河东沙区侧柏树干液流与蒸腾驱动因子的时滞效应研究

树干液流传导反映树木蒸腾过程是估算林木耗水量的重要基础,忽视液流的时滞现象则会造成显著误差。以宁夏河东沙区防护林树种——侧柏(Platycladus orientalis)为研究对象,利用热扩散茎流计监测林木生长旺季的树干液流(J_s)日变化格局,采用时间错位对比法分析侧柏树干液流与蒸腾驱动因子之间的时滞效应,旨在为准确理解液流时滞和冠层蒸腾的关系,提高预测森林蒸腾耗水的准确度提供依据。结果表明:典型晴天下,侧柏树干液流与蒸腾驱动因子太阳辐射(E_s)、饱和水汽压亏缺(D)、大气温度(θ_a)均呈现"迟滞回环"的关系,液流密度J_s对D、θ_a的响应回环过程呈顺时针方向,而J_s与E_s间响应回环过程则为逆时针。采用高斯方程拟合E_s、D、θ_a、潜在蒸散(E_p)及J_s日变化过程,结果表明Js峰值时刻明显滞后于E_s和E_p,提前于θ_a和D;E_s与E_p日变化过程和峰值时刻同步,但明显早于θ_a和D。错位对比法分析得出,典型晴天下侧柏树干液流与主要驱动因子E_s、D、θ_a的实际时滞分别为0.75、-0.75和-0.25 h。侧柏树干液流与E_s之间的时滞ΔJ_s-E_s与日间耗水量(W_d)和日平均水汽压饱和亏缺(D)显著相关;与D之间的时滞ΔJ_s-D与W_d、日间太阳辐射总量(E_(sT))、D显著相关,且对侧柏树干液流时滞具有促进作用。     

温带落叶林的植物物候特征及其对气候变化的响应

植被物候是气候变化对生物圈产生长期或短期影响的重要指示因子。气候变化已经明显改变了许多物种的营养生长和繁殖物候,尤其是在温带地区。研究温带森林物候变化及其对全球变暖的响应,对认识森林物种共存,协同进化以及森林保护和经营等有重要意义。通过概述温带森林下物候研究的进展发现,光照和积温是影响木本植物展叶及繁殖物候的关键因素,林下层树木通过更早展叶,以尽量减少生长季林冠层遮阴对下层树木生长的影响,更早时期开花的树木具有从顶部向四周次第开花的时空格局,林冠层树种开花具有较好的同步性。而草本植物的物候通常受融雪时间和冠层动态的影响更大,并且,温带森林下不同生活史对策的草本植物的物候特征对气候变化的响应也不尽相同,存在明显的季节动态。繁殖物候、光照的季节变化、光合特征、授粉成功之间的联系决定了林下不同繁殖特性的草本植物的繁殖成功率。量化的、多指标、多对象的定位监测是精准物候研究的基础,物候变化的机理和建立可预测的物候模型将是未来研究的重点。     

两种园林植物断根和剪枝后蒸腾和光合特性的变化研究

选择移栽难易程度差异很大的两种园林树木(小叶榕Ficusmicrocarpa和拟单性木兰parakmeriaomeiensis)进行断根和不同程度的剪枝,模拟移栽,形成不同的成活水平个体,比较不同成活水平下蒸腾、光合等生理机能的差异,以期从生理生态角度了解影响成活的关键因子。结果表明,①小叶榕相对拟单性木兰移栽成活率高,主要是处理后,小叶榕气孔导度能很快降低,导致蒸腾速率降低,耗水量显著减小;而拟单性木兰处理后气孔导度和蒸腾速率都升高,主要依靠剪枝来降低植株蒸腾耗水,降低幅度不大。小叶榕日平均蒸腾速率显著高于拟单性木兰,两者日平均耗水量都随剪枝程度的增加而减少,即重度剪枝最容易成活。②拟单性木兰净光合速率与气孔导度都显著相关,而小叶榕净光合速率与气孔导度的相关性较差。小叶榕日平均光合速率显著高于拟单性木兰,种或处理间日变化趋势也都存在差异。③小叶榕处理后水分利用效率变化趋势不同于对照,但处理之间日变化趋势类似,随处理的增加日平均值降低;拟单性木兰处理以及对照间变化趋势有很大差异,但日平均值相差不大,都显著高于对照。仍需进一步研究内部机理,找到解决成活的更有效措施。     

