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1.
A study on the modern dynamics and shoreline changes from 1954 to 2014 of the Molise coast (central Adriatic Sea) has been carried out. Short to long-term shoreline changes and associated surface area variations have been assessed in GIS environment for the study coast, subdivided in nine coastal segments (S1-S9), by using 100-m regularly spaced transects. In addition, the possible influence of natural and anthropogenic factors, especially of climatic variability and engineered shoreline defense structures, has been investigated. The Molise coast has experienced notable long-term erosion (period 1054-2014) that caused an overall coastal land loss of approximately 940,000 m2. Erosion was, yet, limited to coastal segments S1 and S7, nearest to the mouths of major rivers, namely Trigno and Biferno, while the major part of the study coast has remained essentially stable or even advanced. Increased shoreline protection by defense structures has generally favoured shoreline stability and frequently generated shoreline advance, except for segments S1 and S7. Observed differences in shoreline change rates over time at the decadal to interannual scale, have not find a response in the analysis of available data on meteo-marine conditions of the Molise coast and climate variability indices, pointing out the need to improve knowledge on meteomarine conditions and on climatic variability forcing of the study area. From 2004 to 2014, the Molise shoreline remained essentially stable. Nonetheless, most recent shoreline changes (period 2011-2014) and modern shoreline dynamics indicate that erosion has become more widespread, involving at least part of segments S2-S3 and S8-S9, located south of the river mouth segments. The localized long-term shoreline retreat and most recent shoreline erosion appear to be primarily related to channel adjustments of the Biferno and Trigno rivers that occurred since the 1950s under the control of human interventions on the rivers, especially the construction respectively of a dam and a check dam along their lower courses, that trap of most of their solid load, affecting so adversely the sediment budget of the river mouths areas and adjacent beaches. Overall data acquired on the recent shoreline evolution and modern shoreline dynamics of the Molise coast and on related causal factors provide a good basic knowledge for regional coastal management purposes, and for further scientific purposes. Particularly, they suggest the opportunity to deepen a number of aspects such as the relationship between the coast and river catchments feeding it, the possible influences on the Molise shoreline dynamics of the neighbouring coasts, the efficiency/obsolescence of defense structures and the present-day vulnerability to coastal erosion of the Molise coast.  相似文献   

2.
Abstract: The Atlantic rainforest of southern Bahia is one of the last remnants of the lowland forest of eastern Brazil that once covered the entire coastal area from Rio Grande do Norte to Rio Grande do Sul ( lat 8°–28° S) and has been deforested to a small fraction of its original cover (1–12%). All recent vegetation surveys have been based on optical satellite data, which is hampered by cloud cover and by southern Bahia's intricate mix of forest patches with other tree crops, especially cocoa. We describe the application of radar remote-sensing data to distinguish forest patches from cocoa planted in the shade of natural-forest trees. Radar, unlike optical sensors, is not obstructed by cloud cover and can acquire information about forest structure by penetrating into the vegetation canopy. The vegetation map generated from radar data clearly separates forest patches based on the degree of structural disturbance such as the density of shaded trees, the openness of the canopy, and the density of the monodominant Erythyrina shaded trees. The structural classification based on the radar data, and shown on the map, can help researchers assess the degree of fragmentation of the original Atlantic coastal forest and delineate areas of less disturbance with higher potential for conservation of biodiversity. This information can then be applied to conservation planning, especially the design and monitoring of nature reserves and the modeling of biological corridors.  相似文献   

3.
Landsat satellite imagery was analyzed to generate a detailed record of 10 years of vegetation disturbance and regrowth for Pacific coastal areas of San Mateo and Santa Cruz Counties. The Landsat Ecosystem Disturbance Adaptive Processing System (LEDAPS) methodology, a transformation of Tasseled-Cap data space, was applied to detected changes in perennial coastal shrubland, woodland, and forest cover from 1999 to 2009. Results showed several principal points of interest, within which extensive contiguous areas of similar LEDAPS vegetation change (either disturbed or restored) were detected. Regrowth of evergreen shrub and tree cover was prevalent along the several long stretches of the coast highway (CA Route 1) between the cities of Half Moon Bay and Santa Cruz. A number of state parks areas showed measurable vegetation restoration as well. The most prominent loss of perennial coastal vegetation over decade was in the Pescadero Marsh area, where the continued presence of levees has historically reduced flood conveyance capacity into and through the marshlands. Based on these examples, the LEDAPS methodology was determined to be capable of fulfilling much of the need for continual, low-cost monitoring of emerging changes to coastal ecosystems.  相似文献   

