Extremes in the resource consumption process: A mathematical model of the irruption of the moose population on Isle Royale. Part 1—Ecological background and model development

The development of a mathematical model of the moose-browse interaction on Isle Royale, Michigan, from the time of the establishment of the moose population in the early 1900s until the time of the arrival of wolves on the island about the mid-twentieth century is described. A brief history of the observations that were made during that period of the dynamics of the moose population and its resulting impact upon the vegetation is given. The simplifying assumptions that were made, the processes that were judged important enough to be represented in the model, the state variables that were selected and the input variables that were considered are delineated. A rationale is developed for the mathematical form of each of the terms used in the state equations. The literature that was utilized and the methodologies that were employed to obtain estimates of the parameters of the equations and the values of the initial conditions of the state variables are described.     

Turbulence investigation in the roughness sub-layer of a near wall flow

The turbulence behaviour along a wall roughened by pyramidal elements was analysed in the region extending from the apex of the roughness elements up to the external limit of the roughness sub-layer. The data used for the analysis were obtained by particle image velocimetry technique. The rough wall turbulent boundary layer flow is characterized by a relatively low Reynolds number. All the results on the rough wall were compared with data referring to the canonical flow on a smooth wall turbulent boundary layer. Mean values and turbulence quantities for the two flows collapse when approaching the external limit of the roughness sublayer. The quadrant analysis of the Reynolds shear stress, in the region near the surface, shows that the contribution of the sweep motions is about equivalent for the two flows (except for wall distances lower than 40 viscous units). The contribution of the ejection motions appears to be more important over the smooth wall than over the rough wall with increasing differences approaching the wall. The probability density functions of the streamwise fluctuating velocity field for the rough wall case appear to be positively skewed in the zone very close to the pyramid apex, in contrast with the behavior observed for the smooth wall case at corresponding distances from the wall. The integral and Taylor scales for the rough wall case appear to be strongly reduced by the presence of the roughness, while the Kolmogorov microscale shows higher values.     

海南岛儿童发钙量与土壤交换性钙关系的探讨

通过对海南岛上万名学龄前儿童头发钙含量以及所处地域农耕旱地土壤交换性钙的调查与分析发现：土壤钙与儿童头发钙含量呈极显著正相关;海南岛儿童头发钙均值低于国内其它地方的报道值;土壤环境钙含量的高低,能间接反映人体钙的营养状况。     

论我国水生态的保护与修复:任务与对策

本文聚焦我国水生态保护与修复工作,结合长期实践,分析当前的形势和任务,指出了当前要着力解决的水生态问题主要是:在大坝上游,河流变湖库,生境变化导致生物物种变化;水库水流流速变缓,水体自净能力降低,导致富营养化及藻类水华。在大坝下游,清水下泄,冲刷下游河道导致局部河道河势变化较大;水库蓄水使坝下游春季水温下降、秋季水温升高,水温变化过程滞后;水库蓄水导致大坝下游的水文过程改变,水的流量、流速、流态发生变化。在河道(航道)整治工程中,裁弯取直,缩短水流在河道中的停留时间,河岸衬砌和硬化,减少水向沿河堤岸的渗透,在大降水和洪水时易造成涝灾和洪灾。在涉湖工程中,阻隔河湖,填湖造地。在明确存在问题和迫切需求的基础上,提出了切实可行的对策建议。当前水生态保护与修复要围绕一条主线,即"人要发展,鱼要生存"的"人鱼线",要采取"调、控、退、通、改、拆"的综合措施,即生态调度,控制水污染,退建还水、退田还湖,河湖连通,对已建涉水工程进行生态化改造,对在保护区、重点风景名胜区、特有鱼类栖息地修建的小型工程要采取坚决措施拆除,恢复原貌。     

Characterizing the vegetation community in the provenance slope with high-frequency debris flow北大核心CSCD

