Does sheep selectivity along grazing paths negatively affect biological crusts and soil seed banks in arid shrublands? A case study in the Patagonian Monte,Argentina

Domestic animals potentially affect the reproductive output of plants by direct removal of aboveground plant parts but also could alter the structure and fertility of the upper soil and the integrity of biological crusts through trampling. We asked whether sheep selectivity of plant patches along grazing paths could lead to negative changes in biological crusts and soil seed banks. We randomly selected ten floristically homogeneous vegetation stands distributed across an area (1250 ha) grazed by free ranging sheep. Vegetation stands were differently selected by sheep as estimated through sheep-collaring techniques combined with remote imagery mapping. At each stand, we extracted 15 paired cylindrical soil cores from biological crusts and from neighboring soil without crusts. We evaluated the crust cover enclosed in each core and incubated the soil samples at field capacity at alternating 10-18 °C during 24 months. We counted the emerged seedlings and identified them by species. Sheep selectivity along grazing paths was largest at mid-distances to the watering point of the paddock. Increasing sheep selectivity was associated with the reduction of the cover of biological crusts and the size and species number of the soil seed bank of preferred perennial grasses under biological crusts. The size of the soil seed bank of annual grasses was reduced with increasing sheep selectivity under both crust and no crust soil conditions. We did not detect changes in the soil seed banks of less- and non- preferred species (shrubs and forbs) related to sheep selectivity. Our findings highlight the negative effects of sheep selectivity on biological crusts and the soil seed bank of preferred plant species and the positive relationship between biological crusts and the size of the soil seed bank of perennial grasses. Accordingly, the state of conservation of biological crusts could be useful to assess the state of the soil seed banks of perennial grasses for monitoring, conservation and planning the sustainable management of grazing lands.     

Seed bank composition of open and overgrown calcareous grassland soils--a case study from Southern Belgium

The success of calcareous grassland recreation following abandonment depends to a large extent on the composition of the soil seed bank. We studied the species richness and composition of the seed bank along a chronosequence from well-developed calcareous grassland to scrub and forest vegetation, which had developed on calcareous grassland over the last 225 years. The seed bank density was highest in the calcareous grassland soils (930 seeds/m(2)). However, the seed bank was mainly composed of rather common species of nutrient poor grassland, which were poorly represented in the surface vegetation. There were no significant differences in soil seed bank density and species composition between the calcareous grassland and scrub vegetation that was less than 15 years old, largely because several grassland species had persisted in the scrub vegetation and were therefore able to replenish the soil seed bank. In contrast seed density and species richness declined significantly after 40 years of grassland abandonment. Indeed, forest soils had the lowest seed densities (214 seeds/m(2)) with only a few grassland species represented. This reflects the lack of grassland species in the field layer of the forest, and, therefore, the lack of seed production and seed bank replenishment. It is clear that recreation of calcareous grassland on long abandoned sites cannot rely on germination of target species from the seed bank alone. Even in the calcareous grassland soil, seeds of target species are not abundant as the majority produces transient seeds, which decay rapidly if they do not germinate immediately. Successful grassland recreation on such sites therefore may require seeds of target species to be introduced artificially (e.g. as seed mixtures, green hay, etc.). Alternatively, reinstatement of traditional practices, such as grazing or mowing, will increase the natural dispersal potential of these species, allowing population reestablishment in the long term.     

Comparison of the Vegetation and Seed Bank on Hedge Banks of Different Ages in Brittany,France

