城市生态系统的格局和过程

城市生态系统是人类活动影响下的复合生态系统。人为的开发利用活动和局地的生物物理条件形成了城市特有的景观格局。城市发展改变了自然的生物栖息环境,从根本上改变了生态系统的结构、过程、功能,以及所提供的各种服务。这种改变最终又会影响人类自身的健康生存。文章综述了国内外有关城市景观格局和和重要生态过程之间相互作用机制的研究进展。通过研究发现,在大尺度上,净初级生产力是衡量城市发展对区域物流、能流影响的有效指标;在小尺度上,城市物流、能流过程和城市内部的景观结构、格局有很强的相关性。城市发展显著的改变了自然的栖息地及生物生长过程,本地植被种类减少,外来种增加,外来种的数量和人口数量有显著的相关性;而城市动物群落主要由一些小型哺乳动物、无脊椎动物、鸟类、两栖爬行类动物组成,其中,鸟类是研究城市化过程的有效指示物种。城市生态系统特有的景观格局及生态等过程显著的改了局部区域的微气候和水文过程,城市气温增加,风速降低,云、雾量增加,地表径流增加,流速加大,污染负荷增强等。因此,控制人类活动,优化土地利用的结构和空间格局,是改善城市生态系统结构和过程,保证可持续发展的基本出发点。但是,目前有关城市-格局-过程之间的影响机制还不十分清楚,尤其是定量研究较少。因此,结合传统的城市生态学,景观生态学的理论和其他相关学科领域的发展,为深入分析城市格局-过程之间的关系提供了重要的研究前景。     

Linking deforestation scenarios to pollination services and economic returns in coffee agroforestry systems.

The ecological and economic consequences of rain forest conversion and fragmentation for biodiversity, ecosystem functioning, and ecosystem services like protection of soils, water retention, pollination, or biocontrol are poorly understood. In human-dominated tropical landscapes, forest remnants may provide ecosystem services and act as a source for beneficial organisms immigrating into adjacent annual and perennial agro-ecosystems. In this study, we use empirical data on the negative effects of increasing forest distance on both pollinator diversity and fruit set of coffee to estimate future changes in pollination services for different land use scenarios in Sulawesi, Indonesia. Spatially explicit land use simulations demonstrate that depending on the magnitude and location of ongoing forest conversion, pollination services are expected to decline continuously and thus directly reduce coffee yields by up to 18%, and net revenues per hectare up to 14% within the next two decades (compared to average yields of the year 2001). Currently, forests in the study area annually provide pollination services worth 46 Euros per hectare. However, our simulations also revealed a potential win-win constellation, in which ecological and economic values can be preserved, if patches of forests (or other natural vegetation) are maintained in the agricultural landscape, which could be a viable near future option for local farmers and regional land use planners.     

中国乡村城市化过程中的景观生态学问题与对策研究

城市和乡村的发展共同构成了社会-经济-自然复合生态系统的可持续发展,然而长期以来,重城市轻乡村的发展思想导致了城乡二元结构矛盾不断尖锐。随着经济的高速发展和加入WTO后的国际化冲击,未来的10~20年间,中国将进入一个乡村城市化快速发展的时期,这将使得中国本来紧张的人地关系变得更加严重,资源消耗与环境污染问题日益突出,乡村发展将面临更大的挑战,同时也将是中国乡村发展的重要契机。本文依据景观生态学原理与方法,通过研究乡村城市化进程中乡村景观的组合、结构、功能以及变化中存在的问题,提出了城乡协调、全盘布局和保留自身特色等相应的生态规划治理措施和对策,尽可能使乡村景观格局呈现多样性、异质性,并在功能上表现出稳定性和连续性,最终实现城乡景观和谐发展的局面。     

Definition and calculation of uncertainty in ecological process models

We present a method of multi-criteria assessment for the analysis of process model uncertainty that combines analysis of model structure, parameters and data requirements. There are three components in calculation and definition of uncertainty.

