Spatio-temporal variation of macrobenthic communities in the mangrove-fringed Segara Anakan lagoon, Indonesia, affected by anthropogenic activities

The benthic macrofauna of the Segara Anakan lagoon, Java, Indonesia and its fringing mangroves were investigated between May 2004 and August 2006. This lagoon has been affected by various human activities for decades, in particular fishing, effluents from agriculture and industry, and illegal deforestation. In total, 163 taxa were identified, including 127 species occurring in the mangrove forest and 59 species in the subtidal. Mean density of macrobenthos was 2.5-fold higher in the central (881.8 ± 1,151.3 ind. m^?2) than in the eastern mangrove site (356.3 ± 218.8 ind. m^?2). Community structures differed significantly between these sites and showed a serial shift during the investigation period. Gastropods dominated the community in the central mangrove location, whereas sipunculids, polychaetes and gastropods had similar densities in the eastern site. Differences in community composition were best explained by three sediment properties: pore water salinity, δ¹³C_org, and C_org/N. It is suggested that small-scale heterogeneity of food availability and quality is a main factor determining the small-scale variability of the community composition. Compared to other Indo-West Pacific mangroves, species richness and densities are high. However, the dominance of specific taxa, especially of opportunistic species and the comparably low species richness in the subtidal of the lagoon may be attributed to the high sediment input by rivers in the central part and to the large-scale cutting of mangroves. Continuous tree logging will probably lead to a further spread of two fast growing understorey plants and thus to an increase of uniform swamp sites and a decrease of micro-habitats for benthic macrofauna. To our knowledge, this is the first detailed study on the spatio-temporal variation of benthic macrofauna in mangroves of Indonesia.     

Local level community forest management an effective tool in conserving forest biodiversity

Forest vegetation of a community managed forest (Van Panchayat) located between 1570–1700 m elevation was studied along four aspects. Quercus leucotrichophora was the dominant species on three of the four aspects. Across the aspects the total tree density ranged between 150 and 490 ind/ha, sapling density between 100 and 370 ind/ha, seedling density varied from 241 to 1090 ind/ha. The shrub density ranged from 166 to 690 ind/ha and herb density from 6000–17100 ind/ha. The total basal area of tree layer varied from 9.78 to 13.5 m²/ha showing that the forest is young. Species diversity value for tree layer varied from 0.14 to 1.69. Quercus leucotrichophora were regenerating well due to effective forest management by local communities. The higher proportion of seedlings of Quercus leucotrichophora indicating that the Van Panchayat are playing important role in conservation of oak forest of Himalaya.     

Effects of anthropogenic disturbances on vegetation diversity and structure: a case study in the remnant forests surrounding the village ecosystems of Tripura,Northeast India

The types, frequency, and intensity of human interference with forests markedly affect the vegetation dynamics. Assessment of the impact of anthropogenic disturbances on forest diversity and structure is one of the central issues related to human behaviour and forest ecosystems and management. Both species diversity and density are very sensitive to local anthropogenic disturbance, which can be reduced depending on land-use systems. We used 10 line transects of 500 × 10 m to record and measure all plants ≥10 cm girth in some of the remaining forests surrounding villages in Tripura, Northeast India. To predict the effects of anthropogenic pressure on species richness and forest structure, we recorded and scored all ongoing disturbances in all transects of the forests. Although our study did not show any significant effects on the mean number of species, all diversity indices were significantly affected by disturbance. Total stem density (F = 128.60, p < 0.0001) and basal area (F = 65.30, p < 0.0001) of the forest stands were significantly affected by disturbance. Disturbance significantly (F = 84.81, p < 0.0001) impacts the abundance of mature voluminous trees removed by extensive illegal logging. Further, stem density at the middle (F = 10.01, p < 0.05) and upper (F = 131.70, p < 0.0001) canopy stories was also reduced by high disturbance intensity. The present analysis will be useful to policy-makers and planners for implementation of sustainable forest management at both the local and regional scale.     

