MORPHOLOGICAL FEATURES OF SMALL STREAMS: SIGNIFICANCE AND FUNCTION1

ABSTRACT: Throughout the United States, land managers are becoming increasingly aware of the importance of small streams for a wide range of resource benefits. Where channel morphology is modified or structural features are added, stream dynamics and energy dissipation need to be considered. Unit stream power, defined here as the time-rate loss of potential energy per unit mass of water, can be reduced by adding stream obstructions, increasing channel sinuosity, or increasing flow resistance with large roughness elements such as woody root systems, logs, boulders, or bedrock. Notable morphological features of small streams are pools, riffles, bed material, and channel banks. Pools, which vary in size, shape, and causative factors, are important rearing habitat for fish. Riffles represent storage locations for bed material and are generally utilized for spawning. The particle sizes and distributions of bed material influence channel characteristics, bedload transport, food supplies for fish, spawning conditions, cover, and rearing habitat. Riparian vegetation helps stabilize channel banks and contributes in various ways to fish productivity. Understanding each stream feature individually and in relation to all others is essential for proper stream management. Although engineered structures for modifying habitat may alter stream characteristics, channel morphology must ultimately be matched to the hydraulic, geologic, and (especially) vegetative constraints of a particular location.     

A Daily Time Series Analysis of StreamWater Phosphorus Concentrations Along an Urban to Forest Gradient

During a 1-year period, we sampled stream water total phosphorus (TP) concentrations daily and soluble reactive phosphorus (SRP) concentrations weekly in four Seattle area streams spanning a gradient of forested to urban-dominated land cover. The objective of this study was to develop time series models describing stream water phosphorus concentration dependence on seasonal variation in stream base flows, short-term flow fluctuations, antecedent flow conditions, and rainfall. Stream water SRP concentrations varied on average by ±18% or ±5.7 μg/L from one week to another, whereas TP varied ±48% or ±32.5 μg/L from one week to another. On average, SRP constituted about 47% of TP. Stream water SRP concentrations followed a simple sine-wave annual cycle with high concentrations during the low-flow summer period and low concentrations during the high-flow winter period in three of the four study sites. These trends are probably due to seasonal variation in the relative contributions of groundwater and subsurface flows to stream flow. In forested Issaquah Creek, SRP concentrations were relatively constant throughout the year except during the fall, when a major salmon spawning run occurred in the stream and SRP concentrations increased markedly. Stream water SRP concentrations were statistically unrelated to short-term flow fluctuations, antecedent flow conditions, or rainfall in each of the study streams. Stream water TP concentrations are highly variable and strongly influenced by short-term flow fluctuations. Each of the processes assessed had statistically significant correlations with TP concentrations, with seasonal base flow being the strongest, followed by antecedent flow conditions, short-term flow fluctuations, and rainfall. Times series models for each individual stream were able to predict ∼70% of the variability in the SRP annual cycle in three of the four streams (r² = 0.57–0.81), whereas individual TP models explained ∼50% of the annual cycle in all streams (r² = 0.39–0.59). Overall, time series models for SRP and TP dynamics explained 82% and 76% of the variability for these variables, respectively. Our results indicate that SRP, the most biologically available and therefore most important phosphorus fraction, has simpler and easier-to-predict seasonal and weekly dynamics.     

Controls on Temperature in Salmonid‐Bearing Headwater Streams in Two Common Hydrogeologic Settings,Kenai Peninsula,Alaska