4.
Encroachment of tall grasses and shrubs in coastal dunes has resulted in loss of vegetation heterogeneity. This is expected to have negative effects on animal diversity. To counteract encroachment and develop structural heterogeneity grazing is a widely used management practice. Here, we aim to functionally interpret changes in vegetation composition and configuration following grazing management on habitat suitability for sand lizards. Aerial photographs taken over a period of 16 years were used to quantify changes in vegetation composition. A GIS-based method was developed to calculate habitat suitability for sand lizards in a spatially explicit manner, encompassing differences in vegetation structure and patch size. From 1987 to 2003 dune vegetation shifted from small patches of moss and sand to larger patches covered by shrubs and grasses. Grazing management did not have any significant effect on the overall level of heterogeneity, measured as habitat suitability for sand lizards. However, on a more local scale highly suitable patches in 1987 were deteriorating whereas unsuitable patches became more suitable in 2003. This inversion results from a broad shift with shrubs being a limiting habitat element in 1987 to sandy patches being the limiting element in 2003. Future changes are believed to negatively impact sand lizards. The habitat suitability model has proven to be a useful tool to functionally interpret changes in coastal dune vegetation heterogeneity from an animal’s perspective. Further research should aim to include multiple species operating on different scale levels to fully capture the natural landscape dynamics.  相似文献   

5.
The initial recovery of vegetation after a wildfire in a coastal dune area in NW Jutland, Denmark, was studied over a 5-yr period by means of permanent plots representing various dune communities along a topographical gradient. The impact of the fire varied with the position of the plots. Fens and south-facing dunes were little affected while dune heath plots were severely affected including loss of the O-horizon. Post-fire conditions included presence of remaining soil organic matter, a soil seed bank and surviving below-ground plant parts. The soil surface remained stable during the study period. The initial five years of recovery comprised of an initial three-year recruitment phase during which cover and number of species increased and the quantitative species composition changed markedly, followed by two years of a declining rate of change. 38 species of vascular plants were recorded, 35 are regular components in dune, dune heath and heath fen and were recruited from the seed bank, from locally dispersed seeds and/or by sprouting from surviving vegetative parts. The remaining three species were ‘aliens’, dispersed from sources outside the area. Crustose lichens had an important role in the initial recovery by stabilizing the surface and probably inhibiting seed germination, whereas mosses mostly had a subordinate role. The seral position of the plots, as well as the expected time needed for full recovery of pre-fire vegetation, vary with topography and initial soil conditions. Five years after the fire the fen and the south-facing dune probably need less than a decade for full recovery. The remaining plots are judged to be relatively early seral; their full recovery into mature dry or moist dune heath vegetation and O-horizon is expected to need several centuries.  相似文献   

6.
Climate change and engineering activities have modified the hydrology and morphology of estuaries. However, the potential effects of these modifications on vegetation succession in estuarine marshes are still poorly understood. Therefore, we studied temporal changes in tidal habitats of the Elbe estuary over a period of 30 years. We compared vegetation maps from 1980 to 2010 and calculated the change in area of habitats with respect to three salinity and three elevational zones. To analyze the direction of the temporal change, we differentiated between progressive and regressive succession. By using regression tree models (conditional inference trees), we identified the most influential factors determining progressive or regressive succession of low marshes. The total area of the estuarine tidal marshes at the Elbe increased by 2 % from 1980 to 2010, but changes were unequal among the salinity zones. In the salt and brackish zones, the area covered by high marshes increased substantially but decreased in the tidal freshwater zone, while that covered by low marshes decreased in all the salinity zones. Additionally, we determined high persistence of tidal flats and high marshes, whereas only 19 to 28 % of the low marshes found in 1980 remained in 2010. In salt and brackish marshes, more than two-thirds of the area that had been identified as low marshes in 1980 had progressively developed into high marshes. In contrast, 44 % of the area of low marshes in tidal freshwater marshes showed regressive succession back into tidal flats. The distance to the navigation channel was the main factor determining successional direction in salt and brackish marshes. Here, greater proximity to the channel was correlated with higher risk of regressive succession. In tidal freshwater marshes, we identified both the distance to the navigation channel and the situation on the river shore (i.e. inner bank, outer bank or straight bank) as the main factors for marsh succession. Here, considerable engineering activities in the channel had simultaneously decreased the mean low water level and increased the mean high water level between 1980 and 2010, which led to an increase in tidal amplitude. It is quite likely that these changes negatively modified marsh distribution, increased regressive succession and, thus, lowered the quality of tidal freshwater marshes.  相似文献   

