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1.
The Contribution of Headwater Streams to Biodiversity in River Networks1   总被引:1,自引:0,他引:1  
Abstract: The diversity of life in headwater streams (intermittent, first and second order) contributes to the biodiversity of a river system and its riparian network. Small streams differ widely in physical, chemical, and biotic attributes, thus providing habitats for a range of unique species. Headwater species include permanent residents as well as migrants that travel to headwaters at particular seasons or life stages. Movement by migrants links headwaters with downstream and terrestrial ecosystems, as do exports such as emerging and drifting insects. We review the diversity of taxa dependent on headwaters. Exemplifying this diversity are three unmapped headwaters that support over 290 taxa. Even intermittent streams may support rich and distinctive biological communities, in part because of the predictability of dry periods. The influence of headwaters on downstream systems emerges from their attributes that meet unique habitat requirements of residents and migrants by: offering a refuge from temperature and flow extremes, competitors, predators, and introduced species; serving as a source of colonists; providing spawning sites and rearing areas; being a rich source of food; and creating migration corridors throughout the landscape. Degradation and loss of headwaters and their connectivity to ecosystems downstream threaten the biological integrity of entire river networks.  相似文献   

2.
3.
Abstract: Knowledge of headwater influences on the water‐quality and flow conditions of downstream waters is essential to water‐resource management at all governmental levels; this includes recent court decisions on the jurisdiction of the Federal Clean Water Act (CWA) over upland areas that contribute to larger downstream water bodies. We review current watershed research and use a water‐quality model to investigate headwater influences on downstream receiving waters. Our evaluations demonstrate the intrinsic connections of headwaters to landscape processes and downstream waters through their influence on the supply, transport, and fate of water and solutes in watersheds. Hydrological processes in headwater catchments control the recharge of subsurface water stores, flow paths, and residence times of water throughout landscapes. The dynamic coupling of hydrological and biogeochemical processes in upland streams further controls the chemical form, timing, and longitudinal distances of solute transport to downstream waters. We apply the spatially explicit, mass‐balance watershed model SPARROW to consider transport and transformations of water and nutrients throughout stream networks in the northeastern United States. We simulate fluxes of nitrogen, a primary nutrient that is a water‐quality concern for acidification of streams and lakes and eutrophication of coastal waters, and refine the model structure to include literature observations of nitrogen removal in streams and lakes. We quantify nitrogen transport from headwaters to downstream navigable waters, where headwaters are defined within the model as first‐order, perennial streams that include flow and nitrogen contributions from smaller, intermittent and ephemeral streams. We find that first‐order headwaters contribute approximately 70% of the mean‐annual water volume and 65% of the nitrogen flux in second‐order streams. Their contributions to mean water volume and nitrogen flux decline only marginally to about 55% and 40% in fourth‐ and higher‐order rivers that include navigable waters and their tributaries. These results underscore the profound influence that headwater areas have on shaping downstream water quantity and water quality. The results have relevance to water‐resource management and regulatory decisions and potentially broaden understanding of the spatial extent of Federal CWA jurisdiction in U.S. waters.  相似文献   

4.
Abstract: Streams draining mountain headwater areas of the western Mojave Desert are commonly physically isolated from downstream hydrologic systems such as springs, playa lakes, wetlands, or larger streams and rivers by stream reaches that are dry much of the time. The physical isolation of surface flow in these streams may be broken for brief periods after rainfall or snowmelt when runoff is sufficient to allow flow along the entire stream reach. Despite the physical isolation of surface flow in these streams, they are an integral part of the hydrologic cycle. Water infiltrated from headwater streams moves through the unsaturated zone to recharge the underlying ground‐water system and eventually discharges to support springs, streamflow, isolated wetlands, or native vegetation. Water movement through thick unsaturated zones may require several hundred years and subsequent movement through the underlying ground‐water systems may require many thousands of years – contributing to the temporal isolation of mountain headwater streams.  相似文献   