The composition of the herbaceous cover and the seed bank of old and recent hedge banks in Brittany were studied and compared. Concentration method was used for seed bank samples. Grimes plant strategies were used to explain observed patterns. The analysis of the seed bank of these hedge banks showed that the species richness and diversity differed in relation to the date of construction of the hedge banks. The seed banks of recent hedge banks were richer and more diversified than those of old hedge banks. Differences in the floristic composition of the established plant cover between the recent and old hedge banks were determined by multivariate analyses. The species exclusively found in the seed bank and in the herb cover of recent hedge banks were mainly grassland species, whereas the species that only occurred on old hedge banks tended to be woodland species. The floristic composition of the two compartments (established vegetation and seed bank) was very different. A multivariate analysis revealed that the difference between the composition of the seed banks of recent and old hedge banks was less than that between the composition of the established vegetation and seed bank of hedge banks of the same age. Both seed bank and vegetation of recent hedge banks were dominated by ruderal species, whereas old hedge vegetation was dominated by stress-tolerant woodland species, indicating that mechanically constructed hedge banks may impose limitations on colonization by late woodland species.     

Measuring Plant Diversity in the Tall Threetip Sagebrush Steppe: Influence of Previous Grazing Management Practices

In July 2000, a 490-ha wildfire burned a portion of a long-term grazing study that had been established in 1924 at the US Sheep Experiment Station north of Dubois, Idaho, USA. Earlier vegetation measurements in this tall threetip sagebrush (Artemisia tripartita spp. tripartita) bunchgrass plant community documented significant changes in vegetation due to grazing and the timing of grazing by sheep. A study was initiated in May 2001 using 12 multiscale modified Whittaker plots to determine the consequences of previous grazing practices on postfire vegetation composition. Because there was only one wildfire and it did not burn all of the original plots, the treatments are not replicated in time or space. We reduce the potential effects of psuedoreplication by confining our discussion to the sample area only. There were a total of 84 species in the sampled areas with 69 in the spring-grazed area and 70 each in the fall- and ungrazed areas. Vegetation within plots was equally rich and even with similar numbers of abundant species. The spring-grazed plots, however, had half as much plant cover as the fall- and ungrazed plots and the spring-grazed plots had the largest proportion of plant cover composed of introduced (27%) and annual (34%) plants. The fall-grazed plots had the highest proportion of native perennial grasses (43%) and the lowest proportion of native annual forbs (1%). The ungrazed plots had the lowest proportion of introduced plants (4%) and the highest proportion of native perennial forbs (66%). The vegetation of spring-grazed plots is in a degraded condition for the environment and further degradation may continue, with or without continued grazing or some other disturbance. If ecosystem condition was based solely on plant diversity and only a count of species numbers was used to determine plant diversity, this research would have falsely concluded that grazing and timing of grazing did not impact the condition of the ecosystem.     

The Impact of Different Grazing Periods in Dry Grasslands on the Expansive Grass <Emphasis Type="Italic">Arrhenatherum elatius</Emphasis> L. and on Woody Species

Dry grasslands are one of the most species rich and endangered types of vegetation in Europe. In the Czech Republic, dry grasslands are mainly of anthropogenic origin and were formed as a result of grazing after the clear-cutting of thermophilous oak woods. Gradual changes in the farming landscape throughout the 20th century, particularly in the 1960s, resulted in the abandonment of the relatively infertile habitats of dry grasslands. After abandonment, dry grasslands decline and degrade due to the gradual overgrowth of woody species and expansion of perennial tall grasses. In the year 2000, a grazing management program was introduced in the protected areas within the territory of Prague City to maintain the species diversity of dry grasslands. The responses of the expansive grass species, Arrhenatherum elatius L. and multiple woody species (especially, Prunus spinosa L.) to differences in grazing periods were monitored for over a decade. Grazing in spring through the end of June had the greatest impact on the reduction of A. elatius and woody species. Grazing in the height of summer through autumn did not reduce the cover of these plants, and may support the prosperity of both A. elatius and the woody species due to higher levels of nutrients.     

