Structure and formation of conifer stands in the upper timberline ecotone on the North Chuya Ridge,Central Altai

The structure of Siberian stone pine (Pinus sibirica Du Tour.) and Siberian larch (Larix sibirica Ledeb.) stands and specific features of their formation have been studies in the forest-tundra ecotone on the North Chuya Ridge (2235–2475 m a.s.l.). Changes in the structure of these stands along the transition from the upper boundary of closed forests to the high-mountain tundra have proved to have an ambiguous pattern. Both tree species form mixed clusters of similar-aged trees in the lower part of the ecotone but grow singly, in scatters, in its upper part. The formation of conifer stands (tree clusters) in the lower part of the ecotone, on the slopes of the Aktru River valley, began during climate warming in the second half of the 19th century. The expansion of confers to its upper part took place markedly later, in the early 20th century (Siberian larch) or even in the 1930s (Siberian stone pine).     

Tree vegetation of the forest-tundra ecotone in the Western Sayan mountains and climatic trends

Parameters of reproduction of the Siberian stone pine (Pinus sibirica), including radial and apical tree increments, the age structure of stands, the amount of young growth, and its distribution along an altitudinal gradient, have been studied in the forest-tundra ecotone of the Western Sayan. The results show that, over the past 30 years, P. sibirica undergrowth has expanded to the mountain tundra belt, the apical and radial tree increments and stand density have increased, and the life form of many P. sibirica plants has changed from prostrate to erect (single-or multistemmed). These changes correlate with the dynamics of summer temperatures and monthly (in May and June) and annual precipitation. The rise of summer temperatures by 1°C promotes the expansion of P. sibirica undergrowth for approximately 150 m up the altitudinal gradient.     

Climate changes and overstory recruitment of larch at the northern timberline in the Yamal Peninsula

Studies on overstory recruitment of the Siberian larch (Larix sibirica Ledeb.) in open larch forests depending on the time of seedling establishment (1880–1980) have been performed in the northern timberline ecotone on the Yamal Peninsula. Two periods of active larch establishment have been revealed: from 1900 to 1935 and from 1950 to 1970. A comparison of these data with the results of instrumental observations on temperature and precipitation has shown that the overstory recruitment of larch in valleys of northern rivers is largely dependent on July air temperatures in the first years of tree life, when seedlings grow 10–20 cm high.     

Climate changes and tree stand dynamics at the upper limit of their growth in the North Ural mountains

The composition and structure of tree stands near the timberline have been studied on different slopes and at different elevations in the Tylaisko-Konzhakovsko-Serebryanskii Massif, the North Urals. It has been found that the upper limits of tree stands with different degrees of canopy closure have risen considerably (by about 100 m of elevation) since the mid-19th century, although the formation of these stands started as early as the late 18th century. Woodless areas in the eastern part of the massif started to be colonized by Larix sibirica in the late 18th to early 19th centuries; those in the western part, by Picea obovata in the mid-19th century; and in the southern part, by Betula tortuosa in the late 19th century. Analysis of meteorological data provides evidence for warming and increasing humidity of the climate since the late 19th century. Favorable climatic changes that facilitated the expansion of the forest have taken place both in the summer (prolongation of the growing period) and in winter seasons (increase of air temperature and precipitation). The observed differences in the composition and dynamics of tree stands between the studied areas of the mountain range are most probably explained by different requirements of tree species for the depth of snow cover and the degree of soil freezing.     

Dynamics of the timberline in high mountain areas of the nether-polar Urals under the influence of current climate change

Changes in the altitudinal position of the timberline in high mountain areas of the Nether-Polar Urals and basic factors that influence such changes have been revealed on the basis of comparison of the age structure of Siberian larch (Larix sibirica) and arctic birch (Betula tortuosa) tree stands and photographs made in different years. On the mountain slopes studied, an upward shift of the timberline took place in areas covered in winter with thick snow (in the late 18th century), with Siberian larch being the pioneer species. Larch began colonizing areas with a thin snow cover in the 20th century. Birch appeared later and has since strengthened its positions. The increase in winter temperatures and precipitation facilitated the expansion of the forest.     

Changes in the upper limits of tree species distribution in the Western Caucasus (Belaya River basin) related to recent climate warming

To estimate the response of tree vegetation to recent climate warming in the Western Caucasus, the population state of several tree species at the upper limit of their distribution has been estimated in three mountain massifs in the Belaya River basin. For this purpose, the average and maximum stem diameters, vitality, and age of trees have been determined in the transitional zone between middle-mountain and high-mountain beech-fir forests (1400–1700 m a.s.l.) and in crooked birch forests growing at the upper timberline (1810–2025 m). The upper limits of Acer platanoides, A. pseudoplatanus, Ulmus glabra, and Betula litwinowii have shown a tendency to rise. The upper limit of Abies nordmanniana has remained unchanged over the past few decades, which may be explained by a decrease in moisture supply during the warm season.     