Carbon stock variation of Pinus roxburghii Sarg. Forest along altitudes of Garhwal Himalaya, India

The present study was undertaken in Pinus roxburghii forest along three different altitudes i.e., 1100, 1300 and 1500 meter above mean sea level of Garhwal Himalaya to understand the effect of altitudes on carbon stocks (live trees and soil) in Pinus roxburghii forest. Tree density of this forest ranged between 590 tree ha^?1 (upper altitude) to 640 tree ha^?1 (lower altitude). The highest total carbon density (TCD) of above and belowground carbon was 66.33 ± 29.92 Mg ha^?1 at lower altitude followed by 57.64 ± 16.75 Mg ha^?1 in middle altitude and 52.92 ± 6.52 Mg ha^?1 in upper altitude. Soil organic carbon was highest (33.20 ± 2.77 Mg ha^?1) at lower altitude followed by middle (22.61 ± 7.17 Mg ha^?1) and upper altitude (12.65 ± 6.10 Mg ha^?1). Total carbon stock (trees + soil) of Pinus roxburghii forest was maximum (99.53 Mg ha^?1) at lower altitude and minimum (65.57 Mg ha^?1) at upper altitude.     

Differences in plant species diversity between conifer (<Emphasis Type="Italic">Pinus tabulaeformis</Emphasis>) plantations and natural forests in middle of the Loess plateau

We compared differences in plant species diversity between conifer (Pinus tabulaeformis) plantations and natural secondary forests in the middle of the Loess plateau. The goal of the study was to examine the differences in the effect of stand development on species diversity and in species responses to changes between forest types and between forest layers. To clarify the effects of differences in forest management, we emphasized the functional types of plant species occurring in each forest type. The result as follow: (1) The H′ and S of tree layer were significantly lower in natural conifer forest than old conifer and secondary forest, but were not different compared with mid aged conifer forest. The H′ and S of shrub layer were significantly lower in mid aged conifer forest compared with other forest types. The H′ of herb layer showed no significant differences in the four forest types. The evenness index (J′) of tree layer of mid aged conifer forest was lower than other forest communities and its J′ of shrub layer was highest although its richness of shrub layer was lower than in the other forest types. (2) The analysis of β diversity index also indicated large differences between conifer plantations and natural forests. Although the tree layer species were similar in old plantation and natural conifer forests, they differed greatly between the natural conifer and secondary forests. The natural conifer and secondary forest species composition in shrub layer differed significantly from those in plantation and secondary plots. Tree species were significantly less common in plantations than in abandoned coppice forests. Species composition in the herb layer of different forest types was similar. (3) The management of P. tabulaeformis plantations alters plant species composition considerably; the number of sub tall-tree species is increased in old aged conifer forest, especially species dispersed by animals. Plantation management appears to affect ecological processes through seed dispersal. From the perspective of management, the change in the structure and composition of the canopy in plantations could affect the behavior of dispersers and regeneration.     

Hydrodynamics of the Segara Anakan lagoon

A hydrodynamic field study of the Segara Anakan lagoon combined with a reconstruction of the recorded data using a three-dimensional numerical model provides insights into the processes driving the hydrodynamics of the lagoon. This includes bathymetry, quantification of water volume fluxes with ADCP measurements, sea level recordings, salinity measurements, water budget and numerical model results for the periods of the field studies. Salinity measurements at the Klaces village show a complete flushing of the western lagoon during neap tide in the rainy season, which is assumed to increase the sedimentation. Mean salinities of the western part are in the order of 10 and 21 g/kg, while the eastern part has a larger mean salinity of 27 and 32 g/kg for the rainy and dry season, respectively. A numerical sensitivity study shows a relationship between the Citanduy discharge and the nett water volume transport between the eastern and western part of Segara Anakan. A nett transport to the west, which differs from neap to spring tides, turns to zero when Citanduy discharges are between 300 and 400 m³/s. The flushing time of the lagoon depends on the monsoon season and is about 1–3 days at the western outlet and increases to 12 days in the central parts.     

The effect of natural and anthropogenic disturbance in forest canopy and its effect on species richness in forests of Uttarakhand Himalaya,India

Pinus roxburghii (chir-pine) and Quercus leucothchophora (banj-oak) are dominant forests of mountainous part of the Uttarakhand Himalaya. The continued anthropogenic disturbance is opening the canopy, forming canopy gaps and as a result forest fragments are developing. Thus, the present study aims to analyze variations in species richness and vegetational parameters in relation to canopy gaps in forests. Total species richness was greater in open canopied forest compared to moderate and close canopied forests. In comparison between oak and pine forest, it was greater in oak forest while the proportion of common species was low between oak—pine forests. Mean species richness did not significantly vary from one canopy gap to another as well as in oak and pine dominated forest. This indicated that dominant forest types played an important role to form the community structure. The shrubs richness were greater in closed canopy and between the forests it was greater in pine forest. Tree and shrub density was low in open canopy while herb density was high in moderate canopy. Thus, this study indicated that the dominant canopy species play an important role in deciding the community structures especially the distribution of under canopy species. These parameters should be considered for conservation and maintenance of plant biodiversity of a region.     