In order to explore the characteristics and species diversity of the vegetation growing in the provenance slope with high-frequency debris flow, we selected the slopes on the north and south sides of the valley in the Jiangjiagou watershed as the research object. The structural characteristics and quantitative compositions of the communities of vegetation growing on different positions of the slopes were investigated. The species composition, important value, species diversity, and ground and underground biomass of the different communities on the slope were statistically analyzed, and the correlations of the ground and underground biomass with the diversity were determined. The results showed that: of the total 49 species found, the herbs were the dominant flora, of which 33 were found on the northern slope and 23 were found on the southern slope. There were significant differences in the species composition and composition of the important values along the different positions on the slope, being mainly affected by the pioneer herbaceous plants. We found that Heteropogon contortus, Eulaliopsis binata, Arthraxon hispidus, and Sesbania cannabina were highly adaptable to debris flow. These four species are common to the area and can be used as the main configuration species for future ecological restoration. There were differences in the community characteristics and species diversity at different positions on the slope. For the southern slope of the valley, the Shannon-Wiener diversity index was in the order: stable zone (2.311) > instable zone (2.161) > deposit zone (2.036), and in the order: deposit zone (2.626) > stable zone (1.338) > instable zone (1.057) for the northern slope. There were significant differences in the biomass, being in the order: stable zone > instable zone > deposit zone, with higher values in the northern slope than in the southern slope. Based on the aforementioned results, we suggest that the restoration of vegetation in the area of the provenance slope having high-frequency debris flow should focus on the prevention and protection of the instable zone of the slope by natural processes of recovery. The deposit zone needs rational remediation measures. We should introduce local shrub and arbor to increase the species composition and promote the development of plant communities with vertical structures. © 2018 Science Press. All rights reserved.     

武汉市“三废”排放的库兹涅茨特征及原因探析

通过分析武汉市1985-2001年近16年的“三废”排放量变化情况，发现武汉市“三废”的排放总量在波动中呈现上升趋势，但各类排放的时间规律相差明显。建立模型，将其人均GDP与“三废”排放的标准化值进行拟合，其计算结果显示：武汉市“三废”排放与人均GDP之间存在比较明显的三次曲线特征，在1985-1994年曲线呈持续下降趋势，而1995-2001年曲线初步显现出标准的环境库兹涅茨曲线。理论计算表明：武汉市“三废”排放的环境库兹涅茨曲线的转折点为人均GDP为12338．46元左右。通过环境经济学分析发现武汉市的经济发展、产业结构变动、能源结构变动和环境政策等4项因素是武汉市出现非标准环境库兹涅茨曲线的主要原因。     

黑土微生物量和酶活性对邻苯二甲酸二丁酯污染的响应

邻苯二甲酸二丁酯(di-n-butyl phthalate,DBP)是一种在环境中广泛存在的有毒有机化合物,已被我国列为优先控制污染物之一。本研究探讨了不同浓度的DBP污染对黑土呼吸、微生物量以及黑土酶活性的影响。结果表明,DBP污染处理的黑土呼吸速率和微生物量碳较对照均显著增加;微生物氮在DBP污染过程中呈"降低-升高-降低"波动性变化;微生物磷与DBP污染浓度呈显著负相关;DBP对黑土多酚氧化酶表现为先促进后抑制,对转化酶和蛋白酶活性表现为低浓度促进而高浓度抑制;在DBP污染过程中脲酶呈现被激活状态;黑土过氧化氢酶和酸性磷酸酶均受到DBP污染的显著抑制。通过相关性分析发现,土壤微生物量、土壤酶活性与DBP污染浓度之间存在着高度的相关性。由此可推断,DBP污染改变了黑土呼吸、微生物量和酶学活性的代谢特征,进而有可能影响了黑土的生态系统功能,威胁到黑土的可持续利用。     

三峡水库1956—2017年坝下径流特征分析

采用三峡水库坝下宜昌水文站1956—2017年日径流监测数据和长江上游80个国家气象站年均降水量长系列资料,利用一元线性分析法、滑动平均法、线性趋势回归法、非参数统计检验法、Mann-Kendall秩次相关、累积距平法和滑动T检验法分析三峡工程运行前后坝下径流的变化特征,其成果可为三峡工程调度提供参考。结果表明:三峡水库坝下宜昌水文站径流量变化趋势具有显著阶段性。三峡水库蓄水运行前宜昌水文站径流量和上游降水量相关性超过90%,该时期径流量主要受上游天然来水影响。三峡水库运行后,径流量主要受人类活动影响。宜昌水文站径流年际变化减少趋势不显著,长江上游降水量呈显著减少趋势。坝下径流量和上游降水量在2002年发生了突变且变化趋势相同,但径流量减少趋势不显著。径流年内分布发生了一定变化,其中汛期径流量占年径流总量百分比从72%下降到66%,枯期径流量占年径流量总量百分比从11%增加到15%。三峡工程调蓄是人类活动的直接表现,枯期宜昌水文站径流量增大是三峡工程生态效益的具体体现。     

Laboratory experiments on waveform inversion of an internal solitary wave over a slope-shelf

