Understanding the Large‐Scale Influence of Levees on Floodplain Connectivity Using a Hydrogeomorphic Approach

The widespread construction of levees has reduced river–floodplain connectivity and altered associated fluvial processes in many river systems. Despite the recognition that levees can alter floodplain connectivity, few studies have examined the role of levees in reducing floodplain areas at large watershed scales. This paper explores the application of a hydrogeomorphic floodplain inundation model in the Wabash Basin, located in the Midwestern United States, to assess changes in floodplain area in levee‐protected areas. We evaluate 10‐ and 30‐m topographic resolutions and spatially examine the influence of levees on floodplain area in relation to river network attributes. Generally, floodplains in levee‐protected areas were influenced by topographic resolution, stream order, and elevation details of levees found in topography datasets. We show, when compared to Federal Emergency Management Agency maps, our approach underpredicts floodplain area when using 10‐m resolution topography data but only slightly overpredicts when using 30‐m resolution data. After removing details of levees from topography data, we found changes in floodplain area varied spatially, but basin‐aggregate results changed little compared to topography datasets that contain levees, though larger floodplain areas were produced in some regions where levees were removed. This work contributes to a growing research emphasis on using hydrogeomorphic floodplain models to understand floodplain disconnectivity.     

Characterizing the Extent of Spatially Integrated Floodplain and Wetland Systems in the White River,Indiana, USA

Floodplain delineation may inform protection of wetland systems under local, state, or federal laws. Nationally available Federal Emergency Management Agency Flood Insurance Rate Maps (FIRMs, “100‐year floodplain” maps) focus on urban areas and higher‐order river systems, limiting utility at large scales. Few other national‐scale floodplain data are available. We acquired FIRMs for a large watershed and compared FIRMs to floodplain and integrated wetland area mapping methods based on (1) geospatial distance, (2) geomorphic setting, and (3) soil characteristics. We used observed flooding events (OFEs) with recurrence intervals of 25‐50 to >100 years to assess floodplain estimate accuracy. FIRMs accurately reflected floodplain areas based on OFEs and covered 32% of river length, whereas soil‐based mapping was not as accurate as FIRMs but characterized floodplain areas over approximately 65% of stream length. Geomorphic approaches included more areas than indicated by OFE, whereas geospatial approaches tended to cover less area. Overall, soil‐based methods have the highest utility in determining floodplains and their integrated wetland areas at large scales due to the use of nationally available data and flexibility for regional application. These findings will improve floodplain and integrated wetland system extent assessment for better management at local, state, and national scales.     

Hydrological,Physical, and Chemical Functions and Connectivity of Non‐Floodplain Wetlands to Downstream Waters: A Review

We reviewed the scientific literature on non‐floodplain wetlands (NFWs), freshwater wetlands typically located distal to riparian and floodplain systems, to determine hydrological, physical, and chemical functioning and stream and river network connectivity. We assayed the literature for source, sink, lag, and transformation functions, as well as factors affecting connectivity. We determined NFWs are important landscape components, hydrologically, physically, and chemically affecting downstream aquatic systems. NFWs are hydrologic and chemical sources for other waters, hydrologically connecting across long distances and contributing compounds such as methylated mercury and dissolved organic matter. NFWs reduced flood peaks and maintained baseflows in stream and river networks through hydrologic lag and sink functions, and sequestered or assimilated substantial nutrient inputs through chemical sink and transformative functions. Landscape‐scale connectivity of NFWs affects water and material fluxes to downstream river networks, substantially modifying the characteristics and function of downstream waters. Many factors determine the effects of NFW hydrological, physical, and chemical functions on downstream systems, and additional research quantifying these factors and impacts is warranted. We conclude NFWs are hydrologically, chemically, and physically interconnected with stream and river networks though this connectivity varies in frequency, duration, magnitude, and timing.     