Vegetation structure, niche width, niche overlap and types of competition in temperate grazingland of Garhwal Himalaya, India

Temperate grazing lands in Garhwal Himalaya are rich in herbaceous vegetation and extensively used for grazing by locally owned livestock. Capillipedium parviflorum is a frequently occurring and dominant grass species under Pine canopy in associate with several herbs. Beside, grazing and extensive fire, interactions among associated species is a determining factor of community structure. In this article the dominant diversity pattern of vegetation in temperate grazing land along two different altitudes and having different slope aspects is described. Interaction between dominant and co-dominant species with their niche appearance and niche overlap measurement was also observed and presented here. South and west facing slopes had maximum species and diversity values while east north facing slopes had higher species dominant. Capillipedium parviflorum and other grasses had maximum niche values and were dominant and co dominant species. Three types of interspecific interaction were observed between species and grasses were observed as succeeding species in all sites of different topographic conditions. Dominant diversity pattern along with resource partitioning and interspecific competition is discussed and presented here.     

Restoration of Mountain Big Sagebrush Steppe Following Prescribed Burning to Control Western Juniper

Western juniper (Juniperus occidentalis ssp. occidentalis Hook) encroachment into mountain big sagebrush (Artemisia tridentata spp. vaseyana (Rydb.) Beetle) steppe has reduced livestock forage production, increased erosion risk, and degraded sagebrush-associated wildlife habitat. Western juniper has been successfully controlled with partial cutting followed by prescribed burning the next fall, but the herbaceous understory and sagebrush may be slow to recover. We evaluated the effectiveness of seeding perennial herbaceous vegetation and sagebrush at five sites where juniper was controlled by partially cutting and prescribed burning. Treatments tested at each site included an unseeded control, herbaceous seed mix (aerially seeded), and the herbaceous seed mix plus sagebrush seed. In the third year post-treatment, perennial grass cover and density were twice as high in plots receiving the herbaceous seed mix compared to the control plots. Sagebrush cover and density in the sagebrush seeded plots were between 74- and 290-fold and 62- and 155-fold greater than the other treatments. By the third year after treatment, sagebrush cover was as high as 12 % in the sagebrush seeded plots and between 0 % and 0.4 % where it was not seeded. These results indicate that aerial seeding perennial herbaceous vegetation can accelerate the recovery of perennial grasses which likely stabilize the site. Our results also suggest that seeding mountain big sagebrush after prescribed burning encroaching juniper can rapidly recover sagebrush cover and density. In areas where sagebrush habitat is limited, seeding sagebrush after juniper control may increase sagebrush habitat and decrease the risks to sagebrush-associated species.     

Are There Benefits to Mowing Wyoming Big Sagebrush Plant Communities? An Evaluation in Southeastern Oregon

Wyoming big sagebrush (Artemisia tridentata ssp. wyomingensis Beetle & Young) communities frequently are mowed in an attempt to increase perennial herbaceous vegetation. However, there is limited information as to whether expected benefits of mowing are realized when applied to Wyoming big sagebrush communities with intact understory vegetation. We compared vegetation and soil nutrient concentrations in mowed and undisturbed reference plots in Wyoming big sagebrush plant communities at eight sites for three years post-treatment. Mowing generally did not increase perennial herbaceous vegetation cover, density, or biomass production (P > 0.05). Annual forbs and exotic annual grasses were generally greater in the mowed compared to the reference treatment (P < 0.05). By the third year post-treatment annual forb and annual grass biomass production was more than nine and sevenfold higher in the mowed than reference treatment, respectively. Our results imply that the application of mowing treatments in Wyoming big sagebrush plant communities does not increase perennial herbaceous vegetation, but may increase the risk that exotic annual grasses will dominate the herbaceous vegetation. We suggest that mowing Wyoming big sagebrush communities with intact understories does not produce the expected benefits. However, the applicability of our results to Wyoming big sagebrush communities with greater sagebrush cover and/or degraded understories needs to be evaluated.     