Forest structure and regeneration along the altitudinal gradient in the Binsar Wildlife Sanctuary,Uttarakhand Himalaya,India

Forest structure and regeneration were studied along the altitudinal gradient in the Binsar Wildlife Sanctuary of Uttarakhand Himalaya in India. Stratified random sampling of tree species was done by placing minimum 15 quadrats of 10 × 10 m at each aspect and altitude. The results reveal that along the altitudinal gradient there were three types of forest communities in the Sanctuary viz., chir pine (Pinus roxburghii), oakchir pine (Quercus leucotrichophora and Pinus roxburghii) and oak (Quercus floribunda and Quercus leucotrichophora). Rhododendron arboreum, Cornus macrophylla and Lyonia ovalifolia were main associates of these forest communities. The oak forests had larger shrub population as compared to chir pine. Myrsine africana was the most dominant shrub across the altitudinal gradient, whereas few shrub species were restricted to a certain altitudinal range. The regeneration of chir pine was best at lower altitude on south and east aspects, which indicates that it mostly regenerates on warm and dry slopes. In general, the regeneration potential in most of the tree species declines with the altitude. The density of saplings and seedlings also represented the dominant species at each altitudinal range, which indicates the cyclic regeneration of forests in the Sanctuary area.     

Communities of trees along a tropical forest restoration gradient

Increasing rates of deforestation in tropical forests have been linked to agriculturalists. A critical concern generating debate is how well communities of trees recover into a more species rich ecosystem after restoration planting. The aim of the study was to evaluate the pattern of recovery of communities of tree, assess the influence of Acanthus pubescens, Lantana camara and Pennisetum purpureum, on the recovery as well as how restoration planting facilitates recruitment of other native tree seedlings along a gradient of forest restoration in Kibale National Park, Uganda after evictions of illegal settlers. We studied six restoration forests ranging in age from 3 to 16 years, naturally regenerating and three primary forests. Our results showed that recovery with natural regeneration was more effective than restoration planting although the latter enhanced recruitment of other native tree seedling. Tree recovery was generally correlated with age so that species density and diversity increased although at different rates. A reverse pattern was found for dominance but no clear pattern was found for tree density (individual/ha). Communities of tree showed directional patterns of change however community composition were still distinct among the different forests. A. pubescens, L. camara and P. purpureum negatively correlated with species density, tree density and diversity but a positive correlation was found for dominance. Restoration planting can reestablish forests with high species density, tree density and diversity, but this is dependent on age and the extent of the herbs, grasses and shrubs cover in tropical forests.     

Irregularity in frequency of mast seed years in <Emphasis Type="Italic">Quercus floribunda</Emphasis> a late successional species of Central Himalaya

The mid altitudinal oak, Quercus floribunda forms predominantly evergreen forest in Central Himalaya between 2000–2400 m. It is late successional, mature phase species that has limited regeneration on disturbance prone sites. This oak produces mast seed crops at an interval of 2–3 years. During masting in Q. floribunda the seed fall and germination is upto ten times greater than in normal years, emphasizing the importance of mast year crop in forest maintenance. However, no mast year in this species since the last nine years (1997–2005) is a matter of serious concern. The rise in the summer and winter temperature over a period of 15 years appear to have affected the frequency of masting in this oak. The importance of masting can be adjudged from the fact that 97% of the surviving seedlings m⁻² are of the mast year crop.     

Climate changes and tree stand dynamics at the upper limit of their growth in the North Ural mountains

The composition and structure of tree stands near the timberline have been studied on different slopes and at different elevations in the Tylaisko-Konzhakovsko-Serebryanskii Massif, the North Urals. It has been found that the upper limits of tree stands with different degrees of canopy closure have risen considerably (by about 100 m of elevation) since the mid-19th century, although the formation of these stands started as early as the late 18th century. Woodless areas in the eastern part of the massif started to be colonized by Larix sibirica in the late 18th to early 19th centuries; those in the western part, by Picea obovata in the mid-19th century; and in the southern part, by Betula tortuosa in the late 19th century. Analysis of meteorological data provides evidence for warming and increasing humidity of the climate since the late 19th century. Favorable climatic changes that facilitated the expansion of the forest have taken place both in the summer (prolongation of the growing period) and in winter seasons (increase of air temperature and precipitation). The observed differences in the composition and dynamics of tree stands between the studied areas of the mountain range are most probably explained by different requirements of tree species for the depth of snow cover and the degree of soil freezing.     