森林改善空气环境质量功能监测与评价研究

为全面了解森林内部属性与森林改善空气环境功能之间的内在关系与规律,以及不同森林在改善空气环境质量方面的能力强弱,以中国林业科学研究院热带林业试验中心为研究对象,于2014年7─8月按照2 km×2 km网格间距进行系统抽样,设置51个样地对森林空气温度、湿度、负氧离子含量等空气环境状况指标及林分特征因子进行了监测,并对林分类型、发展阶段、郁闭度等林分因子与森林空气温度、湿度、负氧离子含量之间的关系进行了分析。采用温湿指数、安培空气质量评价系数、森林空气负氧离子评价模型对不同类型森林的空气环境质量进行了评价。结果表明：1）森林改善空气环境质量功能主要与其林分类型、发展阶段、林分郁闭度有关;2）在调节空气温湿度,改善空气环境舒适度方面,针叶混交林最强,然后依次为针阔混交林、阔叶混交林、针叶纯林、阔叶纯林、灌木林,以未成林最弱。在吸附污染物、净化空气方面,针阔混交林最强,然后依次为阔叶混交林、阔叶纯林、针叶混交林、针叶纯林、灌木林,以未成林最弱;3）不同发展阶段的林分在改善空气环境舒适度方面的差异不明显,但随着林分发展阶段的深入,森林的结构越来越复杂,森林净化空气的能力也逐渐增强;4）森林在调节空气温湿度,改善空气环境舒适度等方面的功能随郁闭度的增加而增强,当郁闭度大于0.4时,森林改善空气环境舒适度的功能显著增强。郁闭度在0.4~0.7之间的林分净化空气的功能较强,尤以郁闭度为0.5时最强,当林分郁闭度低于0.4或高于0.7以后,林分净化空气的功能均有所降低;5）总体来说混交林改善空气环境质量的功能要显著强于纯林。研究结果对于经营单位如何通过林分状态的调控来增强森林的空气环境改善功能具有重要指导意义。     

土壤水分对幼龄桉树蒸腾和生长的影响

通过盆栽试验,研究不同土壤水分对幼龄桉树蒸腾和生长的影响.结果表明,土壤含水量越高,桉树蒸腾量越大.随着桉树增长,植株蒸腾量逐月增大,但蒸腾强度却逐月下降,说明桉树叶面积增长速度,大于蒸腾量增长速度.不同土壤水分条件下桉树的蒸腾量、蒸腾时间长短和最大蒸腾量出现的时间不同.水分亏缺对桉树的影响是多方面的,外观表现为幼嫩枝梢萎蔫,分枝少、叶片少、叶面积下降、叶色变深、变暗,植株下部叶子叶尖枯死;生理上表现为气孔导度降低,阻力增大,气孔关闭,光合速率降低,养分吸收量减少.总的影响是生长量显著降低.在试验条件下,桉树每生产1 kg干物质所付出的蒸腾耗水量与土壤水分有关,土壤干旱时桉树的水分利用效率最低.     

菌根类型对森林树木净初级生产力的影响

菌根是土壤真菌与植物根系形成的共生体,存在于绝大多数植物（90%）的根系和生境中。菌根共有7种类型,在生态系统的过程和功能方面都扮演着十分重要的角色。为了增强对菌根在森林生态系统中重要功能的理解,文章基于全球森林数据库,在全球尺度上研究了不同菌根类型对森林树木净初级生产力（NPP）的影响。结果表明,森林树木NPP随菌根类型的不同而不同,AM类型菌根森林的NPP[679.49 g.m-2.a-1（以C计）]要显著高于含ECM类型菌根的森林[479.00 g.m-2.a-1（以C计）];菌根类型的不同对森林树木地上和地下及其各组分NPP的影响和贡献也存在着显著的不同,AM类型菌根对地下NPP的贡献要高于ECM菌根,而ECM菌根对地上NPP的贡献则较大。菌根类型对地上、地下NPP组分的影响分析则表明,AM类型的菌根对树叶和细根NPP的贡献较大,而ECM类型菌根则对树木主干和枝NPP的贡献较大。可见,森林树木总体NPP及其各组分NPP都随着菌根类型的不同而存在显著的差异。     