5.
Abstract: Cool summertime stream temperature is an important component of high quality aquatic habitat in Oregon coastal streams. Within the Oregon Coast Range, small headwater streams make up a majority of the stream network; yet, little information is available on temperature patterns and the longitudinal variability for these streams. In this paper we describe preharvest spatial and temporal patterns in summer stream temperature for small streams of the Oregon Coast Range in forests managed for timber production. We also explore relationships between stream and riparian attributes and observed stream temperature conditions and patterns. Summer stream temperature, channel, and riparian data were collected on 36 headwater streams in 2002, 2003, and 2004. Mean stream temperatures were consistent among summers and generally warmed in a downstream direction. However, longitudinal trends in maximum temperatures were more variable. At the reach scale of 0.5‐1.7 km, maximum temperatures increased in 17 streams, decreased in seven streams and did not change in three reaches. At the subreach scale (0.1‐1.5 km), maximum temperatures increased in 28 subreaches, decreased in 14, and did not change in 12 subreaches. Models of increasing temperature in a downstream direction may oversimplify fine‐scale patterns in small streams. Stream and riparian attributes that correlated with observed temperature patterns included cover, channel substrate, channel gradient, instream wood jam volume, riparian stand density, and geology type. Longitudinal patterns of stream temperature are an important consideration for background characterization of water quality. Studies attempting to evaluate stream temperature response to timber harvest or other modifications should quantify variability in longitudinal patterns of stream temperature prior to logging.  相似文献   

6.
Headwater streams are the primary sources of water in a drainage network and serve as a critical hydrologic link between the surrounding landscape and larger, downstream surface waters. Many states, including North Carolina, regulate activity in and near headwater streams for the protection of water quality and aquatic resources. A fundamental tool for regulatory management is an accurate representation of streams on a map. Limited resources preclude field mapping every headwater stream and its origin across a large region. It is more practical to develop a model for headwater streams based on a sample of field data that can then be extrapolated to a larger area of interest. The North Carolina Division of Water Quality has developed a cost‐effective method for modeling and mapping the location, length, and flow classification (intermittent and perennial) of headwater streams. We used a multiple logistic regression approach that combined field data and terrain derivatives for watersheds located in the Triassic Basins ecoregion. Field data were collected using a standard methodology for identifying headwater streams and origins. Terrain derivatives were generated from digital elevation models interpolated from bare‐earth Light Detection and Range data. Model accuracies greater than 80% were achieved in classifying stream presence and absence, stream length and perennial stream length, but were not as consistent in predicting intermittent stream length.  相似文献   

7.
Abstract: Headwater streams make up a large proportion of the total length and watershed area of fluvial networks, and are partially characterized by the large volume of organic matter (large wood, detritus, and dissolved organic matter) and invertebrate inputs from the riparian forest, relative to stream size. Much of those inputs are exported to downstream reaches through time where they potentially subsidize river communities. The relative rates, timing, and conversion processes that carry inputs from small streams to downstream reaches are reasonably well quantified. For example, larger particles are converted to smaller particles, which are more easily exported. Also, dissolved organic matter and surface biofilms are converted to larger particles which can be more easily intercepted by consumers. However, the quality of these materials as it affects biological activity downstream is not well known, nor is the extent to which timing permits biological use of those particles. These ecological unknowns need to be resolved. Further, land uses may disrupt and diminish material transport to downstream reaches by removing sources (e.g., forest harvest), by affecting transport and decomposition processes (e.g., flow regulation, irrigation, changes in biotic communities), and by altering mechanisms of storage within headwaters (e.g., channelization). We present conceptual models of energy and nutrient fluxes that outline small stream processes and pathways important to downstream communities, and we identify informational gaps that, if filled, could significantly advance the understanding of linkages between headwater streams and larger rivers. The models, based on empirical evidence and best professional judgment, suggest that navigable waters are significantly influenced by headwater streams through hydrological and ecological connectivities, and land use can dramatically influence these natural connectivities, impacting downstream riverine ecosystems.  相似文献   

8.
Spatial patterns in major dissolved solute concentrations were examined to better understand impact of surface coal mining in headwaters on downstream water chemistry. Sixty sites were sampled seasonally from 2012 to 2014 in an eastern Kentucky watershed. Watershed areas (WA) ranged from 1.6 to 400.5 km2 and were mostly forested (58%–95%), but some drained as much as 31% surface mining. Measures of total dissolved solutes and most component ions were positively correlated with mining. Analytes showed strong convergent spatial patterns with high variability in headwaters (<15 km2 WA) that stabilized downstream (WA > 75 km2), indicating hydrologic mixing primarily controls downstream values. Mean headwater solute concentrations were a good predictor of downstream values, with % differences ranging from 0.55% (Na+) to 28.78% (Mg2+). In a mined scenario where all headwaters had impacts, downstream solute concentrations roughly doubled. Alternatively, if mining impacts to headwaters were minimized, downstream solute concentrations better approximated the 300 μS/cm conductivity criterion deemed protective of aquatic life. Temporal variability also had convergent spatial patterns and mined streams were less variable due to unnaturally stable hydrology. The highly conserved nature of dissolved solutes from mining activities and lack of viable treatment options suggest forested, unmined watersheds would provide dilution that would be protective of downstream aquatic life.  相似文献   