Pyrophytic Properties of the Larch Larix gmelinii in Terms of Life Strategies

The pyrophytic properties of the larch Larix gmelinii (Rupr.) Rupr. growing in the northern taiga of central Siberia have been studied. It has been found that, although the trees are highly combustible and fire resistance of tree stands is low, the species is highly pyrophytic, which allows it to retain its geographic range. At early stages of postfire successions, the larch exhibits characteristics of pyrogenic explerent and is subject to r-selection. When forest restoration is close to completion, the species passes into the patient state and acquires properties subject to K-selection.     

Grazing,forest density,and carbon storage: towards a more sustainable land use in Caatinga dry forests of Brazil

Grazing is the main land use in semi-arid regions of the world, and sustainable management practices are urgently needed to prevent their degradation. However, how different grazing intensities affect forest density and ecosystem functions is often not sufficiently understood to allow for management adaptations that safeguard the ecosystems and their functions in the long run. We assessed the aboveground carbon stocks and plant densities along a grazing gradient in the semi-arid seasonally dry tropical forest of north-eastern Brazil (Caatinga). On 45 study plots, we analysed the aboveground carbon stocks of the vegetation and determined forest density and recruitment as well as the population structure of the most abundant tree species. Grazing intensity was accounted for based on the weight of livestock droppings and classified as low, intermediate, or high. Mean aboveground carbon stock was 15.74?±?1.92 Mg ha^?1 with trees and shrubs accounting for 89% of the total amount. Grazing at high intensities significantly reduced aboveground carbon stocks of herbs but not of other plant functional types. Instead, aboveground carbon stocks of trees and shrubs were negatively related to altitude above sea level, which is a proxy for reduced water availability along with lower anthropogenic impact. The population structure of the most common tree species was characterised by abundant recruitment, irrespective of grazing, whereas the recruitment of less frequent woody species was negatively affected by grazing. Overall, our data imply that grazing and forage management need to be adapted, including the reduction of free-roaming livestock and storage of fodder, to maintain carbon storage and forest density.     

Fuelwood consumption in two tribal villages of the Nanda Devi Biosphere Reserve of the Indian Himalaya and strategies for fuelwood sustainability

Fuelwood is the only important source of energy in the mountainous region of the Garhwal Himalaya, India. Since the commercial source of energy is generally beyond the reach of ordinary people due to their poor socio-economic conditions and due to limited supply and lack of communication facilities for transport of LPG, the villages of the inner region of the Garhwal Himalaya depend on their fuel requirement from the forest. In the present study, two villages of the Nanda Devi Biosphere Reserve in Uttarakhand part of the Indian Himalaya i.e. Lata and Dunagiri located at 2,415 and 3,600 m altitudes, respectively, were selected for the study of socio-economic profile and vegetation and for estimation of per capita fuelwood consumption and the degree of disturbance. The study was conducted from 2002–2005. The population of these villages is migratory and belongs to the Bhotiya community, a scheduled tribe consisting of two subgroups known as Tolcha and Marchha. They grow traditional crops as well as cash crops. Important tree species used for fuelwood include Cedrus deodara, Pinus wallichiana, Cupressus torulosa, Taxus wallichiana, Acer indicum, Quercus dilatata and Viburnum cotinifolium. Maximum density among trees was shown by Pinus wallichiana (169.6 trees ha⁻¹) in village Lata and by Cedrus deodara (89.6 trees ha⁻¹) in village Dunagiri. The average per capita consumption of fuelwood in villages Lata and Dunagiri was 4.03 and 4.77 kg capita⁻¹ day⁻¹. Maximum number of trees (29 and 31% lopping for Lata and Dunagiri, respectively) belonged to disturbance class 1 (1–20% lopping) followed by the disturbance class 2 (20–40% lopping). Due to location of these villages in the buffer zone of the biosphere reserve, the fuelwood consumption may cause an adverse impact on the ecological status of this reserve, which urgently requires employing strategies for the conservation and management of this biosphere in terms of fuelwood sustainability e.g. regulation of livestock stock and grazing, using alternative sources of fuels, plantation of multipurpose trees and adoption of ecotourism.     

Monitoring regional vegetation change using reflectance measurements from multiple solar zenith angles.