(1)

Assessment criteria: Uncertainty in a process model is reduced as the model can simultaneously simulate an increased number of assessment criteria selected to test specific aspects of the theory being investigated, and within acceptable limits set for those criteria. This reduces incomplete specification of the model—the characteristic that a model may explain some, but not all, of the observed features of a phenomenon. The calculation required is computation of the Pareto set which provides the list of simultaneously achieved criteria within specified ranges.     

Spatial distribution of ecological security status assessment of West-Liaohe River based on geographic information system

Eco-security assessment is a hot research area in resource and environmental science, which involves data with much spatial, non-linear, and random features. Geographic information system (GIS), as a useful tool to analyze and manage spatial information, has a superior advantage in this field. A case study in the western part of the Liaohe River featuring a method of eco-security spatial differences (ESSD) based on GIS is developed in this paper. The method includes four steps: 1) developing the pressure-state-response (P-S-R) framework with site data; 2) digitizing West-Liaohe River and setting its GRID database of ecosecurity assessment indicators; 3) figuring out the relative membership degree (RMD) of eco-security indicators by using the analytical hierarchy process with the weight of indicator; 4) classifying the security zone and mapping the assessment result of eco-security status in grid by GIS method of assigning and clustering. The visual spatial differences of eco-security based on GIS enables decision makers to know the status of eco-security better in making policies for achieving sustainability.     

桐麦间作人工群落空间分布格局的灰色局势决策

本文采用多目标灰色局势决策的方法，以桐麦间作人工群落为例，以１０年为泡桐生长的一个轮伐期，以６种不同的空间配置方式作为对策，以林业经济收益、农业经济收益和系统总体效益为目标，进行多目标决策，定量研究其空间分布格局。研究结果表明，以农为主间作型泡桐的最佳株行距为５×４０ｍ，以桐为主间作型的为５×５ｍ，农桐并重间作型的为５×２０ｍ。     

Paleobiology, community ecology, and scales of ecological pattern

The fossil record provides a wealth of data on the role of regional processes and historical events in shaping biological communities over a variety of time scales. The Quaternary record with its evidence of repeated climatic change shows that both terrestrial and marine species shifted independently rather than as cohesive assemblages over scales of thousands of years. Larger scale patterns also show a strong individualistic component to taxon dynamics; assemblage stability, when it occurs, is difficult to separate from shared responses to low rates of environmental change. Nevertheless, the fossil record does suggest that some biotic interactions influence large-scale ecological and evolutionary patterns, albeit in more diffuse and protracted fashions than those generally studied by community ecologists. These include: (1) the resistance by incumbents to the establishment of new or invading taxa, with episodes of explosive diversification often appearing contingent on the removal of incumbents at extinction events; (2) steady states of within-habitat and global diversity at longer time scales (10(7)-l0(8) yr), despite enormous turnover of taxa; and (3) morphological and biogeographic responses to increased intensities of predation and substratum disturbance over similarly long time scales. The behavior of species and communities over the array of temporal and spatial scales in the fossil record takes on additional significance for framing conservation strategies, and for understanding recovery of species, lineages, and communities from environmental changes.     

城乡建设用地景观格局多尺度差异

以郑州市域内1989、2000和2009年的典型城市与农村样地（25 km×25 km）为例,应用空间自相关、半方差、以及九项轨迹方差分析方法,对城市与农村建设用地景观多尺度上的空间格局特征与动态差异进行研究,结果表明,（1）城市建设用地景观聚集强度较高,斑块规模较大,而农村建设用地景观在大部分尺度上呈随机或均匀分布,斑块规模较小。（2）随着尺度的增加,城市及农村建设用地景观分布状态都由聚集向均匀过渡,但前者过渡较为平缓,而后者在小尺度范围（0~2km）变化速度较快,在较大尺度（3~16 km）上其分布状态的波动较小。（3）1989-2009年,城市与农村建设用地景观动态特征有较大差异,并表现出尺度依赖性。城市较小尺度上（0~4 km）建设用地景观的分布状态表现出聚集-分散的趋势,在较大尺度上（8~14 km）表现出分散-聚集的变化趋势。农村小尺度上（0~4 km）建设用地景观的分布状态也表现出聚集-分散的趋势,但大尺度（4~16 km）上状态变化很小且没有规律。     