9.
Abstract: The volume and sustainability of streamflow from headwaters to downstream reaches commonly depend on contributions from ground water. Streams that begin in extensive aquifers generally have a stable point of origin and substantial discharge in their headwaters. In contrast, streams that begin as discharge from rocks or sediments having low permeability have a point of origin that moves up and down the channel seasonally, have small incipient discharge, and commonly go dry. Nearly all streams need to have some contribution from ground water in order to provide reliable habitat for aquatic organisms. Natural processes and human activities can have a substantial effect on the flow of streams between their headwaters and downstream reaches. Streams lose water to ground water when and where their head is higher than the contiguous water table. Although very common in arid regions, loss of stream water to ground water also is relatively common in humid regions. Evaporation, as well as transpiration from riparian vegetation, causing ground‐water levels to decline also can cause loss of stream water. Human withdrawal of ground water commonly causes streamflow to decline, and in some regions has caused streams to cease flowing.  相似文献   

10.
We reviewed the scientific literature on non‐floodplain wetlands (NFWs), freshwater wetlands typically located distal to riparian and floodplain systems, to determine hydrological, physical, and chemical functioning and stream and river network connectivity. We assayed the literature for source, sink, lag, and transformation functions, as well as factors affecting connectivity. We determined NFWs are important landscape components, hydrologically, physically, and chemically affecting downstream aquatic systems. NFWs are hydrologic and chemical sources for other waters, hydrologically connecting across long distances and contributing compounds such as methylated mercury and dissolved organic matter. NFWs reduced flood peaks and maintained baseflows in stream and river networks through hydrologic lag and sink functions, and sequestered or assimilated substantial nutrient inputs through chemical sink and transformative functions. Landscape‐scale connectivity of NFWs affects water and material fluxes to downstream river networks, substantially modifying the characteristics and function of downstream waters. Many factors determine the effects of NFW hydrological, physical, and chemical functions on downstream systems, and additional research quantifying these factors and impacts is warranted. We conclude NFWs are hydrologically, chemically, and physically interconnected with stream and river networks though this connectivity varies in frequency, duration, magnitude, and timing.  相似文献   

11.
In the Piedmont of North Carolina, a traditionally water‐rich region, reservoirs that serve over 1 million people are under increasing pressure due to naturally occurring droughts and increasing land development. Innovative development approaches aim to maintain hydrologic conditions of the undisturbed landscape, but are based on insufficient target information. This study uses the hydrologic landscape concept to evaluate reference hydrology in small headwater catchments surrounding Falls Lake, a reservoir serving Raleigh and the greater Triangle area. Researchers collected one year of detailed data on water balance components, including precipitation, evapotranspiration, streamflow, and shallow subsurface storage from two headwater catchments representative of two hydrologic landscapes defined by differences in soils and topographic characteristics. The two catchments are similar in size and lie within the same physiographic region, and during the study period they showed similar water balances of 26‐30% Q, ?4 to 5% ΔS, 59‐65% evapotranspiration, and 9‐10% G. However, the steeper, more elevated catchment exhibited perennial streamflow and nongrowing season runoff ratios (Q/P) of 33%, whereas the flat, low‐lying stream was drier during the growing season and exhibited Q/P ratios of 52% during the nongrowing season. A hydrologic landscape defined by topography and soil characteristics helps characterize local‐scale reference hydrology and may contribute to better land management decisions.  相似文献   