长白山阔叶红松林林分空间结构

在长白山北坡自然保护区内,建立面积为1hm~2的阔叶红松林永久标准样地,对胸径1cm以上的每株树木进行定位监测,用空间结构参数——混交度、大小比数和角尺度分析该林分的空间结构.结果表明:该林分树木个体总体上处于中度混交水平,处于乔木上层和中层的树种混交度以强度混交和完全混交为主,明显高于乔木下层树种的混交度;基于胸径计算的大小比数和基于树高计算的大小比数的结果基本一致,都能客观反映该林分在垂直方向上的分化;树木个体整体分布格局与树木大小密切相关,当树木个体起测径阶1cm≤DBH6cm时,林分呈聚集分布的格局,当起测径阶DBH6cm时,林分呈随机分布的格局.图1表4参25     

东北北部温带森林和干草地土壤养分分布及影响因素

土壤是陆地生态系统碳储存的重要场所,其养分变化与全球陆地碳循环密切相关。土壤养分是植物生长的重要保证,而土壤各养分之间是紧密联系的。理解土壤养分变化与环境因素的关系有助于更好地了解陆地生态系统碳、氮、磷循环。本研究以东北北部自东向西沿降水量梯度变化纬度带上的温带森林与干草地生态系统为研究对象,利用气象数据和野外土壤实测数据,分析了纬度带上不同植被类型土壤的有机碳、全氮、碳氮比、速效磷的空间分布格局及其与环境因子（年降水量、年均温、土壤pH值）的关系。研究纬度带上降水量自东向西逐渐减少,植被类型从温带森林过渡到干草原,与降水量和植被类型对应,植被生物量也自东向西呈现从高到低的分布梯度。研究结果表明：从整个研究带上来说,降水量与土壤pH值是土壤养分空间分布的决定因素,沿纬度带从东到西,随着降水量逐渐减少,土壤pH值逐渐增加,而土壤有机碳、全氮、碳氮比、速效磷含量逐渐减少。但如果将森林和草地分别讨论则发现,森林和草地生态系统的土壤养分环境控制因素有较大差别。对于草地生态系统而言,降水量和土壤pH值仍然是其土壤养分含量的控制因子,但森林生态系统由于所处区域降水量充足,降水量不再是其土壤养分的控制因子,降水量只与森林土壤碳氮比呈显著正相关。研究还发现森林土壤的速效磷含量与温度呈正相关,与土壤pH值呈负相关,说明温度对东北北部温带森林的土壤养分含量具有一定的控制作用。     

亚热带与温带森林小流域生态系统汞的生物地球化学循环及其同位素分馏

汞是通过大气进行长距离传输的全球污染物,引起国际社会和学术界高度关注.陆地森林生态系统是全球物质循环最为活跃的地方,但其对全球汞的生物地球化学循环影响的认识还不清楚.同时,森林生态系统汞的生物地球化学循环过程可能对全球大气汞同位素组成产生重要的影响,但目前这两方面的研究还非常缺乏,制约了对全球尺度汞的生物地球化学循环深化规律的把握.本课题组拟在我国温带和亚热带选择3个森林小流域,首先系统开展森林小流域汞的质量平衡研究工作,深入刻画森林流域汞的生物地球化学演化规律,在此基础上开展流域汞的生物地球化学循环过程中汞同位素的分馏特征研究,最终建立流域尺度森林系统汞及其同位素的生物地球化学模型.该项目将极大推进对森林流域尺度汞的生物地球化学循环的认识,探讨森林生态系统与大气汞交换过程对全球尺度汞生物地球化学循环和对大气汞同位素组成的影响,为最终建立基于汞同位素的全球汞生物地球化学循环模型提供基础数据.     