Headwater streams are the most numerous in terms of both number and length in the conterminous United States and play important roles as spawning and rearing grounds for numerous species of anadromous fish. Stream temperature is a controlling variable for many physical, chemical, and biological processes and plays a critical role in the overall health and integrity of a stream. We investigated the controls on stream temperature in salmon‐bearing headwater streams in two common hydrogeologic settings on the Kenai Peninsula, Alaska: (1) drainage‐ways, which are low‐gradient streams that flow through broad valleys; and (2) discharge‐slopes, which are high gradient streams that flow through narrow valleys. We hypothesize local geomorphology strongly influences surface‐water and groundwater interactions, which control streamflow at the network scale and stream temperatures at the reach scale. The results of this study showed significant differences in stream temperatures between the two hydrogeologic settings. Observed stream temperatures were higher in drainage‐way sites than in discharge‐slope sites, and showed strong correlations as a continuous function with the calculated topographic metric flow‐weighted slope. Additionally, modeling results indicated the potential for groundwater discharge to moderate stream temperature is not equal between the two hydrogeologic settings, with groundwater having a greater moderating effect on stream temperature at the drainage‐way sites.     

Can Warmwater Streams Be Rehabilitated Using Watershed-Scale Standard Erosion Control Measures Alone?

Degradation of warmwater streams in agricultural landscapes is a pervasive problem, and reports of restoration effectiveness based on monitoring data are rare. Described is the outcome of rehabilitation of two deeply incised, unstable sand-and-gravel-bed streams. Channel networks of both watersheds were treated using standard erosion control measures, and aquatic habitats within 1-km-long reaches of each stream were further treated by addition of instream structures and planting woody vegetation on banks (“habitat rehabilitation”). Fish and their habitats were sampled semiannually during 1–2 years before rehabilitation, 3–4 years after rehabilitation, and 10–11 years after rehabilitation. Reaches with only erosion control measures located upstream from the habitat measure reaches and in similar streams in adjacent watersheds were sampled concurrently. Sediment concentrations declined steeply throughout both watersheds, with means ≥40% lower during the post-rehabilitation period than before. Physical effects of habitat rehabilitation were persistent through time, with pool habitat availability much higher in rehabilitated reaches than elsewhere. Fish community structure responded with major shifts in relative species abundance: as pool habitats increased after rehabilitation, small-bodied generalists and opportunists declined as certain piscivores and larger-bodied species such as centrarchids and catostomids increased. Reaches without habitat rehabilitation were significantly shallower, and fish populations there were similar to the rehabilitated reaches prior to treatment. These findings are applicable to incised, warmwater streams draining agricultural watersheds similar to those we studied. Rehabilitation of warmwater stream ecosystems is possible with current knowledge, but a major shift in stream corridor management strategies will be needed to reverse ongoing degradation trends. Apparently, conventional channel erosion controls without instream habitat measures are ineffective tools for ecosystem restoration in incised, warmwater streams of the Southeastern U.S., even if applied at the watershed scale and accompanied by significant reductions in suspended sediment concentration.     

Recovery of temperate-stream fish communities from disturbance: A review of case studies and synthesis of theory

To evaluate the relative effect of autecologic factors, site-specific factors, disturbance characteristics, and community structure on the recovery of temperate-stream fish communities, we reviewed case histories for 49 sites and recorded data on 411 recovery end points. Most data were derived from studies of low-gradient third- or fourth-order temperate streams located in forested or agricultural watersheds. Species composition, species richness, and total density all recovered within one year for over 70% of systems studied. Lotic fish communities were not resilient to press disturbances (e.g., mining, logging, channelization) in the absence of mitigation efforts (recovery time >5 to >52 yr) and in these cases recovery was limited by habitat quality. Following pulse disturbances, autecological factors, site-specific factors, and disturbance-specific factors all affected rates of recovery. Centrarchids and minnows were most resilient to disturbance, while salmonid populations were least resilient of all families considered. Species within rock-substrate/nest-spawning guilds required significantly longer time periods to either recolonize or reestablish predisturbance population densities than did species within other reproductive guilds. Recovery was enhanced by the presence of refugia but was delayed by barriers to migration, especially when source populations for recolonization were relatively distant. Median population recovery times for systems in which disturbances occurred during or immediately prior to spawning were significantly less than median recovery times for systems in which disturbances occurred immediately after spawning. There was little evidence for the influence of biotic interactions on recovery rates.     