Internal solitary waves (ISWs) have been detected in many parts of the world oceans, particularly over slope-shelf topography, on which signature of waveform inversion has been identified. The effects of these waves on engineering operations and ecological process have also been reported in the literature. This article reports the results of a series of numerical modeling and laboratory experiments on waveform evolution of a depression ISW in a nearly stratified two-layer fluid system, in which specific water depth ratios above the horizontal plateau of the trapezoidal obstacle were arranged to facilitate the occurrence of waveform inversion. Classifications of waveform instability (no instability, shear instability and overturning with breaking) on the slope are confirmed in the present laboratory study. Numerical results for waveform variation are also found in fair agreement with the laboratory measurements for cases without waveform inversion and minor internal breaking. Moreover, laboratory results revealed that the depth ratio of the stratified two-layer fluid above the plateau and the magnitude of the incident ISW were the two most important factors for promoting waveform inversion beyond a turning point, in addition to the requirement of a sufficient distance from the shoulder of the trapezoidal obstacle. These factors also influenced the outcome of the shoaling process, energy dissipation, internal wave breaking and turbulent mixing on the front slope, as well as the likelihood of waveform inversion on the horizontal plateau. Contrary to the common perception, it was also observed, at least from the results of the present laboratory experiments, that not all the incident ISWs of depression would produce waveform inversion on the plateau, where the upper layer was physical greater than the bottom layer, unless moderate incident wave was provided. The outcome might also be attributed to the limited distance from the shoulder onto the plateau in the present laboratory setup. However, once waveform inversion occurred on the plateau, it was found, among others, that: (1) the amplitude of the transmitted leading crest and trough might be as low as 30 and 20%, respectively, to the amplitude of the incident wave in depression; (2) the characteristic wavelength of the transmitted leading trough doubled while that of the crest was asymptotically one-half of the incident wavelength, despite the wide range variation in the depth ratios above the plateau; and (3) the transmitted potential wave energy of the leading crest contained 30% of the incident energy. Based on the results of present laboratory experiments, the range for the non-dimensional parameter α, which indicates the effect of nonlinearity and the promotion of waveform inversion on horizontal plateau, will be proposed.     

Environmental concerns for marina planning in the Gulf of Suez

This paper presents a case study where numerical modelling was utilized for the first time to estimate shoreline changes during the planning of a private pleasure marina in the Gulf of Suez. The study was made to compliment an environmental impact assessment study (EIA) requested by the Egyptian Environmental Affairs Agency (EEAA). The paper presents data collected during two surveys and the results of the numerical model. The impact of the marina on the sediment budget was investigated using the GENESIS one-line program. One of the main reasons for the study was to confirm that the choice of the marina location ensured minimum erosion of the shoreline. In the model, the sediment transport calibration constants were determined using the results of two surveys. The choice of the formula is discussed in the paper. Two locations for the marina were tested against minimum erosion at the down drift side of the marina. This study was performed in close co-ordination with the EEAA and several solutions were suggested to minimize the expected accretion before the final location was approved.     

Etude expérimentale du comportement nuptial de Platynereis dumerilii (Annelida: Polychaeta): chémoréception, émission des produits génitaux

In the heteronereis form of Platynereis dumerilii, the roles of the swelling of the parapodial cirri of the male or female first segments, of the crenellations of the dorsal parapodial cirri of the male posterior region and those of the male pygidial papilli have been ascertained experimentally with respect to nuptial behaviour. The nuptial dance and the release of the genital products are provoked by chemical stimulating discharges from the partner of the opposite sex. The chemical stimuli act on the swelled or crenellated parapodial cirri, probably through the bipolar chemoreceptor neurons which are numerous on these organs. The excitation is transmitted to the cerebral ganglia by way of the ventral cord; it is translated into motor impulses which act upon the musculature concerned in the nuptial dance and in the release of genital products. Accordingly, the swelling and the crenellations of the parapodial cirri appear to contain chemoreceptors. The specificity of this chemoreception, in regard to the exciting substance emitted by the opposite sex, is probably related to the presence of a protein (Boilly-Marer, 1966a, b) elaborated during heteronereidation by epidermal cells adjacent to the dendritic extensions of the chemoreceptor cells. The pygidial papilli appear to insure sperm evacuation.     