7.
Landsat satellite imagery was analyzed to generate a detailed record of 10 years of vegetation disturbance and regrowth for Pacific coastal areas of Marin and San Francisco Counties. The Landsat Ecosystem Disturbance Adaptive Processing System (LEDAPS) methodology, a transformation of Tasseled-Cap data space, was applied to detected changes in perennial coastal shrubland, woodland, and forest cover from 1999 to 2009. Results showed several principal points of interest, within which extensive contiguous areas of similar LEDAPS vegetation change (either disturbed or restored) were detected. Regrowth areas were delineated as burned forest areas in the Point Reyes National Seashore (PRNS) from the 1995 Vision Fire. LEDAPS-detected disturbance patterns on Inverness Ridge, PRNS in areas observed with dieback of tanoak and bay laurel trees was consistent with defoliation by sudden oak death (Phytophthora ramorum). LEDAPS regrowth pixels were detected over much of the predominantly grassland/herbaceous cover of the Olema Valley ranchland near PRNS. Extensive restoration of perennial vegetation cover on Crissy Field, Baker Beach and Lobos Creek dunes in San Francisco was identified. Based on these examples, the LEDAPS methodology will be capable of fulfilling much of the need for continual, low-cost monitoring of emerging changes to coastal ecosystems.  相似文献   

8.
以MODIS遥感影像数据为数据源,应用遥感和GIS技术对泾河流域植被覆盖格局进行分析。结果表明:(1)泾河流域植被覆盖类型以退化草地和草地为主,二者占流域总面积的53.30%;地面覆盖程度较高的森林、疏林仅占流域总面积的9.83%,流域植被覆盖程度较低。(2)受自然环境条件和人类活动的影响,退化草地、稀疏灌丛、郁闭灌丛和疏林破碎化程度较为严重,对地面覆盖程度最低的退化草地除集中分布于流域北部外,主要零散分布于流域中、南部,其斑块形状复杂多样,数量最多,应成为流域植被恢复的重点。(3)各植被覆盖类型的斑快数量组成以小斑块所占比例最大,最小斑块比率均在50.0%以上;森林的斑块面积多样性最大,草地的斑块面积多样性最小。  相似文献   

9.
Adult sockeye salmon (Oncorhynchus nerka, N = 179) from six Fraser River populations (British Columbia) were intercepted in continental shelf waters ∼215 km from the Fraser River mouth, gastrically implanted with acoustic transmitters, non-lethally biopsied for blood biochemistry, gill Na+,K+-ATPase activity and somatic energy density and then released. Migration behaviour and travel times to the river mouth and into the river were monitored by underwater telemetry receivers positioned at the river mouth and in the river. Post-release survival of salmon was excellent, with 84% (N = 150) of fish reaching the furthest receiving station ∼85 km upriver. Fish from Late-summer run populations (Adams and Weaver Creek) averaged a migration rate of ∼20 km day−1 through the marine area and held at the river mouth and adjacent areas for 7–9 days before entering the river. Summer-run populations (Birkenhead, Chilko, Horsefly and Stellako) had a migration rate ∼33 km day−1 and held for only 1–3 days. Once in river, similar patterns were observed: Late-run populations migrated at ∼28 km day−1 and Summer-run populations at ∼40 km day−1. From point of release to the river mouth, males migrated faster than females, but once in river migration rates did not differ between sexes. Among all females, a correlation was discovered between levels of circulating testosterone and river entry timing. This correlation was not observed among males. Plasma K+, Cl, glucose, lactate and osmolality were also correlated with entry timing in both sexes.  相似文献   