CENTRAL NEBRASKA RIVER BASINS,NEBRASKA1

ABSTRACT: The Central Nebraska Basins (NAWQA) study unit includes the Platte River and two major tributaries, the Loup and Elkhorn Rivers. Platte River flows are variable in the western part of the study unit because of diversions, but the Loup and Elkhorn Rivers originate in an area of dune sand covered by grassland that generates consistent base flows. More frequent runoff in the eastern part of the study unit also sustains stream flow. Ground water in the study unit has no regional confining units and the system is a water table aquifer throughout. Macroinvertebrate and fish taxa at biological sampling sites in the state were related to stream flow. One of the four wetland complexes identified in the study unit includes habitat for threatened and endangered bird species. The study unit is an agricultural area that includes row crops, both irrigated and nonirrigated in the eastern and southern parts, and rangeland in the Sand Hills of the western part. A water quality assessment will be based on the differences in environmental setting in each of four subunits within the study unit.]     

ESTIMATED STREAM FLOW REQUIREMENTS FOR FISHES OF THE WASHITA RIVER BELOW FOSS RESERVOIR,WESTERN OKLAHOMA1

Velocity and depth preference curves for juvenile and adult red shiners (Notropis lutrensis) were developed for each season of the year. The incremental methodology, developed by the U.S. Fish and Wildlife Service for quantifying the amount of physical habitat available for fish, was applied as a basis for recommending minimum stream flows for life stages of red shiners and channel catfish (Ictalurus punctatus) in the Washita River below Foss Reservoir. The minimum stream flow required to support populations of red shiners and channel catfish was estimated to be 0.60 m³/s during all months except April and May, during which flushing flows of 3.0 m³/s were recommended. Under drought conditions a contingency flow of 0.18 m³/s was recommended for August through March, a flushing flow of 2.4 m³/s for April and May, and a flow of 0.30 m³/s for channel catfish spawning in June and July. Standing crop of adult red shiners per weighted usable area averaged 2.73 g/m² (27 kg/ha) for two sites on the Washita River. A positive relation between standing crop and weighted usable area was suggested. Use of the incremental methodology on the study reach below Foss Reservoir required 96 person-hours of labor, excluding training, travel, and developing preference curves, and a total cost of $1,762, of which $1,200 was for the purchase of equipment.     

ANADROMOUS SALMONID RECOVERY IN THE UMATILLA RWER BASIN,OREGON: A CASE STUDY1

ABSTRACT: The Umatilla River Basin Fisheries Restoration Plan was initiated in the early 1980s to mitigate salmonid losses caused by hydroelectric development and habitat degradation. The objectives are to enhance the abundance of endemic steelhead and reintroduce extirpated chinook and coho salmon. The project prompted collaborative effort among federal, state, and tribal agencies, and local water users. It has incorporated habitat restoration, flow enhancement, fish passage improvements, and population supplementation through artificial production. Water exchanges have successfully increased minimum flows during spring and fall migration. While flows remain depressed compared to historic conditions, there is potential for improved habitat, passage, and homing. The mean adult‐to‐adult return rate of hatchery‐reared steelhead exceeded replacement and that of the naturally‐spawning population. Although the smolt‐to‐adult survival rates of hatchery‐reared fish fluctuate, salmonid escapement has increased in recent years, permitting steelhead and spring chinook harvest. Enumeration of potential spawners and observed redds reveals an increase in natural production of all supplemented species. Comparison of hatchery‐reared and naturally‐spawning steelhead populations revealed differences in life history characteristics (in age composition and sex ratios) though run timing and genetic stock compositions of the two components of the populations have not differed. Sustained monitoring is needed to determine benefits of integrating habitat restoration and artificial production in restoring salmonid populations.     