Benefits of protective fencing to plant and rodent communities of the western Mojave Desert,California

Human disturbance in the western Mojave Desert takes many forms. The most pervasive are livestock grazing and off-highway vehicle use. Over the past few decades several areas within this region have been fenced to preclude human disturbance. These areas provide opportunities to study the impact of human activities in a desert ecosystem. This paper documents the response of plant and small mammal populations to fencing constructed between 1978 and 1979 at the Desert Tortoise Research Natural Area, Kern County, California. Aboveground live annual plant biomass was generally greater inside than outside the fenced plots during April 1990, 1991, and 1992. The alien grassSchismus barbatus was a notable exception, producing more biomass in the unprotected area. Forb biomass was greater than that of alien annual grasses inside the fence during all three years of the study. Outside the fence, forb biomass was significantly higher than that of alien grasses only during spring 1992. Percent cover of perennial shrubs was higher inside the fence than outside, while no significant trend was detected in density. There was als more seed biomass inside the fence; this may have contributed to the greater diversity and density of Merriam's kangaroo rats (Dipodomys merriami), long-tailed pocket mice (Chaetodipus formosus), and southern grasshopper mice (Onychomys torridus) in the protected area. These results show that protection from human disturbance has many benefits, including greater overall community biomass and diversity. The significance and generality of these results can be further tested by studying other exclosures of varying age and configurations in different desert regions of the southwestern United States.     

Rapid vegetation regeneration in a seriously degraded Rhanterium epapposum community in northern Kuwait after 4 years of protection

A study was carried out in northern Kuwait to investigate vegetation recovery in sandy depressions in the Sabriya oilfield ('Sabriya-IN'), 4 years after it was completely protected from livestock grazing and other anthropogenic activities which have largely depleted the dwarf shrub vegetation. This vegetation was compared with that in seriously overgrazed depressions outside the oilfield ('Sabriya-OUT'), where negative influences persist, and with a sandy site in central Kuwait which has been protected for over 20 years ('Sulaibiya'). There has been a striking recovery of the dwarf shrub vegetation in Sabriya-IN during the 4 years, with cover values of shrubs as high as at Sulaibiya. Rhanterium epapposum has been the main species to benefit from protection. However, the shrubs have not regenerated from seed, but rather from underground stumps that have probably remained in the soil for decades. Cover values of the annual flora at Sabriya-IN were very similar to those at Sulaibiya, and they were significantly higher than at Sabriya-OUT. However, it appears that some species may have disappeared or become extremely rare at Sabriya-IN when compared with Sulaibiya, as Sabriya-IN and Sabriya-OUT are remarkably similar floristically. Despite the impressive regeneration of the dwarf shrub vegetation at Sabriya-IN, which contradicts the view that vegetation recovery is a slow process in desert ecosystems, it is important to consider what the natural vegetation was in this part of the world. It is suggested that the region was once dominated by an open Acacia woodland, in which perennial grasses comprised most of the ground layer, and that the current dwarf shrub vegetation is a response to decades, if not centuries, of moderate to heavy grazing.     

Germinable Soil Seed Banks and the Restoration Potential of Abandoned Cropland on the Chinese Hilly-Gullied Loess Plateau

Poor vegetation cover is generally considered to be a major factor causing soil erosion on the Loess Plateau in China. It has been argued that tree planting restoration is ineffective, and natural re-vegetation is an alternative ecological solution for restoring abandoned cropland and controlling soil erosion. The aims of this study were to investigate the characteristics of soil seed banks and to assess the natural restoration potential of abandoned cropland in the hilly-gullied Loess Plateau. The soil seed bank was identified by the germination method with the soil samples, which were collected at four sampling times (April, August, and October 2005 and August 2006) from 12 plots abandoned 3–30 years prior to sampling. The seed bank densities of all of the samples in the 0–10 cm soil layer varied from 1,067 ± 225 to 14,967 ± 1,606 seeds m⁻². Fifty-one species (24 annual and 27 perennial species) belonging to 18 families were identified, and 39% of these species belonged to the families Compositae and Gramineae. The pioneer species Artemisia scoparia dominated the seed bank, with an average seed density of 3,722 seeds m⁻², and accounted for 74.4% of the seeds in the bank. The local dominant species (such as Lespedeza davurica, Artemisia gmelinii, Bothriochloa ischaemun and Stipa bungeana) of the later succession stages also existed at densities varying from 17 to 1, 383 seeds m⁻². The combination of soil seed bank characteristics, reproductive traits of the species, the specific landscape conditions indicates that the potential to restoring the abandoned croplands in the hilly-gullied Loess Plateau via natural re-vegetation could be substantial.     