Many traditional models of vegetation canopy reflectance have commonly used one of two approaches. Either the canopy is assumed to consist of discrete objects of known reflectance and geometric-optics are then used to calculate shading effects, or, as in the turbid medium approach, the canopy is treated as a horizontally homogeneous layer of small elements of known optical properties and radiative transfer theory is used to calculate canopy reflectance. This paper examines the effect of solar zenith angle on the reflectance of red and near-infrared radiation from forests using a combination of these modelling approaches. Forests are first modelled as randomly spaced eucalypt crowns over a homogeneous understorey and the fractional coverage of four components: shaded and sunlit canopy and shaded and sunlit understorey are calculated. Reflectance from each fraction is then modelled for a range of solar zenith angles using the Verhoef SAIL model. The overall scene reflection as seen by a nadir viewing satellite sensor is compared for three forest types representing a gradient of crown density from open dry grassy woodlands to dense wetter closed forest with an understorey of mesophytic plants. Modelled trends in scene reflectance change are consistent with aircraft measurements carried out at three different solar zenith angles. Results indicate that an increase in both tree density and solar zenith angle leads to an increase in the dominance of shaded components. In the visible band, both the sparsely treed woodland and the medium density dry forest show similar trends to that predicted by a turbid medium model, however, the wet forest shows a less rapid decrease in reflectance with solar zenith angle. In the near-infrared band, as tree density increases from woodland to wet forest, overall scene reflectance shows increased departure from that modelled using the traditional assumption of smooth homogeneous canopies, changing from an increase with solar zenith angle for the woodland to a decrease with solar zenith angle for the forest types.     

Concentrations of (137)Cs in summer pasture plants that reindeer feed on in the reindeer management area of Finland

Samples of summer pasture plants that reindeer feed on were collected in order to study ¹³⁷Cs concentrations in different plant species and in species nested in certain site types, and to study the regional distribution of ¹³⁷Cs in the Finnish reindeer management area. Plant species were categorized by the site types of mineral soil forest (xeric heath forest and mesic heath forest) and peatland. A third category called ’other plant species’ included plants with various site types, poorly determined species and species with poor statistics. The ¹³⁷Cs concentrations in different site types differed significantly. The mean ¹³⁷Cs concentrations of the whole reindeer management area in the xeric heath forest plant species was 44 ± 27 Bq/kg dw, in the mesic heath forest plant species 75 ± 59 Bq/kg dw and in the peatland plant species 219 ± 150 Bq/kg dw. The peatland species uptake ¹³⁷Cs more efficiently than plant species of mineral soil forests. A particularly efficient collector of ¹³⁷Cs was Trichophorum sp. It is suggested that Trichophorum sp. could be used as an indicator species for reindeer summer fodder plants. The highest concentrations of ¹³⁷Cs were found in Southern Lapland and the lowest in Northern Lapland. Today, the concentrations of ¹³⁷Cs in summer pasture plants that reindeer feed on in Finland are at such a level that there is no need to avoid any plant species. In the case of future nuclear fallout, reindeer grazing in peatlands would increase concentrations of ¹³⁷Cs in reindeer meat.     

Ecological Consequences of Transient Outbreaks of Siberian Larch Casebearer,Coleophora sibiricella Flkv., in the Udmurt Republic

A transient population outbreak of the Siberian larch casebearer, Coleophora sibiricella Flkv. (Lepidoptera, Coleophoridae), in a protective forest strip and its consequences for the host tree species (Larix sibirica Ledeb.) have been studied for the first time. It has been shown that one-time complete defoliation by this pest and the following infestation by xylophagous insects result in tree death, while the productivity of surviving trees decreases significantly. The results of this study indicate that the Siberian larch casebearer is a major phyllophagous pest of larch and that its populations must be monitored by specialists in forest pathology.     

Risks of global warming on montane and subalpine forests in Switzerland – a modeling study

In this study, we assess the risks of global warming on distributions of individual tree species in Switzerland. Applying a spatially explicit model, tree occurrence is predicted as a function of high resolution and physiologically relevant biophysical variables. The study shows that warming only slightly alters the overall abundance of tree species. However, the preferred temperature regimes are unevenly distributed along an elevational gradient, and the sensitivity to temperature varies considerably between the species. As a result, major reorganizations of forests in montane and subalpine belts are expected, i.e., a segregation of the core distributions of Fagus sylvatica and Picea abies is observed. Tree distribution potentials near timberlines, however, remain roughly the same. Electronic Publication