WATER LOSSES ALONG A REACH OF THE PECOS RIVER IN NEW MEXICO1

ABSTRACT: A reach of the Pecos River, located in eastern New Mexico, was examined to evaluate losses of river flows due to evaporation, seepage, and transpiration. An accurate assessment of the water losses along this reach is critical for determining how water rights are adjudicated for water users in the Pecos basin and interstate compact accounting. Water losses significantly impact flows through critical habitat for species protected under the Endangered Species Act. Daily losses of river flows were analyzed for the study reach that extends from immediately below the Pecos River confluence with Taiban Creek to the United States Geological Survey (USGS) gage near Acme. The analysis was completed with consideration for other processes including flood wave travel times and attenuation along with stream bank storage and returns. The analysis was completed using daily stream flow data from USGS gages located along the study reach. Empirical seasonal functions were developed to relate flow loss to the flow rate in the river. The functions were ultimately developed to provide a method for comparing the effects of different river flows on the available water supply.     

Climatic and stream-flow controls on tree growth in a Western montane riparian forest

Humans have severely impacted riparian ecosystems through water diversions, impoundments, and consumptive uses. Effective management of these important areas is becoming an increasingly high priority of land managers, particularly as municipal, industrial, and recreational demands for water increase. We examined radial tree growth of four riparian tree species (Pinus jeffreyi, Populus trichocarpa, Betula occidentalis, and Pinus monophylla) along Bishop Creek, California, and developed models relating basal area increment (BAI) and relative basal area increment (RBAI) to climatic and stream flow variables. Between years 1995–1999, univariate regression analysis with stream flow explained 29 to 61% of the variation in BAI and RBAI among all species except P. trichocarpa; growth by P. trichocarpa was not significantly related to stream flows over this period. Stepwise linear regression indicated that species responded differently to climatic variables, and models based on these variables explained between 33 to 86% of variation in BAI and RBAI during the decade of the 1990s. We examined branch growth of P. trichocarpa for sensitivity to differences in stream flow regimes and found that annual branch growth did not vary between a high- and low-flow site, but that annual branch growth was significantly higher in wet years with greater stream flows. Our results support the establishment of site-specific management goals by land managers that take into account all of the important tree species present in riparian ecosystems and their differential responses to altered hydrologic condition. Instream flow requirements for maintaining tree growth and vigor are only one of the species-specific responses that need to be evaluated, and these assessments should attempt to separate experimentally stream-flow (managed) controls from climatic (unmanaged) controls on growth.     

Application of the ELOHA Framework to Regulated Rivers in the Upper Tennessee River Basin: A Case Study

In order for habitat restoration in regulated rivers to be effective at large scales, broadly applicable frameworks are needed that provide measurable objectives and contexts for management. The Ecological Limits of Hydrologic Alteration (ELOHA) framework was created as a template to assess hydrologic alterations, develop relationships between altered streamflow and ecology, and establish environmental flow standards. We tested the utility of ELOHA in informing flow restoration applications for fish and riparian communities in regulated rivers in the Upper Tennessee River Basin (UTRB). We followed the steps of ELOHA to generate univariate relationships between altered flows and ecology within the UTRB. By comparison, we constructed multivariate models to determine improvements in predictive capacity with the addition of non-flow variables. We then determined whether those relationships could predict fish and riparian responses to flow restoration in the Cheoah River, a regulated system within the UTRB. Although ELOHA provided a robust template to construct hydrologic information and predict hydrology for ungaged locations, our results do not suggest that univariate relationships between flow and ecology (step 4, ELOHA process) can produce results sufficient to guide flow restoration in regulated rivers. After constructing multivariate models, we successfully developed predictive relationships between flow alterations and fish/riparian responses. In accordance with model predictions, riparian encroachment displayed consistent decreases with increases in flow magnitude in the Cheoah River; however, fish richness did not increase as predicted 4 years after restoration. Our results suggest that altered temperature and substrate and the current disturbance regime may have reduced opportunities for fish species colonization. Our case study highlights the need for interdisciplinary science in defining environmental flows for regulated rivers and the need for adaptive management approaches once flows are restored.     