Analysis of consortive relationships between the Siberian stone pine and the nutcracker in the Northern Urals

The data presented have been obtained in the course of long-term stationary quantitative studies on the annual seed reproduction dynamics of Siberian stone pine (Pinus sibirica Du Tour) stands, including the abundance of seedlings, and the local population of the nutcracker (Nucifraga caryocatactes macrorhynchos C.L. Brehm) in forests of the Northern Urals. Statistically significant consortive relationships of the nutcracker population density with the yield of Siberian stone pine seeds and of the abundance of pine seedlings with the yield of seeds and the abundance of nutcrackers over preceding years have been hrevealed for the first time and formalized mathematically.     

Altitudinal and horizontal shifts of the upper boundaries of open and closed forests in the Polar Urals in the 20th century

In the Polar Urals (the Rai-Iz massif and Mounts Tchernaya and Malaya Tchernaya), altitudinal and horizontal shifts of the upper boundary of open and closed larch forests in the 20th century have been studied. Spatiotemporal parameters of these shifts have been assessed with the aid of the ARC/INFO geographic information system (ESRI Inc., United States), using our original large-scale geobotanical maps showing the distribution of different types of forest-tundra communities in the early 1910s and 2000s. The results show that tree vegetation has been actively expanding to higher elevations over the past 90 years. On average, the upper boundaries of open and closed forests have ascended 26 and 35 m and shifted horizontally 290 and 520 m, respectively. These shifts have been conditioned by climate warming and increasing humidity observed since the 1920s.     

Models of taiga-steppe communities on the western coast of Lake Baikal

As a result of monitoring spatial changes in the cenotic structure of plant communities at the taiga-extrazonal steppe boundary on the western coast of Lake Baikal, models of taiga-steppe communities reflecting the formation, dynamics, and genesis of forests in the taiga zone have been constructed. Changes in the species composition of communities, mesophytization of steppes, the expansion of moss synusiae characteristic of polydominant dark conifer and light conifer taiga forests, active forest invasion into steppe areas, and the presence of dark conifer tree species in light conifer forests have been revealed.     

Ecological Consequences of Transient Outbreaks of Siberian Larch Casebearer,Coleophora sibiricella Flkv., in the Udmurt Republic

A transient population outbreak of the Siberian larch casebearer, Coleophora sibiricella Flkv. (Lepidoptera, Coleophoridae), in a protective forest strip and its consequences for the host tree species (Larix sibirica Ledeb.) have been studied for the first time. It has been shown that one-time complete defoliation by this pest and the following infestation by xylophagous insects result in tree death, while the productivity of surviving trees decreases significantly. The results of this study indicate that the Siberian larch casebearer is a major phyllophagous pest of larch and that its populations must be monitored by specialists in forest pathology.     

Communities of trees along a tropical forest restoration gradient

Increasing rates of deforestation in tropical forests have been linked to agriculturalists. A critical concern generating debate is how well communities of trees recover into a more species rich ecosystem after restoration planting. The aim of the study was to evaluate the pattern of recovery of communities of tree, assess the influence of Acanthus pubescens, Lantana camara and Pennisetum purpureum, on the recovery as well as how restoration planting facilitates recruitment of other native tree seedlings along a gradient of forest restoration in Kibale National Park, Uganda after evictions of illegal settlers. We studied six restoration forests ranging in age from 3 to 16 years, naturally regenerating and three primary forests. Our results showed that recovery with natural regeneration was more effective than restoration planting although the latter enhanced recruitment of other native tree seedling. Tree recovery was generally correlated with age so that species density and diversity increased although at different rates. A reverse pattern was found for dominance but no clear pattern was found for tree density (individual/ha). Communities of tree showed directional patterns of change however community composition were still distinct among the different forests. A. pubescens, L. camara and P. purpureum negatively correlated with species density, tree density and diversity but a positive correlation was found for dominance. Restoration planting can reestablish forests with high species density, tree density and diversity, but this is dependent on age and the extent of the herbs, grasses and shrubs cover in tropical forests.     