An inventory of wetland impoundments in the coastal zone of Louisiana,USA: Historical trends

We inventoried wetland impoundments in the Louisiana, USA, coastal zone from the late 1900s to 1985. Historically, impoundment of wetlands for reclamation resulted in direct wetland loss after levees (dikes) failed and the impounded area was permanently flooded, reverting not to wetland, but to open-water habitat. A current management approach is to surround wetlands by levees and water control structures, a practice termed semi-impoundment marsh management. The purpose of this semi-impoundment is to retard saltwater intrusion and reduce water level fluctuations in an attempt to reduce wetland loss, which is a serious problem in coastal Louisiana. In order to quantify the total impounded area, we used historic data and high-altitude infrared photography to map coastal impoundments. Our goal was to produce a documented inventory of wetlands intentionally impounded by levees in the coastal zone of Louisiana in order to provide a benchmark for further research. We inventoried 370,658 ha within the coastal zone that had been intentionally impounded before 1985. This area is equal to about 30% of the total wetland area in the coastal zone. Of that total area, approximately 12% (43,000 ha) is no longer impounded (i.e., failed impoundments; levees no longer exist or only remnants remain). Of the 328,000 ha still impounded, about 65% (214,000 ha) is developed (agriculture, aquaculture, urban and industrial development, and contained spoil). The remaining 35% (114,000 ha) of impoundments are in an undeveloped state (wetland or openwater habitat). In December 1985, approximately 50% (78,000 ha) of the undeveloped and failed impoundments were open-water habitat. This inventory will allow researchers to monitor future change in land-water ratios that occur within impounded wetlands and thus to assess the utility of coastal wetland management using impoundments.     

Spatial Relations Between Floodplain Environments and Land Use – Land Cover of a Large Lowland Tropical River Valley: Pánuco Basin, México

Large lowland river valleys include a variety of floodplain environments that represent opportunities and constraints for human activities. This study integrates extensive field observations and geomorphic data with analysis of satellite remote sensing data to examine spatial relations between land use/land cover (LULC) and floodplain environments in the lower Pánuco basin of eastern Mexico. The floodplain of the lower Pánuco basin was delineated by combining a digital elevation model with a satellite image of a large flood event. The LULC was classified by combining a hybrid classification strategy with image stratification, applied to 15-m-resolution ASTER data. A geomorphic classification of floodplain environments was performed using a dry-stage image (ASTER data) and a 1993 Landsat image acquired during a large flood event. Accuracy assessment was based on aerial photographs (1:38,000), global positioning satellite ground-truthing, and a Landsat 7ETM⁺ image from 2000, which resulted in an overall accuracy of 82.9% and a KHAT of 79.8% for the LULC classification. The geomorphic classification yielded 83.5% overall accuracy, whereas the KHAT was 81.5%. LULC analysis was performed for the entire floodplain and individually within four valley segments. The analysis indicates that the study area is primarily utilized for grazing and farming. Agriculture is primarily associated with coarse-grained (sandy/silty) natural levee and point bar units close to the river channel, whereas cattle grazing occurs in distal and lower-lying reaches dominated by cohesive fine-grained (clayey) deposits, such as backswamps. In the Pánuco valley, wetlands and lakes occur within backswamp environments, whereas in the Moctezuma segments, wetlands and lakes are associated with relict channels. This study reveals considerable variation in LULC related to spatial differences in floodplain environments and illustrates the importance of considering older anthropogenic influences on the landscape. The research design should be applicable for other large lowland coastal plain river valleys where agriculture is a major component of the floodplain landscape.     

Impacts of freshwater wetlands on water quality: A landscape perspective

In this article, we suggest that a landscape approach might be useful in evaluating the effects of cumulative impacts on freshwater wetlands. The reason for using this approach is that most watersheds contain more than one wetland, and effects on water quality depend on the types of wetlands and their position in the landscape. Riparian areas that border uplands appear to be important sites for nitrogen processing and retention of large sediment particles. Fine particles associated with high concentrations of phosphorus are retained in downstream wetlands, where flow rates are slowed and where the surface water passes through plant litter. Riverine systems also may play an important role in processing nutrients, primarily during flooding events. Lacustrine wetlands appear to have the least impact on water quality, due to the small ratio of vegetated surface to open water. Examples are given of changes that occurred when the hydrology of a Maryland floodplain was altered.     