Channel change,sediment transport,and fish habitat in a coastal stream: Effects of an extreme event

A study on sediment transport and channel change was conducted on Zayante Creek and the lower San Lorenzo River in Santa Cruz County, California. A rainstorm with a recurrence interval locally in excess of 150 years occurred during the study year, 1982 WY. Stream surveys indicated that significant aggradation occurred during and after the peak flood. Upper study reaches were substantially recovered after high flows of early April, but the lower study reaches still had significant filling of pools and burial of riffles by sand. Increases in width-depth ratio were minor and localized in upper reaches, but were significant in lower reaches. Large inputs of sand, primarily from landsliding, altered the sediment transport regime. A higher proportion of the bedload is now transported by lower flows than before the January event. Roads and sand quarries contributed significantly to sediment input to the stream. A proposed dam may alter the sediment transport regime of Zayante Creek. Mitigating the effects of this dam on downstream fish habitat may require occasional bankfull discharges.     

Riparian vegetation instream flow requirements: A case study from a diverted stream in the Eastern Sierra Nevada,California, USA

A methodology is described that allows determination of instream flow requirements for maintenance of riparian trees. Tree-ring data revealed strong relationships between tree growth and stream flow volume for riparian species at Rush Creek, an alluvial stream within an arid setting; these relationships allowed development of models that predict growth rates from hydrologic variables. The models can be used to assess instream flow requirements under the assumption that certain levels of growth are necessary to maintain the population. There is a critical need for development and use of instream flow methodologies for riparian vegetation, since present methodologies focus on needs of aquatic animals (e.g., fish) and may underestimate needs of the entire riparian ecosystem.     

STREAM HEALTH AFTER URBANIZATION1

ABSTRACT: Urban development has compromised the quality of physical elements offish habitat in low‐order spawning and rearing streams. In order to identify where priorities should lie in stream rehabilitation, field surveys of a number of streams were conducted near Vancouver, British Columbia. All of the streams were located in watersheds which were urbanized approximately 20 years earlier. The study watersheds ranged from 5 to 77 percent total impervious area (percent TIA). The urban streambeds were found to have less fine material and slightly higher values of intragravel dissolved oxygen than in rural streams. This improved gravel quality is attributed to the higher peak flows generated by impervious areas, and the reduced recruitment of fine material in the urban watersheds. Summer base flow was uniformly low when imperviousness was above 40 percent, evidenced by a decrease in velocity rather than water depth. Large woody debris (LWD) was scarce in all streams with > 20 percent TIA. A healthy buffer zone and abundant LWD were found to stabilize stream banks. The introduction of LWD is considered the most important strategy for stream rehabilitation. Stormwater detention ponds, in contrast, are concluded to have few hydrological benefits if constructed after a stream has reached its urban equilibrium.     

Regulated flushing in a gravel-bed river for channel habitat maintenance: A Trinity River fisheries case study

The operation of Trinity and Lewiston Dams on the Trinity River in northern California in the United States, combined with severe watershed erosion, has jeopardized the existence of prime salmonid fisheries. Extreme streamflow depletion and stream sedimentation below Lewiston have resulted in heavy accumulation of coarse sediment on riffle gravel and filling of streambed pools, causing the destruction of spawning, nursery, and overwintering habitat for prized chinook salmon (Salmo gairdnerii) and steelhead trout (Oncorhynchus tschawytscha). Proposals to restore and maintain the degraded habitat include controlled one-time remedial peak flows or annual maintenance peak flows designed to flush the spawning gravel and scour the banks, deltas, and pools. The criteria for effective channel restoration or maintenance by streambed flushing and scouring are examined here, as well as the mechanics involved.The liabilities of releasing mammoth scouring-flushing flows approximating the magnitude that preceded reservoir construction make this option unviable. The resulting damage to fish habitat established under the postproject streamflow regime, as well as damage to human settlements in the floodplain, would be unacceptable, as would the opportunity costs to hydroelectric and irrigation water users. The technical feasibility of annual maintenance flushing flows depends upon associated mechanical and structural measures, particularly instream maintenance dredging of deep pools and construction of a sediment control dam on a tributary where watershed erosion is extreme. The cost effectiveness of a sediment dam with a limited useful economic life, combined with perpetual maintenance dredging, is questionable.     