Dynamics of growth and seed production in the Siberian stone pine: The level and pattern of variation in characters

A statistical analysis of interannual variation in a set of vegetative and generative characters over 13 years has been performed to characterize the growth and seed production dynamics of the Siberian stone pine (Pinus sibirica Du Tour) in the southeast of the forest zone in western Siberia. The results have shown that the range of fluctuations in most of vegetative characters is ordinary and their distribution is close to normal. The range of fluctuations in many generative characters is enormous (from 0 to +∞), and the distribution of their values is usually skewed, with low values being recorded several times more frequently than high values. Most variable are the proportion of abortive cones and other characters that are determined mainly in the spring of the pollination year. These characters account for a very high level of variation in the total seed production.     

Diet structure of the Siberian roe deer in Yakutia

Daily food requirements and seasonal aspects of the diet have been studied in the Siberian roe deer (Capreolus pygargus Pall.) at the northernmost limits of its range in northeastern Asia, under specific sharply continental climatic conditions of the East Siberian light conifer (larch) taiga forests. Seasonal changes in the composition of the diet and the consumption of different food items are discussed.     

Environmental and climatic signals from stable isotopes in Anatolian tree rings, Turkey

Dendrochronological and oxygen–carbon isotopic analysis was conducted on tree rings collected at two different elevations from three different regions in western Anatolia, Turkey. Tree rings were sampled from Anatolian black pines (Pinus nigra Arn. subsp. pallasiana (Lamb.) Holmboe) of at least 200?years old through an N–S transect covering Bolu-Mengen (north), Kütahya–Simav, and Antalya–Alanya (south) forests. Cellulose was extracted from 5-year blocks of tree rings from lower to upper altitudes in each region. Dendrochronological site chronologies were built for each of the 6 sites, and response to climate was evaluated. The long-term δ¹³C chronologies comprising the period 1800–2008 strongly decline by an average of 2.5‰ from the mid-twentieth century, which corresponds with the commencement of industrialization in Turkey. Oxygen isotope (δ¹⁸O) time series show dominantly low undulations, although some high-frequency variations of up to 6‰ are also observed. As a result of latitude and continent effects, oxygen isotope values of tree rings show an average of 3‰ increase from south (Antalya) to the north (Bolu). Correlations between carbon isotope ratios yield high positive values indicating that industrial pollution exerts a major control on the δ¹³C systematics of the studied trees. Corrected δ¹³C ratios are mostly represented by strong positive responses to variations in temperature and precipitation, whereas δ¹⁸O and summer temperature/precipitation are negatively correlated. Tree-ring widths and δ¹⁸O display similar responses to the temperature and sensitivity to the climatic impact of historic volcanic eruptions, and changes in the solar irradiance record are also detected in other high-resolution paleoclimate archives for the region.     

Impacts of interannual climate variability on past and future forest composition

Numerous analyses of the possible impacts of future climatic changes on tree species composition have been published for both lowland and high-elevation forests. Most of these studies were based on the application of forest "gap" models, and the vast majority of them considered only changes in the average of climatic parameters over time. In this study, we use a unique data set on reconstructed past climatic variations to analyse forest dynamics simulated by the forest gap model ForClim. This analysis forms the basis for a systematic exploration of the ecological effects of changing means vs. changing variability of climate on central European forests. A reconstruction of historical climate covering the last 470 years in the Swiss lowlands (ClimIndex) is extrapolated to a transect across the alpine (cold) treeline and used to simulate the influence of climate variations on the time scale of decades on forest biomass and tree species composition at both sites. While the simulation at the low-elevation site shows little sensitivity to climate variations, the results from upper subalpine forests suggest that two major dieback events would have occurred at elevations above the current but below the climatic tree line, induced by clusters of exceptionally cold summers. The results are in agreement with available dendrochronological data and with documentary evidence on massive negative impacts on flora and fauna at high elevations during these periods. We conclude that ForClim is capable of capturing the effects on tree population dynamics of climate variability at these sites as reconstructed from the ClimIndex record. A factorial design is used to address the sensitivity of ForClim to changes of the long-term averages vs. changes of the variability of monthly temperature and precipitation data. To this end, the simulated tree species composition of near-natural forests is examined along a climate gradient in Europe. The results indicate that there are three types of forest response: (1) little sensitivity to both kinds of change, (2) strong sensitivity to changes in the means, but little sensitivity to changing variability, and (3) strong sensitivity to changing variability at least in parts of the examined climate space. Half of the cases investigated fall under the third category, suggesting that emphasis should be placed on also assessing the sensitivity of ecosystems to future changes in climate variability rather than on changes of average values alone. Electronic Publication     