Grazed Riparian Management and Stream Channel Response in Southeastern Minnesota (USA) Streams

The U.S. Department of Agriculture-Natural Resources Conservation Service has recommended domestic cattle grazing exclusion from riparian corridors for decades. This recommendation was based on a belief that domestic cattle grazing would typically destroy stream bank vegetation and in-channel habitat. Continuous grazing (CG) has caused adverse environmental damage, but along cohesive-sediment stream banks of disturbed catchments in southeastern Minnesota, short-duration grazing (SDG), a rotational grazing system, may offer a better riparian management practice than CG. Over 30 physical and biological metrics were gathered at 26 sites to evaluate differences between SDG, CG, and nongrazed sites (NG). Ordinations produced with nonmetric multidimensional scaling (NMS) indicated a gradient with a benthic macroinvertebrate index of biotic integrity (IBI) and riparian site management; low IBI scores associated with CG sites and higher IBI scores associated with NG sites. Nongrazed sites were associated with reduced soil compaction and higher bank stability, as measured by the Pfankuch stability index; whereas CG sites were associated with increased soil compaction and lower bank stability, SDG sites were intermediate. Bedrock geology influenced NMS results: sites with carbonate derived cobble were associated with more stable channels and higher IBI scores. Though current riparian grazing practices in southeastern Minnesota present pollution problems, short duration grazing could reduce sediment pollution if managed in an environmentally sustainable fashion that considers stream channel response.     

Limits to Understory Plant Restoration Following Fuel-Reduction Treatments in a Piñon–Juniper Woodland

National fuel-reduction programs aim to reduce the risk of wildland fires to human communities and to restore forest and rangeland ecosystems to resemble their historical structure, function, and diversity. There are a number of factors, such as seed bank dynamics, post-treatment climate, and herbivory, which determine whether this latter goal may be achieved. Here, we examine the short-term (2 years) vegetation response to fuel-reduction treatments (mechanical mastication, broadcast burn, and pile burn) and seeding of native grasses on understory vegetation in an upland piñon–juniper woodland in southeast Utah. We also examine how wildlife herbivory affects the success of fuel-reduction treatments. Herbaceous cover increased in response to fuel-reduction treatments in all seeded treatments, with the broadcast burn and mastication having greater increases (234 and 160 %, respectively) in herbaceous cover than the pile burn (32 %). In the absence of seeding, herbaceous cover only increased in the broadcast burn (32 %). Notably, fuel-reduction treatments, but not seeding, strongly affected herbaceous plant composition. All fuel-reduction treatments increased the relative density of invasive species, especially in the broadcast burn, which shifted the plant community composition from one dominated by perennial graminoids to one dominated by annual forbs. Herbivory by wildlife reduced understory plant cover by over 40 % and altered plant community composition. If the primary management goal is to enhance understory cover while promoting native species abundance, our study suggests that mastication may be the most effective treatment strategy in these upland piñon–juniper woodlands. Seed applications and wildlife exclosures further enhanced herbaceous cover following fuel-reduction treatments.     