THE LEVEE LOVE AFFAIR: A STORMY RELATIONSHIP?1

ABSTRACT: A history of flood control in the United States shows an undying affair with levees. This love affair, however, was put severely to the test by the record flooding in the summer of 1993. About 70 percent of levees in the upper Midwest failed during this time, leading to extensive damage to both farmland and urban areas. Consequently, there were repeated calls to re-assess the nation's floodplain management policies. The report of the Intera-gency Floodplain Management Review Committee is one outcome of this and it forms the basis of this commentary on levees. In many respects, levees are effective flood control measures, being relatively cheap to implement and easy to build. At the same time, levees have negative impacts, affecting the hydrological regime both up and down stream, and often exacerbating flooding in other places. Furthermore, technical weaknesses in design, planning, construction, and maintenance have all contributed to levee failures. While the report recommends changes in floodplain management to address some of these issues, it is difficult to see how these will materialize given the current political, economic, and social climate.     

Destruction of Wetlands and Waterbird Populations by Dams and Irrigation on the Murrumbidgee River in Arid Australia

The Lowbidgee floodplain is the Murrumbidgee Rivers major wetland in southeastern Australia. From more than 300,000 ha in the early 1900s, at least 76.5% was destroyed (58%) or degraded (18%) by dams (26 major storages), subsequent diversions and floodplain development. Diversions of about 2,144,000 ML year^–1 from the Murrumbidgee River come from a natural median flow of about 3,380,000 ML year^–1 providing water for Australias capital, hydroelectricity, and 273,000 ha of irrigation. Diversions have reduced the amount of water reaching the Lowbidgee floodplain by at least 60%, from 1888 to 1998. About 97,000 ha of Lowbidgee wetland was destroyed by development of the floodplain for an irrigation area (1975–1998), including building of 394 km of channels and 2,145 km of levee banks. Over 19 years (1983–2001), waterbird numbers estimated during annual aerial surveys collapsed by 90%, from an average of 139,939 (1983–1986) to 14,170 (1998–2001). Similar declines occurred across all functional groups: piscivores (82%), herbivores (87%), ducks and small grebe species (90%), large wading birds (91%), and small wading birds (95%), indicating a similar decline in the aquatic biota that formed their food base. Numbers of species also declined significantly by 21%. The Lowbidgee floodplain is an example of the ecological consequences of water resource development. Yanga Nature Reserve, within the Lowbidgee floodplain, conserved for its floodplain vegetation communities, will lose these communities because of insufficient water. Until conservation policies adequately protect river flows to important wetland areas, examples such as the Lowbidgee will continue to occur around the world.     

VALUE OF WOODY RWER CORRIDORS IN LEVEE PROTECTION ALONG THE MISSOURI RWER IN 19931

ABSTRACT: Following the Midwest flood of 1993, a study was initiated along a 39-mile segment of the Missouri River to determine if there was an association between woody corridors and levee stability. A systematic sample of levee failures revealed that primary levees which did not fail had a significantly wider woody corridor than failed levees. Analysis of the total inventory of failed levees revealed that as the width of the woody corridor decreased, the length of the levee failure increased. Number of levee failures and their severity of damage could be reduced if woody corridors were at least 300 feet wide.     