林分和气候因子对中国天然林根冠比的影响

森林生态系统是陆地生态系统最重要的碳库和碳汇,其地下-地上生物量分配规律即根冠比(R/S),对于精确估算森林生物量,尤其是对建造森林地下生物量估算模型具有重大参考意义.为探究大尺度上气候、林分因子对中国天然林根冠比的影响,对1 109块天然林样地进行统计分析.结果显示:中国天然林R/S平均值为0.26,阔叶森林的R/S要显著高于针叶森林,落叶森林的R/S要显著高于常绿森林,年均温、林龄及水分可利用性与中国天然林R/S呈显著负相关关系,年均温是影响中国天然林R/S变化的最主要因子.综合纬度、经度、年均温、降水、林龄等环境因素的影响发现,纬度、林分密度是影响R/S的最主要因素.因此,精确估算森林碳储量和森林生物量分配时必须考虑纬度和林分密度的影响.     

气温变化对西南山区流域森林水文效应影响的模拟

为了揭示气温变化对西南山区流域森林水文效应的影响,用生物物理/动态植被模型SSiB4/TRIFFID与流域地形指数水文模型TOPMODEL的耦合模型SSiB4T/TRIFFID模拟了西南山区长江上游梭磨河流域森林水文效应对气温变化的响应,分析了气温变化对植被不同演替阶段的流域总径流和总蒸发以及冠层截流蒸发、植被蒸腾和土壤蒸发的影响。结果表明,（1）梭磨河流域森林（常绿针叶林）蒸腾与草和灌木差异小,森林蒸腾潜热比草和灌木仅高1~4 W.m-2,森林冠层截留蒸发高于草和灌木,但土壤蒸发明显低于草和灌木覆盖,森林覆盖流域总蒸发低于草和灌木覆盖甚至低于裸土蒸发,因此增加了流域总径流量,但森林增加径流的作用随土壤蒸发的减小而减小。（2）气温减小1℃将通过减小森林冠层截留蒸发和蒸腾而使森林增加流域总径流量的作用增加;相反,气温增加将增加森林冠层截留蒸发和蒸腾而使森林增加总径流量的作用减小。（3）当温度增加4℃,由于森林总蒸发较草和灌木明显增加,对于较高的土壤蒸发,森林增加总径流量的作用已不明显;对于较低的土壤蒸发,森林减小了流域总径流量。     

岷江上游林草交错带祁连山圆柏群落的物种多样性及乔木种群的分布格局

应用5种多样性指数对祁连山圆柏群落的物种多样性进行了研究，并采用相邻格子样方法测定了乔木种群及其各立木级的分布格局．结果表明，灌木层的物种多样性偏低，而草本层多样性较高，群落总的物种多样性(以丰富度表示)较高．群落总的物种多样性较高是由于较低的灌木层多样性与较高的草本层多样性综合作用的结果，与放牧影响、阳坡的干燥环境有关，可能是林草交错区固有的特点．乔木种群的分布格局呈现出集群分布，但在其发育过程中显示了动态变化的特征，即在幼苗和小树阶段为集群分布，到了大树阶段转变为随机分布或均匀分布．表6参32     

The distribution of lianas and their change in abundance in temperate forests over the past 45 years

Lianas (woody vines) are an important and dynamic component of many forests throughout the world, and increases in CO2, mean winter temperature, and forest fragmentation may promote their growth and proliferation in temperate forests. In this study, we used a 45-year data set to test the hypothesis that lianas have increased in abundance and basal area in the interiors of 14 deciduous temperate forests in Wisconsin (USA) since 1959. We also censused woody plants along a gradient from the forest edge to the interior in seven of these forests to test the hypothesis that the abundance of lianas declines significantly with increasing distance from the forest edge. We found that lianas did not increase in abundance within the interiors of temperate forests in Wisconsin over the last 45 years. However, relative and absolute liana abundance decreased sharply with increasing distance from forest edges. Our findings suggest that forest fragmentation, not climate change, may be increasing the abundance of lianas in northern deciduous temperate forests, and that lianas may further increase in abundance if the severity of forest fragmentation intensifies.     