10.
Analysis of historical cartography, wind direction data, orientation of Volturno River mouth and shoreline changes through time point out a series of morphological changes due both to natural and man-induced processes, which occurred along the Domitia coast (Litorale Domitio) since the second half of the 50s. Meteorological time series analysis showed evidence of a change in wind frequency and direction before and after the second half of the 70s, with almost complete disappearance of wind calm periods. Particularly, after 1975, the angle of wave attack had changed and intensity and frequency increased, thus accelerating coastal erosion in some areas. The morphological response to these environmental changes, likely due to global warming and short climatic crises interacting with anthropic modifications of landscape, resulted in different orientation of the Volturno River mouth which rotated from NW (between the 50s and 70s) to SW (from the mid-70s until the present). Moreover, the NW-SE littoral drift caused a clockwise rotation of the entire coastal segment with fulcrum pole close to the river mouth, since the 50s. These findings may allow the assessment of land-use scenarios tuned with evolution trends in order to properly address planning of these reactive and dynamic landscapes.  相似文献   

11.
At three coastal dune sites at the island of Hiddensee, north-east Germany, vegetation cover was mapped during 2002 and compared to vegetation surveys from the late 1980s and 1930s. Abiotic and biotic factors, which have been identified as being critical for coastal dunes in former studies such as disturbance, salt spray or nutrient availability, were measured. Grazing and land-use history were reviewed by literature and interviews. Tall graminoid communities, mainly Carex arenaria, are a common vegetation unit today. Development, distribution of these dominances and possible causes for its occurrence have not been analysed. Generally, older successional vegetation units increased and pioneer stages decreased from the 1930s until 2002. At the geologically youngest site, the southern dunes, grass encroachment by Carex arenaria was highest (ca. 50% cover in 2002), and age and density of trees lower than at the older, central dunes. Land-use changes such as decrease in grazing pressure, additional feeding of livestock, increase in coastal protection measures and subsequent decrease in shifting sands as well as varying availability of groundwater and amount of salt spray are relevant factors for vegetation changes in coastal dunes over the past 70 years. Site-specific land-use differences such as livestock density and land-use history have a stronger influence than atmospheric N-pollution on the vegetation composition of these acidic, coastal dunes under low to moderate N-deposition loads of 6–8 kg N ha−1 yr−1.  相似文献   

12.
论城市河道整治与景观生态设计   总被引:2,自引:0,他引:2  
针对城市河道整治中存在的一些问题,阐述了城市河道整治的原则,提出城市河道整治要考虑景观生态,设计中要遵循安全性、自然性、生态性、观赏性、亲水性、文化性等原则。在此基础上,重点介绍了河道形态景观、堤岸景观和河道附属景观的设计内容。  相似文献   

13.
The coastal fringe of the eastern part of the Rhône delta is a large area where activities related to ecological and economical interests have increased since the beginning of the century, including the salt industry of theCompagnie des Salins du Midi et des Salines de l’Est, the industrial complex ofFos-sur-Mer, tourism managed by the local authorities ofPort-Saint-Louis andArles, and dune restoration of the La Gracieuse spit undertaken by thePort Autonome de Marseille. The vulnerability of the area for changes brought about by these activities is linked to the extent of several coastal changes, whereas these changes are related to the physical functioning of the coastal fringe, such as coastal processes, but also to external components such as relative sea-level rise, subsidence, reduction of sediment input, human impact, etc. For many decades, coastline ‘protection’ in the eastern part of the Rhône delta has been undertaken to withstand erosional processes that offset beaches and retreat of the littoral. The use of different engineering structures illustrates the heterogeneity and differences in age of the various management policies. At the eastern side of the Rhône, thePort Autonome de Marseille, dealing with management, has opted for soft engineering structures using fences for dune restoration on the La Gracieuse spit. At the western side of the mouth, theCompagnie des Salins du Midi et des Salines de l’Est has been using hard engineering structures to combat erosion (rocks, groins, tetrapods, etc.). A brief assessment of the management efficiency is presented, including an analysis of the data collected concerning the survey of the zones where the structures were built. Their varying efficiency shows the important role played by the sediment supply to the littoral (solid river discharge) and the coastal hydro-sedimentary processes. In the eastern part of the Rhône river the success of the dune restoration is mainly due to the important sediment supply transported in this area by eastern drift currents. At the western part of the mouth, the decrease of sediment supply linked to erosional processes in the shallow off-shore beach has made the structures more vulnerable, and therefore, the activities of the salt industry. For this last threatened zone, three management scenarios are discussed by taking into account economical, physical and ecological parameters.  相似文献   