Spatial and temporal pattern in seagrass community composition and productivity in south Florida

We document the distribution and abundance of seagrasses, as well as the intra-annual temporal patterns in the abundance of seagrasses and the productivity of the nearshore dominant seagrass (Thalassia testudinum) in the south Florida region. At least one species of seagrass was present at 80.8% of 874 randomly chosen mapping sites, delimiting 12,800 km² of seagrass beds in the 17,000-km² survey area. Halophila decipiens had the greatest range in the study area; it was found to occur over 7,500 km². The range of T. testudinum was almost as extensive (6,400 km²), followed by Syringodium filiforme (4,400 km²), Halodule wrightii (3,000 km²) and Halophila engelmanni (50 km²). The seasonal maxima of standing crop was about 32% higher than the yearly mean. The productivity of T. testudinum was both temporally and spatially variable. Yearly mean areal productivity averaged 0.70 g m⁻²day⁻¹, with a range of 0.05–3.29 g m⁻² day⁻¹. Specific productivity ranged between 3.2 and 34.2 mg g⁻¹ day⁻¹, with a mean of 18.3 mg g⁻¹ day⁻¹. Annual peaks in specific productivity occurred in August, and minima in February. Integrating the standing crop for the study area gives an estimate of 1.4 × 1011 g T. testudinum and 3.6 × 10¹⁰ g S. filiforme, which translate to a yearly production of 9.4 × 10¹¹ g T. testudinum leaves and 2.4 × 10¹¹ g S. filiforme leaves. We assessed the efficacy of rapid visual surveys for estimating abundance of seagrasses in south Florida by comparing these results to measures of leaf biomass for T. testudinum and S. filiforme. Our rapid visual surveys proved useful for quantifying seagrass abundance, and the data presented in this paper serve as a benchmark against which future change in the system can be quantified. Received: 30 January 2000 / Accepted: 24 July 2000     

生态环境脆弱地区城市化发展特征及城市发展路径选择流程研究

脆弱的生态环境会对城市化进程产生一定的约束作用。生态环境脆弱地区的城市化,表现出与非脆弱地区不同的特征。以往的研究,更倾向于对各类生态环境脆弱地区分别进行研究。本文试图站在更宏观的角度,通过研究生态环境脆弱和非脆弱地区这两个大类在城市化发展方面的异同,明确生态环境脆弱地区城市化发展的特征及合适的路径。首先在阅读了大量国内外相关文献的基础上,总结了干旱区、喀斯特地区、高寒地区、过渡区等类型的生态环境脆弱区的共性,了解其城市化现状及约束因素,并结合学者们的研究,与生态环境良好地区的城市化进行了对比,提出发展的对策。在理论分析的基础上,文章第二部分初步构建了一个生态环境脆弱地区城市化发展路径的选择方法及流程。研究表明生态环境脆弱区城市化的制约因素主要有地质条件、地形、水文、气候、植被和地上地下自然资源等,与生态环境良好地区城市化的差异主要体现在产业选择、人口承载力及城市建设用地等方面,这些方面也是该类地区在城市化过程中应格外重视的。生态环境脆弱地区城市化路径选择流程主要分为以下几个步骤：归类、分析城市化水平及阶段、分析生态环境演变历程、研究城市化与生态环境的相互作用机制、情景分析、制定战略规划,确定城市的功能、定位等。     