INFLUENCES OF WATERSHED,RIPARIAN‐CORRIDOR,AND REACH‐SCALE CHARACTERISTICS ON AQUATIC BIOTA IN AGRICULTURAL WATERSHEDS1

ABSTRACT: Multivariate analyses and correlations revealed strong relations between watershed and riparian‐corridor land cover, and reach‐scale habitat versus fish and macroinvertebrate assemblages in 38 warmwater streams in eastern Wisconsin. Watersheds were dominated by agricultural use, and ranged in size from 9 to 71 km² Watershed land cover was summarized from satellite‐derived data for the area outside a 30‐m buffer. Riparian land cover was interpreted from digital orthophotos within 10‐, 10‐to 20‐, and 20‐to 30‐m buffers. Reach‐scale habitat, fish, and macroinvertebrates were collected in 1998 and biotic indices calculated. Correlations between land cover, habitat, and stream‐quality indicators revealed significant relations at the watershed, riparian‐corridor, and reach scales. At the watershed scale, fish diversity, intolerant fish and EPT species increased, and Hilsenhoff biotic index (HBI) decreased as percent forest increased. At the riparian‐corridor scale, EPT species decreased and HBI increased as riparian vegetation became more fragmented. For the reach, EPT species decreased with embeddedness. Multivariate analyses further indicated that riparian (percent agriculture, grassland, urban and forest, and fragmentation of vegetation), watershed (percent forest) and reach‐scale characteristics (embeddedness) were the most important variables influencing fish (IBI, density, diversity, number, and percent tolerant and insectivorous species) and macroinvertebrate (HBI and EPT) communities.     

A Regional‐Scale Habitat Suitability Model to Assess the Effects of Flow Reduction on Fish Assemblages in Michigan Streams1

Zorn, Troy G., Paul W. Seelbach, and Edward S. Rutherford, 2012. A Regional‐Scale Habitat Suitability Model to Assess the Effects of Flow Reduction on Fish Assemblages in Michigan Streams. Journal of the American Water Resources Association (JAWRA) 48(5): 871‐895. DOI: 10.1111/j.1752‐1688.2012.00656.x Abstract: In response to concerns over increased use and potential diversion of Michigan’s freshwater resources, and the resulting state legislative mandate, an advisory council created an integrated assessment model to determine the potential for water withdrawals to cause an adverse resource impact to fish assemblages in Michigan’s streams. As part of this effort, we developed a model to predict how fish assemblages characteristic of different stream types would change in response to decreased stream base flows. We describe model development and use in this case study. The model uses habitat suitability information (i.e., catchment size, base‐flow yield, and July mean water temperature) for over 40 fish species to predict assemblage structure in an individual river segment under a range of base‐flow reductions. By synthesizing model runs for individual fish species at representative segments for each of Michigan’s 11 ecological stream types, we developed curves describing how typical fish assemblages in each type respond to flow reduction. Each stream type‐specific, fish response curve was used to identify streamflow reduction levels resulting in adverse resource impacts to characteristic fish populations, the regulatory standard. Used together with a statewide map of stream types, our model provided a spatially comprehensive framework for evaluating impacts of flow withdrawals on biotic communities across a diverse regional landscape.     