14.
The aim of this study was to investigate the variation of channel bed roughness in two rivers, as important parameter in hydraulic modelling especially with regard to flood control. The universities of Ghent (UG) and Antwerp (UA) are conducting scientific research in the river Aa in Belgium and the Biebrza river in Poland in order to better understand the phenomena involved and to come to a more accurate determination of the different parameters influencing flow. In this paper, the determination of the roughness coefficient ‘n’ from the Manning equation is used. This coefficient is not easy to determine and is varying constantly. It is influenced by the meandering character of the river, the bed material and the average grain size, the channel bed forms, the channel obstructions, the geometry changes between sections and the vegetation in the channel. Furthermore, due to these parameters, the roughness of the channel is not equally distributed over the channel, the banks and the floodplains. So, using literature data does not always lead to satisfactory results, due to the different situation in the field (Werner et al. J Hydrol 314:139–157, 2005). Therefore, measurements are necessary to determine the variation of the Manning coefficient. The Manning coefficient is a function of the discharge, but will also vary over the time due to the mentioned influences. In a multidisciplinary research project on the fundamental exchange processes in river ecosystems, hydraulic measurements were performed on a regular base in the river Aa. During these measurement campaigns, velocity and discharge measurements were carried out in multiple cross-sections. Once a month, the discharge and the water levels were measured at the upstream and the downstream end of the test stretch. On the river Biebrza, similar intensive measurement campaigns took place along a 6 km stretch in the upstream part of the river. An accurate determination of the Manning coefficient according a seasonal variation is an important tool in hydraulic modelling.  相似文献   

15.
In 1990, grazing was introduced in a section of Meijendel, a coastal sand dune system near The Hague, The Netherlands. After five years an evaluation was made of the effects of grazing on vegetation development. Three transects were established, two in grazed areas and one in an ungrazed area. Field survey data were classified by means of TWIN-SPAN, ordinated with Detrended Correspondence Analysis and the resulting vegetation types interpreted according to Westhoff & den Held (1969). All associations were found in both the grazed and the ungrazed areas, but at the subassociation and variant level some communities appeared to be restricted to the grazed area. These variants were five grassland variants characterized by disturbance indicators such asSenecio sylvaticus andCynoglossum officinale. The total number of plant species in the 19 permanent plots, which had been observed to have been decreasing since 1960, showed a considerable increase after the introduction of horses and cows in 1990. A marked decrease in the cover ofCalamagrostis epigejos andCarex arenaria since 1990 was evident, while in some plots species such asRibes rubrum andViburnum opulus increased considerably. A series of false-colour aerial photographs were used to compare vegetation structure in the three transects between 1990 and 1995. In the grazed area the tall grass vegetation had almost totally disappeared, whereas the areas of open sand. sand with moss and lichens, and low grass vegetation had increased and the pattern had become more fine-grained. In the ungrazed area the area covered by low grass vegetation had increased at the expense of the area of sand with moss and lichens and the pattern had become more coarse-grained.  相似文献   

16.
Wetlands as an alternative stable state in desert streams   总被引:2,自引:0,他引:2  
Heffernan JB 《Ecology》2008,89(5):1261-1271
Historically, desert drainages of the American southwest supported productive riverine wetlands (ciénegas). Region-wide erosion of ciénegas during the late 19th and early 20th century dramatically reduced the abundance of these ecosystems, but recent reestablishment of wetlands in Sycamore Creek, Arizona, USA, provides an opportunity to evaluate the mechanisms underlying wetland development. A simple model demonstrates that density-dependent stabilization of channel substrate by vegetation results in the existence of alternative stable states in desert streams. A two-year (October 2004-September 2006) field survey of herbaceous cover and biomass at 26 sites located along Sycamore Creek is used to test the underlying assumption of this model that vegetation cover loss during floods is density dependent, as well as the prediction that the distribution of vegetation abundance should shift toward bimodality in response to floods. Observations of nonlinear, negative relationships between herbaceous biomass prior to flood events and the proportion of persistent vegetation cover were consistent with the alternative stable state model. In further support of the alternative-state hypothesis, vegetation cover diverged from an approximately normal distribution toward a distinctly bimodal distribution during the monsoon flood season of 2006. These results represent the first empirically supported example of alternative-state behavior in stream ecosystems. Identification of alternative stable states in desert streams supports recent hypotheses concerning the importance of strong abiotic-disturbance regimes and biogeomorphic mechanisms in multiple-state ecosystems.  相似文献   

