundance and the mean colonization distance (average distance from map cells with low predicted abundance to those with relatively high abundance). Projected salamander abundance was affected more by harvest rate (area harvested each decade) than by the management area boundaries. The alternatives had a varying effect on the mean distance salamanders would have to travel to colonize regenerating stands. Our GIS modeling approach is an example of a spatial analytical tool that could help resource management planners to evaluate the potential ecological impact of management alternatives.     

Postfire Regeneration in <Emphasis Type="Italic">Pinus pinea</Emphasis> L. and <Emphasis Type="Italic">Pinus pinaster</Emphasis> Aiton in Andalucia (Spain)

The objective of this study was to examine postfire regeneration of tree, shrub, and dwarf shrub species, in relation to levels of damage in four planted pine forests (Pinus pinea, Pinus pinaster) in Andalusia. A prefire vegetation map was used for detailing species composition, vertical structure, and density and another for detailing the extent and intensity of fire damage. Between 3 and 7 years after the fires, an inventory was made of the vegetation in each area, using the step-point method. The information thus obtained was used to determine the amount of cover in the dwarf/shrub and tree layers, the frequency of species in each of the layers, floristic richness, and diversity (Shannon index). The botanical composition of the dwarf and shrub layer was analyzed using TWINSPAN. Variables were poorly correlated with level of fire damage, which suggests that the forests in this study followed the autosuccession model. Because of the artificial origin or seminatural condition, regeneration of the dominant tree species is poor, and it seems unlikely that forests will recover to their prefire state. Therefore action is recommended to restore these ecosystems.     

INFLUENCE OF SMALL MAMMALS ON STORMFLOW RESPONSES OF PINE-COVERED CATCHMENTS1

ABSTRACT: The relative abundance of small mammals in five forest land cover types on the upper Coastal Plain of north Mississippi was determined. Burrowing mammals accounted for one-half of the total captures; one shrew species that accounted for over one-fourth of the total captures had a strong affinity for well-stocked pine plantations. The opportunity for detention and retention of rainfall was enhanced by burrowing activity. Reductions of stormflow volumes 12 to 15 years after replacing poor quality, upland hardwoods with loblolly pine were only partially explained by increased interception of rainfall; much of the residual reductions are postulated to be due to small mammal burrows. Small mammal activity deserves further study as an important aspect of forest land hydrology.     

Vegetation Composition,Dynamics, and Management of a Bracken–Grassland and Northern–Dry Forest Ecosystem

We investigated differences in vegetation composition and dynamics for two globally rare ecosystems, bracken–grasslands and northern–dry forests of northern Wisconsin. These ecosystems commonly have been viewed as degraded pine barrens. Bracken–grasslands contained a high dominance of exotic species, low native richness, and no obvious prairie species, suggesting logging-era anthropogenic origins. Differences in cover for common plants among ecosystems were examined using Mann-Whitney U tests of equivalence. Cover of all 8 graminoid species, 4 of 5 Ericaceae and Myricaceae species, and 10 of 17 species of forbs were significantly different between ecosystems. Vegetation changes over a 4-year period were examined through detrended correspondence analysis (DCA) and analysis of variance (ANOVA) repeated measures. DCA analyses of community composition failed to detect significant temporal trends within individual management units, although differences were apparent between ecosystems, regardless of sample year. In addition, no apparent patterns could be detected between years when comparing dominant individual species to management history (prescribed fire). This is contrary to what would be expected for a degraded pine barrens and questions the efficacy of using repeated prescribed fire as a management tool in bracken–grasslands. Methods for conservation and restoration of xeric ecosystems of northern Wisconsin have historically relied heavily on single species (e.g., sharp-tailed grouse) wildlife models, without full consideration of other factors. We suggest that stakeholders involved in these restoration projects examine historic processes and reference conditions prior to formulating management goals. Greater attention to the differentiation and individual management needs of pine barrens, northern–dry forests, and bracken–grasslands is needed.     

Postfire regeneration in Pinus pinea L. and Pinus pinaster aiton in Andalucia (Spain)

The objective of this study was to examine postfire regeneration of tree, shrub, and dwarf shrub species, in relation to levels of damage in four planted pine forests (Pinus pinea, Pinus pinaster) in Andalusia. A prefire vegetation map was used for detailing species composition, vertical structure, and density and another for detailing the extent and intensity of fire damage. Between 3 and 7 years after the fires, an inventory was made of the vegetation in each area, using the step-point method. The information thus obtained was used to determine the amount of cover in the dwarf/shrub and tree layers, the frequency of species in each of the layers, floristic richness, and diversity (Shannon index). The botanical composition of the dwarf and shrub layer was analyzed using TWINSPAN. Variables were poorly correlated with level of fire damage, which suggests that the forests in this study followed the autosuccession model. Because of the artificial origin or seminatural condition, regeneration of the dominant tree species is poor, and it seems unlikely that forests will recover to their prefire state. Therefore action is recommended to restore these ecosystems.     