Hydrodynamic Modeling Improves Green River Reconnection with Floodplain Wetlands for Endangered Fish Species Recovery

We performed two‐dimensional (2D) hydrodynamic modeling to aid recovery of the endangered razorback sucker (Xyrauchen texanus) by reconnecting the Green River with its historic bottomland floodplain wetlands at Ouray National Wildlife Refuge, Utah. Reconnection allows spring flood flows to overtop the river levee every two to three years, and passively transport razorback sucker larvae to the wetlands to grow in critical habitat. This study includes (1) river hydrologic analysis, (2) simulation of a levee breach/weir, overtopping of river flood flows, and 2D flow through the wetlands using Hydrologic Engineering Center River Analysis System 2D, and (3) modeling flow and restoration scenarios. Indicators of hydrologic alteration were used to evaluate river flow metrics, in particular flood magnitudes, frequency, and duration. Results showed a target spring flow of 16,000 cfs (453 m³/s) and a levee breach elevation of 4,663 ft (1,421 m) amsl would result in a median flow >6,000 acre‐feet (7.4 million m³) over five days into the wetlands, which is adequate for razorback sucker larvae transport and rearing. Modeling of flow/restoration scenarios showed using gated water control structures and passive low‐water crossings between wetland units can provide adequate control of flow movement into and storage in multiple units. Levee breaching can be a relatively simple, cost‐effective method to reconnect rivers and historic floodplains, and hydrodynamic modeling is an important tool for analyzing and designing wetland reconnection.     

Evaluating cumulative effects on wetland functions: A conceptual overview and generic framework

This article outlines conceptual and methodological issues that must be confronted in developing a sound scientific basis for investigating cumulative effects on freshwater wetlands. We are particularly concerned with: (1) effects expressed at temporal and spatial scales beyond those of the individual disturbance, specific project, or single wetland, that is, effects occurring at the watershed or regional landscape level; and (2) the scientific (technical) component of the overall assessment process. Our aim is to lay the foundation for a research program to develop methods to quantify cumulative effects of wetland loss or degradation on the functioning of interacting systems of wetlands. Toward that goal we: (1) define the concept of cumulative effects in terms that permit scientific investigation of effects; (2) distinguish the scientific component of cumulative impact analysis from other aspects of the assessment process; (3) define critical scientific issues in assessing cumulative effects on wetlands; and (4) set up a hypothetical and generic structure for measuring cumulative effects on the functioning of wetlands as landscape systems.We provide a generic framework for evaluating cumulative effects on three basic wetland landscape functions: flood storage, water quality, and life support. Critical scientific issues include appropriate delineations of scales, identification of threshold responses, and the influence on different functions of wetland size, shape, and position in the landscape.The contribution of a particular wetland to landscape function within watersheds or regions will be determined by its intrinsic characteristics, e.g., size, morphometry, type, percent organic matter in the sediments, and hydrologic regime, and by extrinsic factors, i.e., the wetland's context in the landscape mosaic. Any cumulative effects evaluation must take into account the relationship between these intrinsic and extrinsic attributes and overall landscape function. We use the magnitude of exchanges among component wetlands in a watershed or larger landscape as the basis for defining the geographic boundaries of the assessment. The time scales of recovery for processes controlling particular wetland functions determine temporal boundaries. Landscape-level measures are proposed for each function.     

Protecting River Corridors in Vermont1

Kline, Michael and Barry Cahoon, 2010. Protecting River Corridors in Vermont. Journal of the American Water Resources Association (JAWRA) 46(2):227-236. DOI: 10.1111/j.1752-1688.2010.00417.x Abstract: The Vermont Agency of Natural Resources’ current strategy for restoring aquatic habitat, water quality, and riparian ecosystem services is the protection of fluvial geomorphic-based river corridors and associated wetland and floodplain attributes and functions. Vermont has assessed over 1,350 miles of stream channels to determine how natural processes have been modified by channel management activities, corridor encroachments, and land use/land cover changes. Nearly three quarters of Vermont field-assessed reaches are incised limiting access to floodplains and thus reducing important ecosystem services such as flood and erosion hazard mitigation, sediment storage, and nutrient uptake. River corridor planning is conducted with geomorphic data to identify opportunities and constraints to mitigating the effects of physical stressors. Corridors are sized based on the meander belt width and assigned a sensitivity rating based on the likelihood of channel adjustment due to stressors. The approach adopted by Vermont is fundamentally based on restoring fluvial processes associated with dynamic equilibrium, and associated habitat features. Managing toward fluvial equilibrium is taking hold across Vermont through adoption of municipal fluvial erosion hazard zoning and purchase of river corridor easements, or local channel and floodplain management rights. These tools signify a shift away from primarily active management approaches of varying success that largely worked against natural river form and process, to a current community-based, primarily passive approach to accommodate floodplain reestablishment through fluvial processes.     