12.
Ephemeral and intermittent streams are abundant in the arid and semiarid landscapes of the Western and Southwestern United States (U.S.). Connectivity of ephemeral and intermittent streams to the relatively few perennial reaches through runoff is a major driver of the ecohydrology of the region. These streams supply water, sediment, nutrients, and biota to downstream reaches and rivers. In addition, they provide runoff to recharge alluvial and regional groundwater aquifers that support baseflow in perennial mainstem stream reaches over extended periods when little or no precipitation occurs. Episodic runoff, as well as groundwater inflow to surface water in streams support limited naturally occurring riparian communities. This paper provides an overview and comprehensive examination of factors affecting the hydrologic, chemical, and ecological connectivity of ephemeral and intermittent streams on perennial or intermittent rivers in the arid and semiarid Southwestern U.S. Connectivity as influenced and moderated through the physical landscape, climate, and human impacts to downstream waters or rivers is presented first at the broader Southwestern scale, and secondly drawing on a specific and more detailed example of the San Pedro Basin due to its history of extensive observations and research in the basin. A wide array of evidence clearly illustrates hydrologic, chemical, and ecological connectivity of ephemeral and intermittent streams throughout stream networks.  相似文献   

13.
Interest in connectivity has increased in the aquatic sciences, partly because of its relevance to the Clean Water Act. This paper has two objectives: (1) provide a framework to understand hydrological, chemical, and biological connectivity, focusing on how headwater streams and wetlands connect to and contribute to rivers; and (2) briefly review methods to quantify hydrological and chemical connectivity. Streams and wetlands affect river structure and function by altering material and biological fluxes to the river; this depends on two factors: (1) functions within streams and wetlands that affect material fluxes; and (2) connectivity (or isolation) from streams and wetlands to rivers that allows (or prevents) material transport between systems. Connectivity can be described in terms of frequency, magnitude, duration, timing, and rate of change. It results from physical characteristics of a system, e.g., climate, soils, geology, topography, and the spatial distribution of aquatic components. Biological connectivity is also affected by traits and behavior of the biota. Connectivity can be altered by human impacts, often in complex ways. Because of variability in these factors, connectivity is not constant but varies over time and space. Connectivity can be quantified with field‐based methods, modeling, and remote sensing. Further studies using these methods are needed to classify and quantify connectivity of aquatic ecosystems and to understand how impacts affect connectivity.  相似文献   

14.
Abstract: The Crown of the Continent is one of the premiere ecosystems in North America containing Waterton‐Glacier International Peace Park, the Bob Marshall‐Great Bear‐Scapegoat Wilderness Complex in Montana, various Provincial Parks in British Columbia and Alberta, several national and state forest lands in the USA, and Crown Lands in Canada. The region is also the headwater source for three of the continent’s great rivers: Columbia, Missouri and Saskatchewan that flow to the Pacific, Atlantic and Arctic Oceans, respectively. Headwaters originate in high elevation alpine environs characterized by high snow accumulations in winter and rainstorms in summer. Most headwaters of the region contain high quality waters with few ions in solution and extremely low nutrient concentrations. Alpine streams have few species of aquatic organisms; however, they often possess rare species and have hydrogeomorphic features that make them vulnerable to climatic change. Subalpine and valley bottom streams of the Crown of the Continent Ecosystem (CCE) flow through well forested watersheds. Along the elevation gradient, the streams and rivers of the CCE flow through series of confining and nonconfining valleys resulting in distinct canyon and floodplain reaches. The alluvial floodplains are characterized by high species diversity and bioproduction maintained by the hydrologic linkages of habitats. The streams and rivers of the CCE have low nutrient concentrations, but may be significantly affected by wildfire, various resource extraction activities, such as logging or mining and exurban encroachment. Wildfire has been shown to increase nutrient loading in streams, both during a fire and then following the fire for as much as 5 years. Logging practices increase nutrient loading and the algal productivity of stream periphyton. Logging and associated roads are also known to increase sediment transport into Crown of the Continent streams directly affecting spawning success of native trout. The CCE is one of the fastest growing regions in the USA because of the many recreational amenities of the region. And, while the region has many remarkably pristine headwater streams and receiving rivers, there are many pending threats to water quality and quantity. One of the most urgent threats comes from the coal and gas fields in the northern part of the Crown of the Continent, where coal deposits are proposed for mountain‐top removal and open‐pit mining operations. This will have significant effects on the waters of the region, its native plants and animals and quality of life of the people.  相似文献   