Assessment of frequent cutting as a plant-community management technique in power-line corridors

Repeated cutting of vegetation at or near ground level in power-line corridors is a common practice for inhibiting tree growth and regeneration. However, few data exist on long-term community responses. In this study, we sampled 20 northern Kentucky power-line corridors and compared their seedling and sapling communities to the edges and interiors of adjacent undisturbed forests. Mean seedling and sapling density in corridors was roughly twice that of adjacent undisturbed forest interiors, suggesting that repeated cutting is not a viable method of inhibiting tree regeneration. Corridor communities were dominated byRobinia pseudoacacia (black locust) andFraxinus americana (white ash), but ordinations indicated strong similaritties among communities in corridors and adjacent forests. Many of the tree species found in adjacent forests, with the exception of a few shade-tolerant species, had highest seedling and sapling densities in corridors. Stump or root sprouting by many species appears to regenerate forests quickly after cutting. However, disturbed soil and detritus accumulations caused by management crews and their equipment may also create a large variety of microsites for seedling establishment. Because repeated cutting selects for dominance by species with highest sprout growth rates, it should not be used as the sole management technique. It may instead be used to alter the vigor, stature, and stored reserves of trees so that herbicides or other methods of tree control can be used more efficiently.     

Differences in bioregional classifications among four aquatic biotic groups: Implications for conservation reserve design and monitoring programs

Bioregional classifications are used extensively for conservation management and monitoring programs. This study used generalised dissimilarity modelling (GDM) to test the ability of different regional classifications of four groups of aquatic biota to be used as surrogates for each other. Classifications were derived for aquatic macrophytes, macroinvertebrates, freshwater fish and frogs using community-level modelling, or GDM, which relates the biotic assemblage structure with environmental variables. Six regions were defined for each biotic group for the State of New South Wales. Regional classifications differed markedly between the different biotic groups because the environmental drivers that were related to species turnover throughout the region differed among groups. Altitude and rainfall were the strongest drivers of species turnover among the groups. Results suggest that physiographic variables should be incorporated in reserve design and monitoring programs to explicitly address differences in classifications between similar biotic groups.     

Prolonged herbicide-induced vegetation changes in a regenerating boreal aspen clearcut

A soil-active herbicide (hexazinone) was applied (0, 2, and 4 kg/ha of active ingredient) in a 3-year-old regenerating boreal Populus tremuloides Michx. (aspen) clearcut to determine its effect on the compositional and structural development of the vegetation. Woody stem densities and plant foliar cover were evaluated prior to and 2, 6, and 17 years after treatment. Herbicide treatment at the 2 and 4 kg/ha rates reduced tree and total woody stem densities relative to the 0 kg/ha level. The 4 kg/ha level reduced stem densities by 27% 17 years after treatment. The primary reductions occurred in Amelanchier alnifolia (Nutt.) Nutt. ex M. Roemer (saskatoon) and Rosa acicularis Lindl. (wild rose); whereas Corylus cornuta Marsh. (beaked hazelnut) and Viburnum edule (Michx.) Raf. (low-bush cranberry) stem densities increased. Notable herbicide-caused foliar cover reductions at the 4 kg/ha level occurred in Eurybia conspicua (Lindl.) Nesom. (showy aster), Mertensia paniculata (Ait.) G. Don. (tall mertensia), Rubus pubescens Raf. (dewberry), and Spiraea betulifolia Pallas (spiraea), but Aralia nudicaulis L. (sarsaparilla), Cornus canadensis L. (bunchberry), and Symphyotrichum ciliolatum (Lindl.) A.&D. Lve (Lindley's aster) increased. Less distinctive but similar changes occurred in the 2 kg/ha treatment. Total plant cover, species richness, and species dominance concentration were similar among treatments. Eight distinctive forest understory-types were recognized among treatments in Year 17. Between the 0 and 4 kg/ha treatments, five understory-types differed in their frequency of occurrence. Hexazinone did not improve the survival of silviculturally planted Picea glauca (Moench) Voss (white spruce) seedlings relative to untreated sites, but the 4 kg/ha treatment level did increase Pinus contorta Dougl. ex Loud. (lodgepole pine) survival from 12 to 34%. Surviving seedlings had significantly greater height and basal diameter growth than those at the 0 kg/ha sites, particularly the 4 kg/ha treatment.     

Woodland restoration in Scotland: ecology, history, culture, economics, politics and change

In the latter half of the 20th century, native pine woodlands in Scotland were restricted to small remnant areas within which there was little regeneration. These woodlands are important from a conservation perspective and are habitat for numerous species of conservation concern. Recent developments have seen a large increase in interest in woodland restoration and a dramatic increase in regeneration and woodland spread. The proximate factor enabling this regeneration is a reduction in grazing pressure from sheep and, particularly, deer. However, this has only been possible as a result of a complex interplay between ecological, political and socio-economic factors. We are currently seeing the decline of land management practices instituted 150-200 years ago, changes in land ownership patterns, cultural revival, and changes in societal perceptions of the Scottish landscape. These all feed into the current move to return large areas of the Scottish Highlands to tree cover. I emphasize the need to consider restoration in a multidisciplinary framework which accounts not just for the ecology involved but also the historical and cultural context.