土壤有机碳储量估算的影响因素和不确定性

土壤作为陆地生态系统最大的碳库和碳循环过程的关键环节,其源、汇的变化直接影响全球碳平衡,因此,土壤有机碳储量估算成为生态领域的重要研究内容之一。通过比较不同研究者在不同研究尺度上报道的有机碳储量的研究结果,发现这些研究结果较不一致。分析认为导致土壤有机碳储量评估结果存在较大差异的原因,主要是来自于采样过程中人类干扰以及气候变化等环境要素的波动,特别是研究者所采取的不同估算方法和背景资料。从环境要素（外因）和估算方法（内因）两个角度出发,提出了土壤有机碳储量的研究意义,阐述了造成土壤有机碳储量估算的不确定性和目前研究中存在的问题,并在此基础上,对其未来研究方向和重点进行了展望。     

天津市南运河河道整治景观设计

依据天津南运河河道沿岸的场地特征,确定了南运河河道整治景观设计的目标和原则,重点探析南运河线形文化景观遗存和滨河环境空间利用的实现形式,并认为南运河河道整治景观设计应建立沿岸连续的步行系统。依据道路与河面的高度差,建立6个不同的景观节点,采取生态护堤措施,保护和完善生态廊道,建立统一完善的历史文化传承体系。     

China's Grain for Green Program has reduced soil erosion in the upper reaches of the Yangtze River and the middle reaches of the Yellow River

Severe soil erosion in the upper reaches of the Yangtze River and the middle reaches of the Yellow River is a serious environmental problem in western China. In order to control this soil erosion, the national government implemented the Grain for Green Program (GGP) in Sichuan Province (in the upper reaches of the Yangtze River) and Shaanxi Province (in the middle reaches of the Yellow River) in 1999. To evaluate the effects of the GGP on soil erosion, the present study investigated the relationships between annual precipitation and three parameters, annual runoff, sediment concentration, and sediment discharge, for these rivers. As the area covered by the GGP increased, sediment concentration and sediment discharge both decreased, and the correlation with the GGP area was much stronger than the correlation with annual precipitation. Therefore, China's GGP appears to be controlling soil erosion in the upper reaches of the Yangtze River and the middle reaches of the Yellow River.     

Fourier analysis of the roll-up and merging of coherent structures in shallow mixing layers

The current study investigates the role of nonlinearity in the development of two-dimensional coherent structures (2DCS) in shallow mixing layers. A nonlinear numerical model based on the depth-averaged shallow water equations is used to investigate temporal shallow mixing layers, where the mapping from temporal to spatial results is made using the velocity at the center of the mixing layer. The flow is periodic in the stream-wise direction and the transmissive boundary conditions are used in the cross-stream boundaries to prevent reflections. The numerical results are examined with the aid of Fourier decomposition. Results show that the previous success in applying local linear theory to shallow mixing layers does not imply that the flow is truly linear. Linear stability theory is confirmed to be only valid within a short distance from the inflow boundary. Downstream of this linear region, nonlinearity becomes important for the roll-up and merging of 2DCS. While the energy required for the merging of 2DCS is still largely provided by the velocity shear, the merging mechanism is one where nonlinear mode interaction changes the velocity field of the subharmonic mode and the gradient of the along-stream velocity profile which, in turn, changes the magnitude of the energy production of the subharmonic mode by the velocity shear implicitly. The nonlinear mode interaction is associated with energy up-scaling and is consistent with the inverse energy cascade which is expected to occur in shallow shear flows. Current results also show that such implicit nonlinear interaction is sensitive to the phase angle difference between the most unstable mode and its subharmonic. The bed friction effect on the 2DCS is relatively small initially and grows in tandem with the size of the 2DCS. The bed friction also causes a decrease in the velocity gradient as the flow develops downstream. The transition from unstable to stable flow occurs when the bed friction balances the energy production. Beyond this point, the bed friction is more dominant and the 2DCS are progressively damped and eventually get annihilated. The energy production by the velocity shear plays an important role from the upstream end all the way to the point of transition to stable flow. The fact that linear stability theory is valid only for a short distance from the inflow boundary suggests that some elements of nonlinearity is incorporated in the mean velocity profile in experiments by the averaging process. The implicit nature of nonlinear interaction in shallow mixing layers and the sensitivity of the nonlinear interaction to phase angle difference between the most unstable mode and its subharmonic allows local linear theory to be successful in reproducing features of the instability such as the dominant mode of the 2DCS and its amplitude.     