The role of traditional institutions in the conservation of forest resources in East Usambara,Tanzania

The study investigated the role of traditional institutions in the conservation of the East Usambara forests in Tanzania. It revealed that there had been institutional changes in management of forests, which had a negative impact on forest conservation. However, the presence of traditional institutions, which included among others traditional healers, traditional leaders, traditional taboos, sacred species and sites, had active roles in conserving the forest. More than 90% of the respondents had retained indigenous tree species on their farms and around households, with an average of six trees per household. This strategy is influenced by socio-economic factors whereby respondent age had a significant positive influence on the number of trees retained by a household. The study concludes by pointing out that traditional institutions have active roles in conserving the forests, and recommends that various strategies must be combined, including the use of traditional institutions, in Conservation of forests. Studying tradition institutions will give us knowledge on how they operate and how they can be combined with other strategies in government, NGOs and even the private sector in conservation of the forests. Socio-economic factors must be taken into account in the management of natural resources.     

A mechanistic model of tree competition and facilitation for Mediterranean forests: Scaling from leaf physiology to stand dynamics

Mechanistic theories of plant competition developed to explain changes in community structure and dynamics along resource availability gradients have been mostly applied to temperate forests and grasslands where light and nutrients are the two main limiting resources. In contrast, the mechanisms underlying the structure and dynamics of water-limited plant communities have been little explored. Also previous mechanistic models rely either on complex simulators, which are difficult to interpret or on simple conceptual models, which ignore too many critical details. In this study, we develop a model of stand dynamics for light and water-limited forests of intermediate complexity and we provide an analytical framework for its analyses. The model is an individual-based simulator that describes the feedback between transpiration, stomatal function and soil water dynamics with asymmetrical competition for light and water. Trees allocate carbon to three main compartments: shoot, stem and roots. We use the model to explore general patterns that may emerge across levels of biological organization from the leaf to the stand. Model predictions are consistent with a number of features of Mediterranean forests structure and dynamics. At the plant-level the leaf-based tradeoff between carbon gain and water loss expresses as a tradeoff between mortality and growth. This tradeoff explains plant morphological changes in above-ground biomass and root to shoot allocation along a water availability gradient. At the community-level, tradeoffs among carbon acquisition and water loss govern the sign of plant interactions along the gradient. Coexistence among morphological types was not observed for the range of parameters and environmental conditions explored. Overall the model provides an unifying explanation for the observed changes in the sign of plant to plant interactions along environmental gradients as well as a process-based formulation that can be linked to empirical studies.     

森林生态系统的水文调节功能及生态学机制研究进展

森林水文调节功能是森林所实现的重要服务功能之一,可是由于森林资源被无节制的开采利用,导致人们不断遭受森林破环所带来的洪旱灾害。因此关于森林生态水文功能研究已成为生态学和水文学的研究重点之一。近年来,国内外对森林水文调节过程及其生态学机制进行了广泛深入的研究,所以文章从森林的水文过程出发,对林冠截留、树干流、凋落物层截留、林地水分涵养和蒸发蒸腾及其影响因子的国内外研究现状进行了归纳分析,研究认为林地各冠层均能够截留降雨,降低雨水动能,从而减少地表径流的产生和对地表的冲击;凋落物层能蓄留水分、抑制蒸散、减缓地表径流;而树干流改变降雨水平空间格局,影响水分入渗以及土壤水源涵养。森林结构复杂,明显改变了降雨分配过程,而森林水文过程及调节功能又受到森林结构的制约,因此定量定性探讨森林生态系统的结构、过程与水文调节功能之间关系,是未来森林生态水文功能研究的重点。     

Managing Stand Structure as Part of Ecologically Sustainable Forest Management in Australian Mountain Ash Forests