Community structure of the benthos in Scottish sea-lochs. II. Spatial pattern

Spatial pattern in the distribution of the macrobenthos was studied from series of van Veen grab samples taken in a systematic pattern in Lochs Etive and Creran, W. Scotland. Each series was collected in a circular pattern at 7 stations. Plots of the variances of the numerical occurrences of the species in the samples against their means revealed greater deviation from the random expectation in the case of the 2 muddy sand stations than for the soft-mud stations, with only 1 exception. Two tests of goodness-of-fit to the Poisson distribution were also applied. The degree of concordance between the 3 measures of non-randomness applied was measured. The best correlation was found between values of the ratio of variance: mean and the test statistic D of the Kolmogorov-Smirnov goodness-of-fit test. It was not possible to discuss the biological basis for the pattern measured, except for a few species shown to be markedly aggregated. No clear trend emerged from attempts to detect differences in pattern with various ecological groupings of the macrobenthos. However, from the use of the test values as sample statistics illustrative of overall community pattern, it was possible to conclude that the fauna of the shallow, usually current-swept areas of muddy sand in the sea-lochs investigated was more aggregated than that on the, usually deeper, soft-mud sediments in quieter water.     

玄武湖菹草种群空间格局分析及其环境效应

大型水生植物的分布格局与空间尺度有着密切关系,传统的分析方法只能分析一种尺度下的格局。引入点格局分析,以种群空间分布坐标点图为基础,分析各种尺度下的种群格局。运用点格局分析对玄武湖菹草种群的空间分布格局进行研究,并结合N、P等水质指标探讨其环境效应。结果表明,玄武湖菹草种群分布集群特征明显,且在尺度232~344m内聚集最为明显。从水质监测结果看,菹草种群有利于改善水体环境,无种群区的TN、TP和NO3-N浓度明显高于有种群区;溶解氧(DO)则表现为有种群区高于无种群区。菹草在空间分布上的明显规律性,会进一步导致水质参数的空间异质性。     

The invasion paradox: reconciling pattern and process in species invasions

The invasion paradox describes the co-occurrence of independent lines of support for both a negative and a positive relationship between native biodiversity and the invasions of exotic species. The paradox leaves the implications of native-exotic species richness relationships open to debate: Are rich native communities more or less susceptible to invasion by exotic species? We reviewed the considerable observational, experimental, and theoretical evidence describing the paradox and sought generalizations concerning where and why the paradox occurs, its implications for community ecology and assembly processes, and its relevance for restoration, management, and policy associated with species invasions. The crux of the paradox concerns positive associations between native and exotic species richness at broad spatial scales, and negative associations at fine scales, especially in experiments in which diversity was directly manipulated. We identified eight processes that can generate either negative or positive native-exotic richness relationships, but none can generate both. As all eight processes have been shown to be important in some systems, a simple general theory of the paradox, and thus of the relationship between diversity and invasibility, is probably unrealistic. Nonetheless, we outline several key issues that help resolve the paradox, discuss the difficult juxtaposition of experimental and observational data (which often ask subtly different questions), and identify important themes for additional study. We conclude that natively rich ecosystems are likely to be hotspots for exotic species, but that reduction of local species richness can further accelerate the invasion of these and other vulnerable habitats.     

广州中心城区公园绿地空间格局及可达性分析

公园绿地是城市中一类典型的开敞空间,对维护城市生态与人居环境健康具有重要意义,其空间布局的合理性与可达性直接影响其对城市居民的服务水平。以广州中心城区为例,借助GIS技术和景观格局指数,分析城市公园绿地景观格局、可达性和服务状况,并探讨城市公园绿地空间格局对其可达性影响。结果表明：①市级公园其面积占公园绿地总面积的84.71%,城市公园绿地聚集度为92.44,广州中心城区公园绿地景观类型和空间格局不尽合理;②网络分析表明研究区仅有22.35%的面积和44.67%的人口能够较便捷地到达邻近公园绿地,广州中心城区公园绿地的可达性和服务效果不理想;③城市公园绿地数量与空间格局是影响其可达性与服务水平的重要因子。     