Der kammerbauvorgang der großforaminifere Heterostegina depressa (Nummulitidae)

Chamber formation of Heterostegina depressa d'Orbigny 1826 mostly starts with the beginning of the daily light period. First, the necessary space for the construction of the new chamber (“Kammerhof”) is provided by rotation of the test inside the ectoplasm sheath, which is well fixed to the substrate. At the same time, protoplasm flows out of the aperture and out of the pores of the last chamber transporting organic particles. Three to 4 h after beginning of chamber formation, these particles coalesce to an organic layer between the ectoplasm sheath and the edge of the test. A short time later, this layer is bound on the outside by an organic membrane. Organic layer and organic membrane indicate the contour of the new chamber together with its chamberlets; they represent the “anlage” of the new chamber. Calcification takes place at this membrane. After the beginning of calcification, which lasts 15 to 20 h, the organic layer is removed and the ectoplasm sheath is retracted to the test.     

Cyclical contributions of the digestive epithelium to faecal pellet formation by the copepod Calanus helgolandicus

Faecal pellet formation within the gut of Stage V and adult females of the copepod Calanus helgolandicus Claus involves (1) cyclical processes of digestion and (2) the contribution of parts of the gut epithelium to the pellets. During an experimental regime in which dim lighting was restricted to day-time and feeding to night-time (17.00 to 09.00 hrs), the copepods responded with cyclical changes in both the quantity of pellets they produced and the fine structure of the contents. During the feeding period, the contents showed changes in the relative amounts of materials originating from disintegrated cells of the digestive epithelium and those derived directly from the ingested food. The vacuolar B-cells of the gut contribute to the content of the pellets and the distal, necrotic N-cells appear to be involved in forming the peritrophic membrane which encloses each pellet. Cells of the gut epithelium which are broken down during feeding are all replaced during the non-feeding period. Other individuals were taken directly from the sea and in these, also, the cells of the gut broke down during feeding and contributed to the faecal pellets. The supply of epithelial cells may limit the duration of the feeding period.     

Specialized feeding mechanism in the pelagic copepod genus Heterorhabdus (Calanoida: Heterorhabdidae), with special reference to the mandibular tooth and labral glands

Scanning and transmission electron microscopy and gut-content analysis revealed the fine structure of the mandibular ventralmost tooth and labral glands of the carnivorous copepods Heterorhabdus spp. on the basis of the specimens collected in the waters near Nansei Islands, Southwestern Japan, in November 1993. The ventralmost tooth is hollow with a subterminal pore and a basal opening. The distal region of the tooth is like a hypodermic needle and is strengthened by opal layers. The tip of the tooth including the subterminal pore usually protrudes from between the labrum and the paragnath, while the basal region of the ventralmost tooth and the whole parts of the other teeth are semi-enclosed in the preoral food chamber formed by the labrum and the paragnath. There is a large glandular cell in the basal region of the mandibular gnathobase, but no direct connection exists between the gnathobasal cells and the tubular lumen of the ventralmost tooth. The posterior surface of the labrum carries a lateral pair of large glandular pores, which are located close to, and appear to fit exactly the basal openings of the ventralmost teeth. Each of these glands has two massive secretory cells, and the secretory materials from these cells appear to be mixed before discharge from the pore. A specialized feeding mechanism is proposed, wherein venom or anaesthetic is injected from the labral-gland pores into the tubular lumen of the mandibular ventralmost tooth, and then relcased into the prey from its subterminal pore immediately after capture using the stout maxillae of prey macrozooplankters such as copepodids and polychaetes.     

The representation of the environment in the context of individual-based modeling

Individual-based modeling includes the explicit representation of the environment, to which individual organisms interact. From the perspective of spatial representation, spatially extended environment is different from discrete individual organisms, thus the success of object-orientation in the representation of individual organisms may not be applicable to the representation of the environment. Over the past few years, the attempt to adopt object-orientation in the representation of the environment has stimulated interesting discussions over what space is and how it may be represented in ecological modeling. The objective of this paper is to evaluate the merit of two traditional approaches used to represent the environment, i.e., the grid model and the patch model, and, in particular, the object-oriented versions of the two approaches in the representation of the environment. Two case studies are provided in support of the discussions of how the environment may be represented. One case study concerns individual fish growth and movement in an aquatic environment and the other concerns the movement of calving elk in a short-grass prairie. The discussion stresses the importance of two issues in the context of individual-based modeling: (1) the distinction between object-orientation used as a programming technique and as a representation means, and (2) the conceptual compatibility between a perceived phenomenon and the approach used to represent the phenomenon. It is suggested that caution be exercised in the practice of treating cells as objects. The paper concludes that two approaches may be appropriate for individual-based modeling. One is a hybrid approach that incorporates the traditional grid model of the environment and an object-oriented model of individual organisms. The other is the all-object approach that combines the object-oriented patches of the environment and the object-oriented individual organisms.