15.
Freshwater ecosystems are linked at various spatial and temporal scales by movements of biota adapted to life in water. We review the literature on movements of aquatic organisms that connect different types of freshwater habitats, focusing on linkages from streams and wetlands to downstream waters. Here, streams, wetlands, rivers, lakes, ponds, and other freshwater habitats are viewed as dynamic freshwater ecosystem mosaics (FEMs) that collectively provide the resources needed to sustain aquatic life. Based on existing evidence, it is clear that biotic linkages throughout FEMs have important consequences for biological integrity and biodiversity. All aquatic organisms move within and among FEM components, but differ in the mode, frequency, distance, and timing of their movements. These movements allow biota to recolonize habitats, avoid inbreeding, escape stressors, locate mates, and acquire resources. Cumulatively, these individual movements connect populations within and among FEMs and contribute to local and regional diversity, resilience to disturbance, and persistence of aquatic species in the face of environmental change. Thus, the biological connections established by movement of biota among streams, wetlands, and downstream waters are critical to the ecological integrity of these systems. Future research will help advance our understanding of the movements that link FEMs and their cumulative effects on downstream waters.  相似文献   

16.
Abstract: In January 2001, the U.S. Supreme Court ruled that the U.S. Army Corps of Engineers exceeded its statutory authority by asserting Clean Water Act (CWA) jurisdiction over non‐navigable, isolated, intrastate waters based solely on their use by migratory birds. The Supreme Court’s majority opinion addressed broader issues of CWA jurisdiction by implying that the CWA intended some “connection” to navigability and that isolated waters need a “significant nexus” to navigable waters to be jurisdictional. Subsequent to this decision (SWANCC), there have been many lawsuits challenging CWA jurisdiction, many of which are focused on headwater, intermittent, and ephemeral streams. To inform the legal and policy debate surrounding this issue, we present information on the geographic distribution of headwater streams and intermittent and ephemeral streams throughout the U.S., summarize major findings from the scientific literature in considering hydrological connectivity between headwater streams and downstream waters, and relate the scientific information presented to policy issues surrounding the scope of waters protected under the CWA. Headwater streams comprise approximately 53% (2,900,000 km) of the total stream length in the U.S., excluding Alaska, and intermittent and ephemeral streams comprise approximately 59% (3,200,000 km) of the total stream length and approximately 50% (1,460,000 km) of the headwater stream length in the U.S., excluding Alaska. Hillslopes, headwater streams, and downstream waters are best described as individual elements of integrated hydrological systems. Hydrological connectivity allows for the exchange of mass, momentum, energy, and organisms longitudinally, laterally, vertically, and temporally between headwater streams and downstream waters. Via hydrological connectivity, headwater, intermittent and ephemeral streams cumulatively contribute to the functional integrity of downstream waters; hydrologically and ecologically, they are a part of the tributary system. As this debate continues, scientific input from multiple fields will be important for policymaking at the federal, state, and local levels and to inform water resource management regardless of the level at which those decisions are being made. Strengthening the interface between science, policy, and public participation is critical if we are going to achieve effective water resource management.  相似文献   

17.
Recovery of lotic macroinvertebrate communities from disturbance   总被引:7,自引:0,他引:7  
Ecosystem disturbances produce changes in macrobenthic community structure (abundances, biomass, and production) that persist for a few weeks to many decades. Examples of disturbances with extremely long-term effects on benthic communities include contamination by persistent toxic agents, physical changes in habitats, and altered energy inputs. Stream size, retention, and local geomorphology may ameliorate the influence of disturbances on invertebrates. Disturbances can alter food webs and may select for favorable genotypes (e.g., insecticidal resistance). Introductions of pesticides into lotic ecosystems, which do not result in major physical changes within habitats, illustrate several factors that influence invertebrate recovery time from disturbance. These include: (1) magnitude of original contamination, toxicity, and extent of continued use; (2) spatial scale of the disturbance; (3) persistence of the pesticide; (4) timing of the contamination in relation to the life history stages of the organisms; (5) vagility of populations influenced by pesticides; and (6) position within the drainage network. The ability of macroinvertebrates to recolonize denuded stream habitats may vary greatly depending on regional life histories, dispersal abilities, and position within the stream network (e.g., headwaters vs larger rivers). Although downstream drift is the most frequently cited mechanism of invertebrate recolonization following disturbance in middle- and larger-order streams, evidence is presented that shows aerial recolonization to be potentially important in headwater streams. There is an apparent stochastic element operating for aerial recolonization, depending on the timing of disturbance and flight periods of various taxa. Available evidence indicates that recolonization of invertebrate taxa without an aerial adult stage requires longer periods of time than for those that possess winged, terrestrial adult stages (i.e., most insects). Innovative, manipulative experiments are needed in order to address recolonization mechanisms of animals inhabiting streams that differ in size, latitude, disturbance frequency and magnitude, as well as the potential influence of early colonists on successional sequences of species following disturbance.  相似文献   

