LONGITUDINAL SUCCESSION IN IMPACT ASSESSMENT OF RIVER SYSTEM FISH COMMUNITIES1

ABSTRACT: Longitudinal succession indicates that diversity and species richiess of fish communities increases with increasing river size (increasing habitat complexity and environmental stability). Cultural impacts tend to decrease diversity, species richness, and evenness. To assess the impacts of cultural events on the fish community of a river system, the bias caused by longitudinal succession was removed by normalizing diversity indices and species richness values for river size. The relationships between an impact index and the community parameters of evenness, normalized diversity, and normalized species richness were determined by regression analysis. Similar relationships were determined using nonnormalized data, and the results compared. The two sets of regressions were similar in terms of variance explained. Evenness, which is unaffected by longitudinal sucession, appears to be the most impact sensitive parameter. A major source of error and confusion rsults from introduced species which are simultaneously sources of biological impact and a component of the community. Numbers of introduced species tend to increase with both impact and river size.     

Composition, species diversity, and biomass of the herbaceous community in dry tropical forest of northern India in relation to soil moisture and light intensity

Herb layer contributes substantially to the species diversity of forests and responds relatively quickly to changes in the environment. The objectives of the present study were to understand the relationships among tree canopy cover, soil moisture, light intensity, herbaceous diversity and biomass in a dry tropical forest of India. For this, 20 locations equally distributed in four sites were selected. Four quadrats, each 1?×?1?m in size, were randomly placed for sampling at each location. For each quadrat, tree canopy cover, incident light, soil moisture, herbaceous diversity, and biomass were determined. Results indicated that the selected locations differed in terms of tree canopy cover, soil moisture, light intensity, herbaceous diversity, and biomass. Principal component analysis (PCA), using importance value indices of the component species yielded four groups corresponding to the four communities. PCA axes were related to the tree canopy cover, light intensity, and soil moisture and suggested that these variables had a profound effect on the organization and determination of herbaceous floristic composition and diversity. Positive relationships of tree canopy cover with soil moisture, herbaceous diversity and biomass, and those of soil moisture with herbaceous diversity and biomass suggested that the tree canopies facilitated the herbaceous communities by modifying environmental conditions that ultimately improved the diversity and production. Further, the study showed a linear relationship of herbaceous diversity with biomass, indicating the importance of species diversity for generating primary production in forest herbs.     

Ground flora trampling studies: Five years after closure

Trampling as an ecological factor is a major concern of the management of park, forest, nature preserves, and wilderness areas as recreational resources. Of particular importance to the management concept of carrying capacity is the relative resistance of native plant communities, to trampling and the resilience or the capacity of impacted communities to recover. This information can be used by management to establish seasonal use limits to prevent irreversible degradation of these resources. The purpose of this study was to follow the recolonizaton of experimental trail surfaces barren of vegetative cover and hiked at three levels of use intensity. Results of this study indicate that soil compaction as measured by soil penetration resistance increased progressively with use level while the total number of species, species diversity, and dominant index scores declined. A major finding was that the greatest degree of change occurred at the first level of hiking, indicating that major floristic measures were most affected by the least amount of hiking. Recolonization of impacted areas that received 100 trampling passes as measured by plant cover, dominant indices, floristic dissimilarity, and species diversity was similar to areas receiving four and eight times more trampling, despite major differences in soil penetration resistance. These data suggest limited use delivered at one time can be as damaging as increasing levels of use delivered at over time.     