17.
Soil properties are well known to affect vegetation, but the role of soil heterogeneity in the patterning of vegetation dynamics is poorly documented. We asked whether the location of an ecotone separating grass-dominated and sparsely vegetated areas reflected only historical variation in degradation or was related to variation in inherent soil properties. We then asked whether changes in the cover and spatial organization of vegetated and bare patches assessed using repeat aerial photography reflected self-organizing dynamics unrelated to soil variation or the stable patterning of soil variation. We found that the present-day ecotone was related to a shift from more weakly to more strongly developed soils. Parts of the ecotone were stable over a 60-year period, but shifts between bare and vegetated states, as well as persistently vegetated and bare states, occurred largely in small (<40 m2) patches throughout the study area. The probability that patches were presently vegetated or bare, as well as the probability that vegetation persisted and/or established over the 60-year period, was negatively related to surface calcium carbonate and positively related to subsurface clay content. Thus, only a fraction of the landscape was susceptible to vegetation change, and the sparsely vegetated area probably featured a higher frequency of susceptible soil patches. Patch dynamics and self-organizing processes can be constrained by subtle (and often unrecognized) soil heterogeneity.  相似文献   

18.
塔里木河下游生态输水过程中荒漠河岸林活力恢复监测   总被引:4,自引:0,他引:4  
根据近5年来对塔里木河下游荒漠河岸林植被的监测数据,分析了生态输水后植物活力的恢复状况.结果显示:应急生态输水增加了塔里木河下游的生物多样性,使原本面临死亡的荒漠河岸植被重新复活,而且不同程度上促进了胡杨群落的自然更新;在近河道50 m的范围内均出现了少量的胡杨、柽柳实生苗,并且在离河道150 m的范围内已经有相当数量的胡杨次生苗,在离河道400 m范围内大约有25%的胡杨均有不同程度的基部新枝萌蘖.通过生态输水后地下水位的逐步抬升,河道两岸的低阶地发育着一定面积的草甸植被,形成了由胡杨、柽柳和草本植物所组成的干旱区非地带性河岸稀疏植被群落,说明应急生态输水对于胡杨为建群种的荒漠河岸林植被的恢复和自然更新产生了积极的影响.  相似文献   

19.
Coastal barrier environments are heavily influenced by human activities yet there are few examples of landscape ecological work investigating human dimensions of settlement disturbance patterns and processes. We investigated the impacts of residential development on vegetation cover for a remote roadless coastal barrier in Carova, North Carolina that is subject to policies from the federal to local levels and addressed three research questions: How has the region’s the policy history influenced patterns of residential parcel development? What are the spatial and temporal patterns of parcel development? How has development impacted patterns of barrier vegetation cover? We traced the influences of the federal 1982 Coastal Barrier Resources Act (CBRA) designed to discourage development in risky coastal areas as well as state/local coastal policies and employed remote sensing change detection, NDVI analysis and spatial analysis and regression techniques. Results showed an acceleration of new housing structures since 1990, contrary to the intended effects of CBRA. An estimated vegetation cover loss of 437 m2 was associated with each newly developed parcel. NDVI varied along spatial and temporal gradients with more recent development having lower NDVI than older development. Recently developed parcels were larger in area, closer to the beach, and contained houses with larger footprints compared to older developed parcels. Our approach represents a place-based analytical framework for coastal barrier landscapes. Beyond the Carova case study, adopting such an approach coupling natural and human systems for the entire eastern US barrier system requires defining a comprehensive set of coastal barrier spatial units to enable typological classification and subsequent systematic investigation to inform debates regarding coastal ecosystem services and sustainability.  相似文献   

20.
Bottom trawl estimates of densities of the portunid crab Liocarcinus depurator along the Mediterranean coast of Spain from 1994 to 2003, covering a depth range of 50–800 m, were analysed using non-linear geostatistical techniques. The variation of the spatial distribution was modelled through variogram analysis using depth as a covariate. Patch size ranged from 20 to 75 km. Crab density showed a general decrease from 1994 to 2003. L. depurator high-density areas were characterised as either occasional or permanent (always present every year). High-density patches appeared to be associated with either upwelling or strong river influence, i.e. the Western Alborán Sea and the Ebro Delta. The application of non-linear geostatistical techniques proved useful in identifying permanent high-density population patches, thereby highlighting regions of special interest for the ecology of the species.  相似文献   

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