18.
Riparian areas of large streams provide important habitat to many species and control many instream processes — but is the same true for the margins of small streams? This review considers riparian areas alongside small streams in forested, mountainous areas of the Pacific Northwest and asks if there are fundamental ecological differences from larger streams and from other regions and if there are consequences for management from any differences. In the moist forests along many small streams of the Pacific Northwest, the contrast between the streamside and upslope forest is not as strong as that found in drier regions. Small streams typically lack floodplains, and the riparian area is often constrained by the hillslope. Nevertheless, riparian‐associated organisms, some unique to headwater areas, are found along small streams. Disturbance of hillslopes and stream channels and microclimatic effects of streams on the riparian area provide great heterogeneity in processes and diversity of habitats. The tight coupling of the terrestrial riparian area with the aquatic system results from the closed canopy and high edge‐to‐area ratio for small streams. Riparian areas of the temperate, conifer dominated forests of the Pacific Northwest provide a unique environment. Forest management guidelines for small streams vary widely, and there has been little evaluation of the local or downstream consequences of forest practices along small streams.  相似文献   

19.
Stanfield, Les W. and Don A. Jackson, 2011. Understanding the Factors That Influence Headwater Stream Flows in Response to Storm Events. Journal of the American Water Resources Association (JAWRA) 1‐22. DOI: 10.1111/j.1752‐1688.2010.00518.x Abstract: Headwater drainage features (first‐ to second‐order streams) are the capillaries of the landscape that, among other things, moderate the timing and volumes of water available to the riparian and aquatic ecosystems. How these features respond to summer rainfall is poorly understood. We studied how geology and an index of land use/land cover influenced peak flows following rainfall events in 110 headwater stream sites that were studied over a four‐month period during a drought year. Highest peak flows were observed in the most urbanized catchments and in poorly drained soils, but specific responses were variable depending on both geology and land disturbance. Redundancy analysis indicated that both surficial geology and land disturbance were important factors influencing peak flows under drought conditions. We conclude that responses of these headwater streams to individual storms during drought conditions are unpredictable from data collected using our methods, but increased peak flows were associated with increased urban and agricultural development, but mitigated by surficial geology. These findings demonstrate the challenges to accurately predict flow conditions in headwater streams during periods of extreme weather that concurrently have the greatest potential effect on biota. The combination of these challenges and importance of such events indicates the need to develop new approaches to study and manage these resources.  相似文献   

20.
Wigington, Parker J., Jr., Scott G. Leibowitz, Randy L. Comeleo, and Joseph L. Ebersole, 2012. Oregon Hydrologic Landscapes: A Classification Framework. Journal of the American Water Resources Association (JAWRA) 1‐20. DOI: 10.1111/jawr.12009 Abstract: There is a growing need for hydrologic classification systems that can provide a basis for broad‐scale assessments of the hydrologic functions of landscapes and watersheds and their responses to stressors such as climate change. We developed a hydrologic landscape (HL) classification approach that describes factors of climate‐watershed systems that control the hydrologic characteristics of watersheds. Our assessment units are incremental watersheds (i.e., headwater watersheds or areas draining directly into stream reaches). Major components of the classification include indices of annual climate, climate seasonality, aquifer permeability, terrain, and soil permeability. To evaluate the usefulness of our approach, we identified 30 rivers with long‐term streamflow‐gauging records and without major diversions and impoundments. We used statistical clustering to group the streams based on the shapes of their annual hydrographs. Comparison of the streamflow clusters and HL distributions within river basin clusters shows that the Oregon HL approach has the ability to provide insights about the expected hydrologic behavior of HLs and larger river basins. The Oregon HL approach has potential to be a useful framework for comparing hydrologic attributes of streams and rivers in the Pacific Northwest.  相似文献   

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