Effects of Trampling Limitation on Coastal Dune Plant Communities

Sandy coastlines are sensitive ecosystems where human activities can have considerable negative impacts. In particular, trampling by beach visitors is a disturbance that affects dune vegetation both at the species and community level. In this study we assess the effects of the limitation of human trampling on dune vegetation in a coastal protected area of Central Italy. We compare plant species diversity in two recently fenced sectors with that of an unfenced area (and therefore subject to human trampling) using rarefaction curves and a diversity/dominance approach during a two year study period. Our results indicate that limiting human trampling seems to be a key factor in driving changes in the plant diversity of dune systems. In 2007 the regression lines of species abundance as a function of rank showed steep slopes and high Y-intercept values in all sectors, indicating a comparable level of stress and dominance across the entire study site. On the contrary, in 2009 the regression lines of the two fenced sectors clearly diverge from that of the open sector, showing less steep slopes. This change in the slopes of the tendency lines, evidenced by the diversity/dominance diagrams and related to an increase in species diversity, suggests the recovery of plant communities in the two fences between 2007 and 2009. In general, plant communities subject to trampling tended to be poorer in species and less structured, since only dominant and tolerant plant species persisted. Furthermore, limiting trampling appears to have produced positive changes in the dune vegetation assemblage after a period of only two years. These results are encouraging for the management of coastal dune systems. They highlight how a simple and cost-effective management strategy, based on passive recovery conservation measures (i.e., fence building), can be a quick (1–2 years) and effective method for improving and safeguarding the diversity of dune plant communities.     

HYDROLOGIC DESIGN CONSIDERATIONS OF CONSTRUCTED WETLANDS FOR URBAN STORMWATER RUNOFF1

ABSTRACT: The successful design of constructed wetlands requires a continuous supply of water or vegetation that can withstand drought conditions. Having a constant water source is the best alternative to insure species diversity throughout the season. Consequently, detention structure designs should be based on times between events as well as on hydrologic return periods, since between events is when most evaporation and infiltration losses are likely to occur. In arid or semi-arid environments, this is a difficult process because of long interevent times and seasonal changes in precipitation patterns. This discussion is predicated on the assumption that phytoplankton, epiphytic algae, and emergent vegetation require moist conditions to be effective at removing nutrients, metals and other pollutants. There are drought tolerant species of vegetation that can be used in constructed wetlands but it may take several days to re-establish the attached bacteria communities necessary for optimum pollutant removal. This paper examines a stochastic framework to examine the probability of extended dry periods based on historic rainfall data. The number of consecutive dry days is selected for a specified level of assurance. By multiplying this value by the sum of daily system losses, an overall pond volume can be determined that ensures a minimum depth of water. To illustrate the utility of the approach, the method is applied to a site in Spokane, Washington.     

缙云山国家级自然保护区蝴蝶生态学研究

2007年和2009年的4-9月对缙云山自然保护区蝶类资源进行了调查,共采集蝴蝶标本112种,隶属11科67属.区系分析表明,本地区蝶类资源东洋种占优势,古北种所占比例较小,具备东洋界的基本属性.结合1998-2000年统计数据,以年份为影响因子对蝴蝶群落多样性指数进行单因素方差分析,结果表明缙云山自然保护区10年来各多样性指数无显著变化,蝴蝶群落结构相对稳定.     

IMPACT OF WATERSHED URBANIZATION ON STREAM INSECT COMMUNITIES1

ABSTRACT: The impact of urbanization on stream insect communities was determined by sampling 22 sites in northern Virginia representing a range of human population densities. Watershed development had little effect on the total insect numbers (no./m²), but shifted the taxonomic composition markedly. Relative abundance of Diptera (mainly chironomids) increased at more highly urbanized sites, while most other insect orders including Ephemeroptera (mayflies), Coleoptera (bettles), Megaloptera (dobsonflies), and Plecoptera (stone-flies) decreased. Trichoptera (caddisfiles) exhibited a variable response. Genus diversity and richness (number of genera) were significantly higher in less urbanized streams. Two genera of chironomids were positively correlated with increased urbanization, while 14 other genera (scattered through several orders) were negatively related to human population density. Principal components analysis demonstrated a gradient from more urbanized to less urbanized stations based on generic and order level biological data. Results of this study indicate that watershed urbanization has a major impact on benthic insect communities even in the absence of point source discharges.     

Island biogeographical theory: Can it be used to predict lotic recovery rates?