Sheep grazing effects on Mojave Desert vegetation and soils

Some effects of domestic sheep grazing on vegetation and soils were measured at four sites in the western Mojave Desert. Although sheep have grazed the Mojave Desert for the past 50–100 years, the effects of grazing on the desert ecosystem are largely unknown.The results reflect only short-term effects of grazing, because fenced control areas were not available for study. Heavy grazing caused a 60% reduction in the above-ground biomass under creosote bushes (Larrea tridentata), and sheep trampling caused an increase in soil strength and decreased intershrub annual densities 24 and 28% in two areas. The average area per individual (cover) of burrobush (Ambrosia dumosa) decreased 16–19% and the cover of spiny hop-sage (Grayia spinosa) decreased 29% under grazing pressure. The volumes of individual Ambrosia were 21 and 65% less in two heavily grazed areas than in two lightly grazed areas; the volume of goldenhead (Acamptopappus sphaerocephalus) was 68% less in one heavily grazed area.These changes indicate that the range quality of the Mojave Desert is deteriorating under sheep grazing pressures. Trampling reduces annual cover and disrupts the soil surface, thus promoting wind erosion. The loss of annual biomass and shrub cover should adversely affect reptiles and rodents by removing food sources and protection. Soil strength increases may retard future growth of annuals, further contributing to erosion and food source losses. Studies should be initiated to determine the long-term effects of grazing in the Mojave Desert.     

Combining Aboriginal and Non-Aboriginal Knowledge to Assess and Manage Feral Water Buffalo Impacts on Perennial Freshwater Springs of the Aboriginal-Owned Arnhem Plateau, Australia

Aboriginal land managers have observed that feral Asian water buffalo (Bubalis bubalis Lydekker) are threatening the ecological and cultural integrity of perennial freshwater sources in Arnhem Land, Australia. Here we present collaborative research between the Aboriginal Rangers from Warddeken Land Management Limited and Western scientists which quantified the ground-level impacts of buffalo on seven perennial freshwater springs of the Arnhem Plateau. A secondary aim was to build the capacity of Aboriginal Rangers to self-monitor and evaluate the ecological outcomes of their land management activities. Sites with high buffalo abundance had significantly different ground, ground cover, and water quality attributes compared to sites with low buffalo abundance. The low buffalo abundance sites were characterized by tall herbaceous vegetation and flat ground, whereas wallows, bare ground, and short ungrazed grasses were indicators of sites with high buffalo abundance. Water turbidity was greater when buffalo abundance was high. The newly acquired monitoring skills and derived indicators of buffalo damage will be used by Aboriginal Rangers to assess the ecological outcomes of their future buffalo control efforts on the Arnhem Plateau.     

Use of Remote Sensing Techniques to Determine the Effects of Grazing on Vegetation Cover and Dune Elevation at Assateague Island National Seashore: Impact of Horses

The effects of grazing by feral horses on vegetation and dune topography at Assateague Island National Seashore were investigated using color-infrared imagery, lidar surveys, and field measurements. Five pairs of fenced and unfenced plots (300 m²) established in 1993 on sand flats and small dunes with similar elevation, topography, and vegetation cover were used for this study. Color-infrared imagery from 1998 and field measurements from 2001 indicated that there was a significant difference in vegetation cover between the fenced and unfenced plot-pairs over the study period. Fenced plots contained a higher percentage of vegetation cover that was dominated by American beachgrass (Ammophila breviligulata). Lidar surveys from 1997, 1999, and 2000 showed that there were significant differences in elevation and topography between fenced and unfenced plot-pairs. Fenced plots were, on average, 0.63 m higher than unfenced plots, whereas unfenced plots had generally decreased in elevation after establishment in 1993. Results demonstrate that feral horse grazing has had a significant impact on dune formation and has contributed to the erosion of dunes at Assateague Island. The findings suggest that unless the size of the feral horse population is reduced, grazing will continue to foster unnaturally high rates of dune erosion into the future. In order to maintain the natural processes that historically occurred on barrier islands, much larger fenced exclosures would be required to prevent horse grazing.     