Ecologically sustainable management of temperate forests is a complex task that involves balancing potentially conflicting land uses such as wood production and nature conservation. We argue that a variety of strategies implemented at different spatial scales is required for biodiversity conservation in temperate forests where wood production is permitted. This is a form of "risk-spreading;" if one option is ineffective for a given species, it may still be conserved as a result of the implementation of other approaches. At the largest scale, there is a clear need for reserves to protect representative samples of forest ecosystems. Within landscapes broadly designated for timber harvesting, intermediate-scale strategies such as the implementation of networks of streamside reserves and wildlife corridors are important for biodiversity conservation. At smaller spatial scales within harvested areas, critical habitat components for forest-dependent organisms like large old trees and logs must be provided. We focus on the importance of these fine-scale attributes for the conservation of biodiversity within logged forests using the mountain ash (  Eucalyptus regnans) forests of the Central Highlands of Victoria as a case study. Forest managers must develop silvicultural practices that maintain and perpetuate critical stand attributes essential for the conservation of forest-dependent organisms. To this end, a shift is required from the extensive use of clearfelling to the adoption of new silvicultural techniques that maintain more structurally complex multi-aged stands. The maintenance of key structural features should be used as a template to guide harvesting methods to ensure that production forests contribute to biodiversity conservation, not only in mountain ash forests, but also temperate wood production elsewhere around the world.     

The Potential for Species Conservation in Tropical Secondary Forests

Abstract: In the wake of widespread loss of old‐growth forests throughout the tropics, secondary forests will likely play a growing role in the conservation of forest biodiversity. We considered a complex hierarchy of factors that interact in space and time to determine the conservation potential of tropical secondary forests. Beyond the characteristics of local forest patches, spatial and temporal landscape dynamics influence the establishment, species composition, and persistence of secondary forests. Prospects for conservation of old‐growth species in secondary forests are maximized in regions where the ratio of secondary to old‐growth forest area is relatively low, older secondary forests have persisted, anthropogenic disturbance after abandonment is relatively low, seed‐dispersing fauna are present, and old‐growth forests are close to abandoned sites. The conservation value of a secondary forest is expected to increase over time, as species arriving from remaining old‐growth forest patches accumulate. Many studies are poorly replicated, which limits robust assessments of the number and abundance of old‐growth species present in secondary forests. Older secondary forests are not often studied and few long‐term studies are conducted in secondary forests. Available data indicate that both old‐growth and second‐growth forests are important to the persistence of forest species in tropical, human‐modified landscapes.     

Conservation of Epiphyte Diversity in an Andean Landscape Transformed by Human Land Use

Abstract: Epiphytes are diverse and important elements of tropical forests, but as canopy‐dwelling organisms, they are highly vulnerable to deforestation. To assess the effect of deforestation on epiphyte diversity and the potential for epiphyte conservation in anthropogenically transformed habitats, we surveyed the epiphytic vegetation of an Ecuadorian cloud forest reserve and its surroundings. Our study was located on the western slopes of the Andes, a global center of biodiversity. We sampled vascular epiphytes of 110 study plots in a continuous primary forest; 14 primary forest fragments; isolated remnant trees in young, middle‐aged, and old pastures; and young and old secondary forests. It is the first study to include all relevant types of habitat transformation at a single study site and to compare epiphyte diversity at different temporal stages of fragmentation. Epiphyte diversity was highest in continuous primary forest, followed by forest fragments and isolated remnant trees, and lowest in young secondary forests. Spatial parameters of habitat transformation, such as fragment area, distance to the continuous primary forest, or distance to the forest edge from inside the forest, had no significant effect on epiphyte diversity. Hence, the influence of dispersal limitations appeared to be negligible or appeared to operate only over very short distances, whereas microclimatic edge effects acted only in the case of completely isolated trees, but not in larger forest fragments. Epiphyte diversity increased considerably with age of secondary forests, but species assemblages on isolated remnant trees were impoverished distinctly with time since isolation. Thus, isolated trees may serve for recolonization of secondary forests, but only for a relatively short time. We therefore suggest that the conservation of even small patches of primary forest within agricultural landscape matrices is essential for the long‐term maintenance of the high epiphyte diversity in tropical cloud forests.