Classic island biogeographic theory predicts that equilibrium will be reached when immigration and extinction rates are equal. These rates are modified by number of species in source area, number of intermediate islands, distance to recipient island, and size of intermediate islands. This general model has been variously modified and proposed to be a stochastic process with minimal competitive interaction or heavily deterministic. Predictive models of recovery (regardless of the end point chosen) have been based on the appropriateness of the MacArthur-Wilson models. Because disturbance frequency, severity, and intensity vary in their effect on community dynamics, we propose that disturbance levels should first be defined before evaluating the applicability of island biogeographical theory. Thus, we suggest a classification system of four disturbance levels based on recovery patterns by primary and secondary succession and faunal organization by primary (invasion of vacant areas) and secondary (remnant of previous community remains) processes. Level 1A disturbances completely destroy communities with no upstream or downstream sources of colonizers, while some component of near surface interstitial or hyporheic flora and fauna survive level 1B disturbances. Recovery has been reported to take from five years to longer than 25 years, when most invading colonists do not have an aerial form. Level 2 disturbances destroy the communities but leave upstream and downstream colonization sources (level 2A) and, sometimes, a hyporheic pool of colonizers (level 2B). Recovery studies have indicated primary succession and faunal structuring patterns (2A) with recovery times of 90–400 days or secondary succession and faunal structuring patterns (2B) with recovery times of 40–250 days. Level 3 disturbances result in reduction in species abundance and diversity along a stream reach; level 4 disturbances result in reduction of abundance and diversity in discrete patches. Both disturbance types lead to secondary succession and secondary faunal organization. Recovery rates can be quite rapid, varying from less than 10 days to 100 or more days. We suggest that island biogeographical models seem appropriate to recovery by secondary processes after level 3 and 4 disturbances, where competition may be an important organizing factor, while models of numerical abundance and resource tracking are probably of better use where community development is by primary succession (levels 1 and 2). Development of predictive recovery models requires research that addresses a number of fundamental questions. These include the role of hydrologic patterns on colonization dynamics, the role of nonaerial colonizers in recovery from level 1 disturbances, and assessment of the impact of changes in the order of invasion by colonizers of varying energetic efficiencies. Finally, we must be able to assemble these data and determine whether information that guides community organization at one level of disturbance can provide insights into colonization dynamics at other levels.     

ANALYSIS OF DATA FROM BIOLOGICAL SURVEYS OF STREAMS: DIVERSITY AND SAMPLE SIZE1

ABSTRACT: Brillouin's equation (H) for species diversity from information theory is to be preferred for the purposes of applied ecology over the equation of Shannon (H′) or the more commonly used approximate equation (H″). By its use, the difficult problem of delimiting the extent of the community being sampled in a stream survey can be avoided. Moreover, Brillouin's equation gives the exact diversity of the fully censused collection, whreas Shannon's diversity can only be approximated with a biased estimator. If we regard a sample as a message from the environment to the ecologist, Brillouin's equation is the proper one for computing its diversity. The product moment correlation coefficient between Brillouin's H for the total number of individuals from a group of samples and for randomly chosen subsets of 100 individuals from each sample was nearly as high as the correlation between H and H″ based on the total samples. This indicates that small sample sizes may give a useful diversity index. Replicated subsamples show that much smaller samples than are normally used can discriminate between communities from polluted and unpolluted environments. The use of smaller samples should reduce the cost of stream surveys.     

SPECTRAL ANALYSIS OF TIDAL FLUCTUATIONS IN GROUND WATER LEVEL1

ABSTRACT: Water level data at 16 ground water wells and two sea water gauging stations, coupled with barometric measurements in an alluvial plain in the central‐west region of Taiwan, are analyzed using spectral analysis in the time and frequency domains. The semi‐diurnal component from water level station is observed to be the most noticeable signal while the diurnal component is the less distinct signal recorded at the water level stations. Both semidiurnal and diurnal components are coupled with atmospheric pressure measurements. From the atmospheric pressure data, spectral analysis indicates that both the raw and the pressure adjusted water levels are almost in phase and retain the same amplitude in this area. It implies that the effect of pressure variations is not significant for the sea water and ground water level nearby; the astronomical tidal components, as expected, are the main factor causing fluctuation of ocean water and ground water levels in the Choshuihsi alluvial plain.     