Edge effects on pine stands in a large city

Manifestations of the edge effect on Scots pine (Pinus sylvestris L.) trees and stands were evaluated in natural southern taiga forests growing in a large industrial city (Yekaterinburg, Russia). For this purpose, 14 transects were laid out, each consisting of 6–10 circular 400-m² plots arranged in a line that extended for 140–260 m into the tree stand, perpendicular to its boundary. During transect surveys, 128 records were made of parameters characterizing the state of pine trees (height, diameter, defoliation level, needle age) and stands (density and timber volume). The boundaries differed in age, i.e., the time of formation (no more than 8 years and more than 20 years ago), and type (adjoining to motorways or wastelands). The edge effect manifested itself only near old boundaries and only for tree height (5 m lower near the boundary than deep in the forest) and for stand density and timber volume (25% lower), independently of the boundary type. Since the test parameters linearly increased with distance from the boundary, it was impossible to estimate the range of the edge effect.     

Climatic signal in radial increment of conifers in forest-steppe of southern Siberia and its dependence on local growing conditions

Radial increments have been studied in Siberian larch (Larix sibirica Ledeb.) and Scots pine (Pinus sylvestris L.) growing in the forest-steppe zone of southern Siberia in areas with different landscapes and levels of soil moisture supply but uniform climatic conditions. It has been shown that the radial increment significantly depends on climatic factors, primarily temperatures in April to July and September, sums of precipitation in April to July and September to November, and hydrothermal coefficient in May. The climatiC response in the wood of conifers in the forest-steppe zone differs depending on their species-specific features and local growing conditions.     

Distribution of frost injuries in the wood of conifers

The distribution of frost injuries in earlywood and latewood, in different zones of stem (with respect to height, diameter, and circumference), and at the bases of large roots has been studied in Siberian spruce (Picea obovata Ledeb.) and Siberian larch (Larix sibirica Ledeb.) at the northern limit of their ranges (the Obsko-Tazovskaya forest-tundra). It has been shown that a high frequency of frost injuries, especially in spruce, is characteristic of the study region. Mass frost injuries occur only in thin trees, mainly in the lower part of stems. Guidelines for collecting and preparing wood samples for the study of frost injuries are proposed.     

Effect of Ecological Factors on the Accumulation of Fluorine by Larches, <Emphasis Type="Italic">Larix gmelinii</Emphasis> (Rupr.) Rupr. and <Emphasis Type="Italic">L. sibirica</Emphasis> Ledeb., in Eastern Siberia

The accumulation of fluorine in the trunk wood and branches of Larix gmelinii growing on soils with a naturally increased fluoride content and of L. sibirica growing in a zone affected by atmospheric emissions from an aluminum plant has been studied. Based on results of in vivo and in vitro experiments, it is concluded that larch trees manifest individual differences in their response to fluorine.     

Differences in plant species diversity between conifer (<Emphasis Type="Italic">Pinus tabulaeformis</Emphasis>) plantations and natural forests in middle of the Loess plateau

We compared differences in plant species diversity between conifer (Pinus tabulaeformis) plantations and natural secondary forests in the middle of the Loess plateau. The goal of the study was to examine the differences in the effect of stand development on species diversity and in species responses to changes between forest types and between forest layers. To clarify the effects of differences in forest management, we emphasized the functional types of plant species occurring in each forest type. The result as follow: (1) The H′ and S of tree layer were significantly lower in natural conifer forest than old conifer and secondary forest, but were not different compared with mid aged conifer forest. The H′ and S of shrub layer were significantly lower in mid aged conifer forest compared with other forest types. The H′ of herb layer showed no significant differences in the four forest types. The evenness index (J′) of tree layer of mid aged conifer forest was lower than other forest communities and its J′ of shrub layer was highest although its richness of shrub layer was lower than in the other forest types. (2) The analysis of β diversity index also indicated large differences between conifer plantations and natural forests. Although the tree layer species were similar in old plantation and natural conifer forests, they differed greatly between the natural conifer and secondary forests. The natural conifer and secondary forest species composition in shrub layer differed significantly from those in plantation and secondary plots. Tree species were significantly less common in plantations than in abandoned coppice forests. Species composition in the herb layer of different forest types was similar. (3) The management of P. tabulaeformis plantations alters plant species composition considerably; the number of sub tall-tree species is increased in old aged conifer forest, especially species dispersed by animals. Plantation management appears to affect ecological processes through seed dispersal. From the perspective of management, the change in the structure and composition of the canopy in plantations could affect the behavior of dispersers and regeneration.