MISSOURI RIVER FLOOD OF 1993: ROLE OF WOODY CORRIDOR WIDTH IN LEVEE PROTECTION1

ABSTRACT. We investigated the relationships between levee damage and woody corridor along a 353‐mile segment of the Missouri River in Missouri during the flood of 1993. Results indicated that woody corridors between riverbanks and primary levees played a significant role in the reduction or prevention of flood related damage to levees. Forty‐one percent of levee failures in this segment occurred in areas with no woody corridor, while 74 percent and 83 percent of failures occurred where woody corridor widths were less than 300 feet and less than 500 feet, respectively. Median failure lengths with a woody corridor present were 50.3 percent shorter than median failure lengths with no woody corridor present. Levees without failures had significantly wider median woody corridor widths than levees that failed. Eligibility for the Corps of Engineers levee maintenance program was not a significant factor in the reduction of levee damage. Discontinuities in woody corridors played a role in 27.5 percent of the levee failures in the study segment. Smaller segments of the river valley were studied to determine if geomorphic differences influenced variations in the protective value of woody corridors.     

Land loss in coastal Louisiana (U.S.A.)

This paper examines causes and consequences of wetland losses in coastal Louisiana. Land loss is a cumulative impact, the result of many impacts both natural and artificial. Natural losses are caused by subsidence, decay of abandoned river deltas, waves, and storms. Artificial losses result from flood-control practices, impoundments, and dredging and subsequent erosion of artificial channels. Wetland loss also results from spoil disposal upon wetlands and land reclamation projects.Total land loss in Louisiana's coastal zone is at least 4,300 ha/year. Some wetlands are converted to spoil banks and other eco-systems so that wetland losses are probably two to three times higher. Annual wetland losses in the Barataria Bay basin are 2.6% of the wetland area. Human activities are the principal determinants of land loss. The present total wetland area directly lost because of canals may be close to 10% if spoil area is included. The interrelationship between hydrology, land, vegetation, substrate, subsidence, and sediment supply are complicated; however, hydrologic units with high canal density are generally associated with higher rates of land loss and the rate may be accelerating.Some cumulative impacts of land loss are increased saltwater intrusion, loss of capacity to buffer the impact of storms, and large additions of nutrients. One measure of the impact is that roughly $8–17 × 10⁶ (U.S.A.) of fisheries products and services are lost annually in Louisiana.Viewed at the level of the hydrologic unit, land loss transcends differences in local vegetation, substrate, geology, and hydrology. Land management should therefore focus at that level of organization. Proper guideline recommendations require an appreciation of the long-term interrelations of the wetland estuarine system.     

MULTIVARIATE CLASSIFICATION OF SMALL ORDER WATERSHEDS IN THE QUABBIN RESERVOIR BASIN,MASSACHUSEVFS1

ABSTRACT: A multivariate approach was used to analyze hydrologic, geologic, geographic, and water-chemistry data from small order watersheds in the Quabbin Reservoir Basin in central Massachusetts. Eighty three small order watersheds were delineated and landscape attributes defining hydrologic, geologic, and geographic features of the watersheds were compiled from geographic information system data layers. Principal components analysis was used to evaluate 11 chemical constituents collected bi-weekly for 1 year at 15 surface-water stations in order to subdivide the basin into subbasins comprised of watersheds with similar water quality characteristics. Three principal components accounted for about 90 percent of the variance in water chemistry data. The principal components were defined as a biogeochemical variable related to wet. land density, an acid-neutralization variable, and a road-salt variable related to density of primary roads. Three subbasins were identified. Analysis of variance and multiple comparisons of means were used to identify significant differences in stream water chemistry and landscape attributes among subbasins. All stream water constituents were significantly different among subbasins. Multiple regression techniques were used to relate stream water chemistry to landscape attributes. Important differences in landscape attributes were related to wetlands, slope, and soil type.     