Effects of soil physical characteristics and biotic interferences on the herbaceous community composition and species diversity on the campus of Banaras Hindu University,India

Soil, water and species diversity relationships are central components of the vegetation ecology. In this connection, the present study was performed on the three sites within the campus of Banaras Hindu University of India, to relate herbaceous species diversity to soil physical characteristic and the intensity of biotic interferences. At each site, three, 10 m × 10 m plots were randomly established and within each plot, four quadrats each 50 cm × 50 cm were randomly placed for sampling. For each quadrat, number of individuals and their herbage cover were recorded by species. Soil physical characteristics (soil moisture, water-holding capacity, soil porosity and bulk density), elements of biotic interferences and α-diversity and its components were determined for each plot. The plots were ordinated by Non-metric Multidimensional Scaling (NMS) using Importance Value Indices of the component species. Results showed that the selected locations differed in terms of soil moisture and species diversity parameters due to differences in biotic interferences. NMS ordination yielded three groups corresponding to the three communities experiencing different intensity of land use. NMS axes were substantially related to the soil and herbaceous diversity parameters and suggested that the elements of soil physical characteristics, intensity of biotic interferences and regional herbaceous species pool had profound effect on the organization and determination of herbaceous floristic composition. Further, the sample locations exhibiting greater soil moisture, water-holding capacity, soil porosity and lesser soil bulk density harboured greater herbaceous diversity. A negative relationship between indices of species diversity and soil bulk density revealed that the dry and compact soils due to greater biotic pressure contributed to the loss of species diversity. Reduction in livestock numbers, grazing pressure and soil bulk density could be helpful in the promotion of soil quality and species diversity.     

Evaluating cumulative effects on wetland functions: A conceptual overview and generic framework

This article outlines conceptual and methodological issues that must be confronted in developing a sound scientific basis for investigating cumulative effects on freshwater wetlands. We are particularly concerned with: (1) effects expressed at temporal and spatial scales beyond those of the individual disturbance, specific project, or single wetland, that is, effects occurring at the watershed or regional landscape level; and (2) the scientific (technical) component of the overall assessment process. Our aim is to lay the foundation for a research program to develop methods to quantify cumulative effects of wetland loss or degradation on the functioning of interacting systems of wetlands. Toward that goal we: (1) define the concept of cumulative effects in terms that permit scientific investigation of effects; (2) distinguish the scientific component of cumulative impact analysis from other aspects of the assessment process; (3) define critical scientific issues in assessing cumulative effects on wetlands; and (4) set up a hypothetical and generic structure for measuring cumulative effects on the functioning of wetlands as landscape systems.We provide a generic framework for evaluating cumulative effects on three basic wetland landscape functions: flood storage, water quality, and life support. Critical scientific issues include appropriate delineations of scales, identification of threshold responses, and the influence on different functions of wetland size, shape, and position in the landscape.The contribution of a particular wetland to landscape function within watersheds or regions will be determined by its intrinsic characteristics, e.g., size, morphometry, type, percent organic matter in the sediments, and hydrologic regime, and by extrinsic factors, i.e., the wetland's context in the landscape mosaic. Any cumulative effects evaluation must take into account the relationship between these intrinsic and extrinsic attributes and overall landscape function. We use the magnitude of exchanges among component wetlands in a watershed or larger landscape as the basis for defining the geographic boundaries of the assessment. The time scales of recovery for processes controlling particular wetland functions determine temporal boundaries. Landscape-level measures are proposed for each function.     

Predicting mosaics and wildlife diversity resulting from fire disturbance to a forest ecosystem

A model for predicting community mosaics and wildlife diversity resulting from fire disturbance to a forest ecosystem is presented. It applies an algorithm that delineates the size and shape of each patch from grid-based input data and calculates standard diversity measures for the entire mosaic of community patches and their included animal species. The user can print these diversity calculations, maps of the current community-type-age-class mosaic, and maps of habitat utilization by each animal species. Furthermore, the user can print estimates of changes in each resulting from natural disturbance. Although data and resolution level independent, the model is demonstrated and tested with data from the Lewis and Clark National Forest in Montana.     