基于景观格局的辽河三角洲湿地生态安全分析

以1990年和2005年遥感影像为基本信息源,在GIS技术支持下,在对辽河三角洲湿地景观格局分析的基础上,采用破碎度、分离度、优势度等景观格局指数和景观类型脆弱度为评价指标研究辽河三角洲湿地景观生态安全。20世纪90年代以来大规模的资源开发,加剧了辽河三角洲湿地景观的破碎化程度,使景观优势度增高,多样性下降。从1990-2005年,研究区主要湿地景观类型水稻田、苇田、滩涂的干扰程度有所增加,生态安全度均有所下降,表明人类对自然湿地生态系统的干扰越来越明显。总体来说,15年来研究区整体景观生态安全度呈现下降趋势。以往湿地生态安全的研究主要集中在环境脆弱性和保护策略方面,研究层次着重在生态系统层面,而从景观格局角度对湿地景观生态安全的研究涉及较少,文章涉及的是景观生态学可持续发展研究的一个新领域,所提出的内容实质上是景观生态安全定量表征的方法探讨。应用景观生态学方法研究生态安全,揭示景观结构与功能关系并进一步分析区域生态环境的变化趋势及其内在因素,不但为辽河三角洲湿地及其生物多样性的保护和资源开发提供了科学依据,而且丰富和发展了我国生态安全研究的理论与方法。     

Spatial pattern and persistence of historical fire boundaries in southern interior British Columbia

Recent ecosystem and fire management research aims to quantify, model and understand historical fire disturbances focusing on their frequency, size and distribution. Less attention, however, has been paid to fire boundaries and their location on the landscape. Our study presents a spatial method to quantify the location, pattern and persistence of historical fire boundaries using tree ring fire scar data in the lower Stein watershed (British Columbia). Data from Pinus ponderosa and Pseudotsuga menziesii collected in 35 one-hectare plots over a 412-hectare study area were analyzed for the period between 1879 and 1947 using local spatial statistics and boundary detection techniques. Results of the analysis using local spatial statistic Moran’s I showed significant clustering of boundaries near topographic breaks. To determine and test whether fire boundaries between plots were persistent, we used boundary detection methods and developed a spatially restricted randomization test. The results revealed that out of 86 possible boundary links, 8 were significantly persistent (P < 0.025) while another 8 were significantly rare (P < 0.025). These spatial methods can help determine the historical spatial configuration of persistent boundaries and can be used to maintain natural forest dynamics.

A coffee agroecosystem model: I. Growth and development of the coffee plant

This paper is the first of three on the coffee production system consisting of (1) the coffee plant, (2) coffee berry borer (CBB) and (3) the role of CBB parasitoids. A previous simulation model of the coffee plant was developed using data from Brazil where coffee phenology is characterized by distinct seasonal flowering (Gutierrez et al., 1998). In contrast, flowering in Colombia is continuous with low seasonality. To capture the differences in coffee phenology and growth in the two climatic regions, the Gutierrez et al. (1998) model was modified using new data from Colombia.The modifications to the model include:

(1)

The effect of solar radiation on floral buds initiation;

(2)

An age structure population model to track the daily input and development of the floral buds;

(3)

The effect of leaf water potential on breaking dormancy in flower buds, and hence on the timing and intensity of flowering;

(4)

The incorporation of both the vegetative and the reproductive demands to predict the photosynthetic rate.

(5)

The effect of low temperature on photosynthesis and defoliation.

Other aspects of the model were re-interpreted and refinements made to generalize its structure for use across coffee varieties and geographic areas. The model, without modification, realistically simulates field data from Brazil and two Colombian locations having different varieties, patterns of rainfall and hence flowering phenology.The model will be used as the base trophic level for incorporating CBB and high tropic levels effects, and for the analysis of management options in the coffee production system.     

川中浅丘农区坡地林农复合系统作物复种方式的综合效益评价

从能量转换效益、价值转换效益方面，对坡地林农复会系统１０种农作物复种方式进行了研究，并用层次分析法（ＡＨＰ）对各复种方式进行综合效益评价。结果表明，１０种复种方式中，小麦一花生的综合效益最高，５种复种方式较高，其余则较低。据此，提出了系列改进措施，供生产中应用。