Flood Risk Perceptions and Overdevelopment in the Floodplain1

ABSTRACT: There is a long standing hypothesis that overdevelopment has occurred in the nation's floodplains due to imperfect information about the potential flood hazard, an expectation of disaster relief and anticipation of future structural protection. This hypothesis is investigated with multiple regression analysis of data for a case study area. In particular the question of whether floodplain residential property values are fully discounted for expected flood damages is addressed by considering the impact of the National Flood Insurance Program on property values. The extent to which flooding risk perceptions are based on low cost information such as distance from and elevation above the river is also considered. Finally, implications for floodplain management policy are discussed.     

Diversion of water flow from a floodplain wetland stream: an analysis of geomorphological setting and hydrological and ecological consequences

Diversion of water has been ongoing in the Mkuze Wetland for several decades. Two canals form the focus of this study; the Mpempe-Demazane Canal and the Tshanetshe Canal. The former involved an ambitious excavation over a distance of 13.5 km in the lower part of the wetland, while the latter was a minor excavation over a distance of approximately 100 m in the upper part of the wetland. Although ambitious and costly, the Mpempe-Demazane Canal resulted in little downward or headward erosion, and there was minor diversion of flow. However, the minor excavation of the Tshanetshe Canal resulted in erosion downstream of the excavation (the Tshanetshe Stream), downward and lateral erosion of the excavated section, and headward erosion that has propagated almost 4 km upstream along the Mkuze River. Most of the flow of the Mkuze River has been captured by the Tshanetshe Canal and Stream. The impact of canalisation on floodplain wetlands is thus more dependent on the location than the scale of activity. The avulsion of the Mkuze River into the Tshanetshe Canal and Stream is due to a large difference in elevation between the Mkuze River and floodplain into which it was diverted, and the fact that in this region the river typically has high discharges. This avulsion may have been inevitable as a result of natural processes of sedimentation. In contrast, the difference in elevation between the Mkuze River and the basin into which it was diverted via the Mpempe Canal was small as is discharge of the Mkuze River in this part of the wetland. Thus, the diversion was unsuccessful. The presence of hippos that create hydraulically efficient pathways that are oriented parallel to the regional hydraulic slope, may accelerate avulsion in large African wetlands. Overall, it is argued that the environmental consequences of excavation need to be viewed against the background that wetlands are dynamic features within the landscape.     

HARVEST INFLUENCES ON FLOODWATER PROPERTIES IN A FORESTED FLOODPLAIN1

ABSTRACT: Floodplain forests directly influence water quality by serving as sinks, sources, or transformers of nutrients. Increases in the demand for timber raise the question of how silvicultural disturbance may affect this function. The objective of this research was to compare biogeochemical relationships between undisturbed vs. disturbed conditions in a floodplain forest. A randomized complete block design consisting of three blocks and two treatments (partial harvest and undisturbed) was installed on the Flint River floodplain, Georgia. The partial cut was conducted during September-October 1993. Automated water samplers were situated to sample during flood events as sheet flow entered and exited treatment plots during the 1994, 1995 and 1996 flood seasons. Prevs. post-contact comparisons indicated that the undisturbed floodplain has minimal influence on water chemistry at this scale of measurement. Although the partial harvest on an 8-ha scale had minimal effect upon sheetfiow water chemistry for three years following harvest, the data suggest that harvests may stimulate a minor increase in Ca and K sink activity.     