The nature of cumulative impacts on biotic diversity of wetland vertebrates

There is no longer any doubt that cumulative impacts have important effects on wetland vertebrates. Interactions of species diversity and community structure produce a complex pattern in which environmental impacts can play a highly significant role. Various examples show how wetlands maintain the biotic diversity within and among vertebrate populations, and some of the ways that environmental perturbations can interact to reduce this diversity.The trophic and habitat pyramids are useful organizing concepts. Habitat fragmentation can have severe effects at all levels, reducing the usable range of the larger habitat generalists while threatening the genetic integrity of small, isolated populations. The complexity of trophic interactions, and the propensity, or necessity, of vertebrates to switch from one food source to another—something we know little about—makes using food chain support as a variable for predicting environmental impacts very questionable.Historical instances illustrate the effects of the accumulation of impacts on vertebrates. At present it is nearly impossible to predict the result of three or more different kinds of perturbations, although long-range effects can be observed. One case in point is waterfowl; while their ingestion of lead shot, harvesting by hunters during migration, and loss of habitat have caused waterfowl populations to decline, the proportional responsibility of these factors has not been determined.Further examples show multiplicative effects of similar actions, effects with long time lags, diffuse processes in the landscape that may have concentrated effects on a component subsystem, and a variety of other interactions of increasing complexity. Not only is more information needed at all levels; impacts must be assessed on a landscape or regional scale to produce informed management decisions. I conclude that a system of replicate wetland reserves that are allowed to interact naturally with the surrounding landscape will be more effective in preserving biotic diversity than isolated sanctuaries.     

A FIELD EVALUATION OF THE EFFECTS OF HEATED DISCHARGES ON FISH DISTRIBUTION1

ABSTRACT. Studies were conducted to determine the distribution of fish in the New and East Rivers in relation to thermal discharges from Appalachian Power Company's fossil fuel plant at Glen Lyn, Virginia. Over 15,300 specimens representing 41 species were collected with seines, electrogear and rotenone at six sampling locations from February, 1973 to October 1973. Sampling frequency was designed to evaluate the effects of ambient temperature upon preferred temperature. Diversity indices were calculated for each location. There was a slight decrease in the diversity indices for those stations located in the thermal discharge. Condition coefficients calculated for Notropis albeolus Jordan, Notropis rubellus Aqasaiz, Notropis spilopterus Cope, Ictalurus punctatus Rafinesque, and Etheostoma blennioides Rafuesque were found to be significantly (p =.05) lower in the thermal discharge for all species tested except E. blennioides. Temperatures were plotted against frequency of capture to determine a particular species temperature selection from field data and indicated that: (1) Some species avoided high temperatures (i.e., Curnpostoma anomalum Rafuesque). (2) Some species were attracted to high temperatures (i.e., Ictalurus punctatus). (3) Some species distribution was not effected by temperatures (i.e., Notropis spilopterus).     

THE EFFECTS OF DIPOTASSIUM ENDOTHALL ON THE ZOOPLANKTON AND WATER QUALITY OF A SMALL POND1

ABSTRACT: Zooplankton samples and dissolved oxygen-temperature readings and water samples were taken from a treatment and control pond on 41 and 42 separate occasions, respectively, both before and after the application of 5.0 mg/l (a.i.) dipotassium endothall to the 0.31 ha treatment pond on May 31, 1973. Seasonal fluctuations in the density of Cladocera and Copepoda coincided quite closely in both ponds and were similar to the fluctuations reported in the literature to be typical of temperate region ponds and lakes. An apparent seasonal shift in generic composition from .Daphnia spp. in May to Ceriodaphnia spp. in June to Chydorus spp. from July/early September occurred in both ponds and is thought to be at least partially due to fish predation. No apparent changes in species composition or generic densities of Cladocerans was noted in the treatment pond that did not also occur in the control pond. There were no noticeable effects of the dipotassium endothall on either the Calanoida or Cyclopoida suborders of Copepoda. A later pulse of Ostracoda in the treatment pond, when compared to the control, may have been due to the dipotassium endothall or to a combination of the effects of the herbicide on the macrophytes and the method of sampling. A decrease in dissolved oxygen below saturation, occurring in the treated pond from 3–21 days after treatment, was attributed to an increase in biological oxygen demand associated with weed-kill. There was no noticeable increase in plant nutrients (N and P) in the treated pond following herbicide application, nor were there any apparent changes in the other chemical parameters studied.     