Importance of Riparian Forests in Urban Catchments Contingent on Sediment and Hydrologic Regimes

Forested riparian corridors are thought to minimize impacts of landscape disturbance on stream ecosystems; yet, the effectiveness of streamside forests in mitigating disturbance in urbanizing catchments is unknown. We expected that riparian forests would provide minimal benefits for fish assemblages in streams that are highly impaired by sediment or hydrologic alteration. We tested this hypothesis in 30 small streams along a gradient of urban disturbance (1–65% urban land cover). Species expected to be sensitive to disturbance (i.e., fluvial specialists and “sensitive” species that respond negatively to urbanization) were best predicted by models including percent forest cover in the riparian corridor and a principal components axis describing sediment disturbance. Only sites with coarse bed sediment and low bed mobility (vs. sites with high amounts of fine sediment) had increased richness and abundances of sensitive species with higher percent riparian forests, supporting our hypothesis that response to riparian forests is contingent on the sediment regime. Abundances of Etheostoma scotti, the federally threatened Cherokee darter, were best predicted by models with single variables representing stormflow (r² = 0.34) and sediment (r² = 0.23) conditions. Lentic-tolerant species richness and abundance responded only to a variable representing prolonged duration of low-flow conditions. For these species, hydrologic alteration overwhelmed any influence of riparian forests on stream biota. These results suggest that, at a minimum, catchment management strategies must simultaneously address hydrologic, sediment, and riparian disturbance in order to protect all aspects of fish assemblage integrity.     

ANALYSIS AND MODELING OF LONG-TERM STREAM TEMPERATURES ON THE STEAMBOAT CREEK BASIN,OREGON: IMPLICATIONS FOR LAND USE AND FISH HABITAT1

ABSTRACT: Steamboat Creek basin is an important source of timber and provides crucial spawning and rearing habitat for anadromous steelhead trout (Oncorhynchus mykiss). Because stream temperatures are near the upper limit of tolerance for the survival of juvenile steelhead, the possible long-term effect of clear-cut logging on stream temperatures was assessed. Twenty-year (1969–1989) records of summer stream temperature and flow from four tributaries and two reaches of Steamboat Creek and Boulder Creek (a nearby unlogged watershed) were analyzed. Logging records for the Steamboat Creek basin and air temperature records also were used in the analysis. A time-series model of the components of stream temperature (seasonal cycle of solar radiation, air temperature, streamflow, an autoregressive term of order 1, and a linear trend variable) was fitted to the water-temperature data. The linear trend variable was significant in all the fitted models except Bend Creek (a tributary fed by cool ground-water discharge) and Boulder Creek. Because no trends in either climate (i.e., air temperature) or streamflow were found in the data, the trend variable was associated with the pre-1969 loss and subsequent regrowth of riparian vegetation and shading canopies.     

Fish Assemblage Responses to Water Withdrawals and Water Supply Reservoirs in Piedmont Streams

Understanding effects of flow alteration on stream biota is essential to developing ecologically sustainable water supply strategies. We evaluated effects of altering flows via surface water withdrawals and instream reservoirs on stream fish assemblages, and compared effects with other hypothesized drivers of species richness and assemblage composition. We sampled fishes during three years in 28 streams used for municipal water supply in the Piedmont region of Georgia, U.S.A. Study sites had permitted average withdrawal rates that ranged from < 0.05 to > 13 times the stream’s seven-day, ten-year recurrence low flow (7Q10), and were located directly downstream either from a water supply reservoir or from a withdrawal taken from an unimpounded stream. Ordination analysis of catch data showed a shift in assemblage composition at reservoir sites corresponding to dominance by habitat generalist species. Richness of fluvial specialists averaged about 3 fewer species downstream from reservoirs, and also declined as permitted withdrawal rate increased above about 0.5 to one 7Q10-equivalent of water. Reservoir presence and withdrawal rate, along with drainage area, accounted for 70% of the among-site variance in fluvial specialist richness and were better predictor variables than percent of the catchment in urban land use or average streambed sediment size. Increasing withdrawal rate also increased the odds that a site’s Index of Biotic Integrity score fell below a regulatory threshold indicating biological impairment. Estimates of reservoir and withdrawal effects on stream biota could be used in predictive landscape models to support adaptive water supply planning intended to meet societal needs while conserving biological resources.