Strategies for assessing the cumulative effects of wetland alteration on water quality

Assessment of cumulative impacts on wetlands can benefit by recognizing three fundamental wetland categories: basin, riverine, and fringe. The geomorphological settings of these categories have relevance for water quality.Basin, or depressional, wetlands are located in headwater areas, and capture runoff from small areas. Thus, they are normally sources of water with low elemental concentration. Although basin wetlands normally possess a high capacity for assimilating nutrients, there may be little opportunity for this to happen if the catchment area is small and little water flows through them.Riverine wetlands, in contrast, interface extensively with uplands. It has been demonstrated that both the capacity and the opportunity for altering water quality are high in riverine wetlands.Fringe wetlands are very small in comparison with the large bodies of water that flush them. Biogeochemical influences tend to be local, rather than having a measurable effect on the larger body of water. Consequently, the function of these wetlands for critical habitat may warrant protection from high nutrient levels and toxins, rather than expecting them to assume an assimilatory role.The relative proportion of these wetland types within a watershed, and their status relative to past impacts can be used to develop strategies for wetland protection. Past impacts on wetlands, however, are not likely to be clearly revealed in water quality records from monitoring studies, either because records are too short or because too many variables other than wetland impacts affect water quality. It is suggested that hydrologic records be used to reconstruct historical hydroperiods in wetlands for comparison with current, altered conditions. Changes in hydroperiod imply changes in wetland function, especially for biogeochemical processes in sediments. Hydroperiod is potentially a more sensitive index of wetland function than surface areas obtained from aerial photographs. Identification of forested wetlands through photointerpretation relies on vegetation that may remain intact for decades after drainage. Finally, the depositional environment of wetlands is a landscape characteristic that has not been carefully evaluated nor fully appreciated. Impacts that reverse depositional tendencies also may accelerate rates of change, causing wetlands to be large net exporters rather than modest net importers. Increases in rates as well as direction can cause stocks of materials, accumulated over centuries in wetland sediments, to be lost within decades, resulting in nutrient loading to downstream aquatic ecosystems.     

Hydrologic Connectivity and the Contribution of Stream Headwaters to Ecological Integrity at Regional Scales1

Abstract: Cumulatively, headwater streams contribute to maintaining hydrologic connectivity and ecosystem integrity at regional scales. Hydrologic connectivity is the water‐mediated transport of matter, energy and organisms within or between elements of the hydrologic cycle. Headwater streams compose over two‐thirds of total stream length in a typical river drainage and directly connect the upland and riparian landscape to the rest of the stream ecosystem. Altering headwater streams, e.g., by channelization, diversion through pipes, impoundment and burial, modifies fluxes between uplands and downstream river segments and eliminates distinctive habitats. The large‐scale ecological effects of altering headwaters are amplified by land uses that alter runoff and nutrient loads to streams, and by widespread dam construction on larger rivers (which frequently leaves free‐flowing upstream portions of river systems essential to sustaining aquatic biodiversity). We discuss three examples of large‐scale consequences of cumulative headwater alteration. Downstream eutrophication and coastal hypoxia result, in part, from agricultural practices that alter headwaters and wetlands while increasing nutrient runoff. Extensive headwater alteration is also expected to lower secondary productivity of river systems by reducing stream‐system length and trophic subsidies to downstream river segments, affecting aquatic communities and terrestrial wildlife that utilize aquatic resources. Reduced viability of freshwater biota may occur with cumulative headwater alteration, including for species that occupy a range of stream sizes but for which headwater streams diversify the network of interconnected populations or enhance survival for particular life stages. Developing a more predictive understanding of ecological patterns that may emerge on regional scales as a result of headwater alterations will require studies focused on components and pathways that connect headwaters to river, coastal and terrestrial ecosystems. Linkages between headwaters and downstream ecosystems cannot be discounted when addressing large‐scale issues such as hypoxia in the Gulf of Mexico and global losses of biodiversity.