Comparison of community composition and species diversity of understorey and overstorey tree species in a dry tropical forest of northern India

The study focuses on understorey-overstorey plant community dynamics in a dry tropical forest to facilitate appropriate management decisions. We compare community composition and species diversity of the understorey vegetation among five dry tropical forest sites in northern India. A total of 1500 quadrats distributed over 15 one-ha permanent plots in five sites differing in the degree of disturbance, were used to enumerate the understorey tree species and the results were compared with overstorey tree layer. The non-metric multidimensional scaling (NMS) ordination revealed that human disturbance intensity, as well as the overall disturbance regimes, and soil water holding capacity controlled the organisation of dry tropical forest understorey composition through effects on soil organic matter. The alpha-diversity and its components decreased with increasing human disturbance intensity, reflecting utilisation pressure and decreased soil fertility, as also revealed by the analysis of overstorey tree layer. There was a significant positive relationship between overstorey and understorey diversity. Results suggest that in the future, the existing understorey tree communities may replace the current dry tropical forest communities under prevailing environmental conditions. The study also asserts that the rate of species accumulation will be greater in more disturbed sites as well as at small spatial scale within each disturbance level.     

Analyzing ecotones to predict biotic productivity

A technique for predicting the potential biotic productivities of areas is presented. The simple method uses maps or other techniques of remote imagery in comparing adjacent terrestrial and/or littoral areas for investigations when ground reconnaissance is not feasible. Theedge index, the statistical basis for comparison, is calculated as the ratio of the length of a particular ecotone (for example, timberline or shoreline) that would be affected by a proposed development to the affected land area. In similar habitats an unbiased comparison results at the ordinal level of measurement. The method predicts that a higher edge index will correspond to a higher productivity. But because it is not established whether or not the relationship is somehow scalar, the method is not recommended for comparing greatly dissimilar or far distant areas.It has long been observed that ecotones (edges) where two biotic communities meet often produce more game than pure stands of either community. The reasons for this edge effect are reviewed. The demonstrably higher productivity of ecotones is partially explained in terms of the mechanisms that also account for diversity of species.An attempt is made to reconcile theoretical treatments of species diversity with the pragmatic methods for reaching conclusions, which are employed by specialists in resource management.     

CONNECTING GROUND WATER INFLUXES WITH FISH SPECIES DIVERSITY IN AN URBANIZED WATERSHED1

ABSTRACT: Valley Creek watershed is a small stream system that feeds the Schuylkill River near Philadelphia, Pennsylvania. The watershed is highly urbanized, including over 17 percent impervious surface cover (ISC) by area. Imperviousness in a watershed has been linked to fish community structure and integrity. Generally, above 10 to 12 percent ISC there is marked decline in fish assemblages with fish being absent above 25 percent ISC. This study quantifies the importance of ground water in maintaining fish species diversity in subbasins with over 30 percent ISC. Valley Creek contains an atypical fish assemblage in that the majority of the fish are warm‐water species, and the stream supports naturally reproducing brown trout, which were introduced and stocked from the early 1900s to 1985. Fish communities were quantified at 13 stations throughout the watershed, and Simpson's species diversity index was calculated. One hundred and nine springs were located, and their flow rates measured. A cross covariance analysis between Simpson's species diversity index and spring flow rates upstream of fish stations was performed to quantify the spatial correlation between these two variables. The correlation was found to be highest at lag distances up to about 400 m and drop off significantly beyond lag distances of about 800 m.