Effects of Protective Fencing on Birds, Lizards, and Black-Tailed Hares in the Western Mojave Desert

Lepus californicus ), perennial plant cover, and structural diversity of perennial plants were evaluated from spring 1994 through winter 1995 at the Desert Tortoise Research Natural Area (DTNA), in the Mojave Desert, California. Abundance and species richness of birds were higher inside than outside the DTNA, and effects were larger during breeding than wintering seasons and during a high than a low rainfall year. Ash-throated flycatchers (Myiarchus cinerascens), cactus wrens (Campylorhynchus brunneicapillus), LeConte's thrashers (Toxostoma lecontei), loggerhead shrikes (Lanius ludovicianus), sage sparrows (Amphispiza belli), and verdins (Auriparus flaviceps) were more abundant inside than outside the DTNA. Nesting activity was also more frequent inside. Total abundance and species richness of lizards and individual abundances of western whiptail lizards (Cnemidophorous tigris) and desert spiny lizards (Sceloporus magister) were higher inside than outside. In contrast, abundance of black-tailed hares was lower inside. Structural diversity of the perennial plant community did not differ due to protection, but cover was 50% higher in protected areas. Black-tailed hares generally prefer areas of low perennial plant cover, which may explain why they were more abundant outside than inside the DTNA. Habitat structure may not affect bird and lizard communities as much as availability of food at this desert site, and the greater abundance and species richness of vertebrates inside than outside the DTNA may correlate with abundances of seeds and invertebrate prey.     

Native Grasses as a Management Alternative on Vegetated Closure Caps

Capped waste sites often are vegetated with commercial turf grasses to increase evapotranspiration and prevent erosion and possible exposure of the barrier. Fertilizer, frequent watering, and mowing may be required to establish the turf grass and prevent invasion by trees and shrubs. Oldfield vegetation of grasses and forbs is a possible sustainable alternative to turf grass communities. To determine if oldfield vegetation can establish on caps, we (1) compared establishment of a dominant oldfield grass and a commercial turf grass under different combinations of new closure cap management: spring or summer planting and presence or absence of amendments to alleviate drought (watering, mulch) or increase soil fertility (fertilizer, lime, a nitrogen-fixing legume); (2) surveyed existing caps to determine if oldfield species establish naturally; and (3) performed a greenhouse experiment to compare growth of two native grasses under low and amended (added water, soil nutrients) conditions. Both the commercial grass and oldfield species established under new cap conditions; fertilizer, water, and mulch improved vegetation establishment in spring or summer, but legumes decreased grass cover. In the greenhouse, both native grasses grew best with amendments; however, substantial stem and root length were obtained with no fertilizer and only once-weekly watering. Existing vegetated caps supported planted grasses and naturally established oldfield species. Overall, the results indicate native grasses can establish on new caps and oldfields can serve as a management model; further work is needed to determine the management strategy to maintain herbaceous vegetation and slow woody species invasion.     

Initial effects of light armored vehicle use on grassland vegetation at Fort Lewis, Washington

Sustainable use of military training lands requires understanding and prediction of the effects of military vehicles on vegetation. We examined the initial impacts of an 8-wheeled, light armored vehicle (LAV) on grassland vegetation at Fort Lewis, Washington. The LAV drove replicate spiral paths at two starting velocities, 10.3 and 5.1 m s(-1). The disturbed width (width of ground impacted by the tires) increased as turning radius decreased, but was unaffected by vehicle velocity. An inverse-exponential model predicted disturbed width (r(2)=0.68) at all turning radii for both velocities combined. In low-velocity spirals, and for straight tracking (turning radius>40 m) and moderate turns (radius 20-40 m) in high-velocity spirals, all vegetation damage was imprint (plants flattened by wheels). During sharp (radius <20 m), high-velocity turns, most or all of the disturbed width was scraped free of surface vegetation and soil, which was piled to the outside of each tire track. Total plant cover (all species) was not affected by track curvature in low-velocity spirals, but decreased in the order straight tracking>moderate turns>sharp turns in high-velocity spirals. In low-velocity spirals, post-tracking cover of several plant growth forms (non-native species, perennial species, sod-forming grasses) was similar to pre-tracking cover, but in high-velocity spirals, post-tracking cover of these growth forms decreased in the order straight > or =moderate=sharp. Cover of native species and forbs decreased more in high- than in low-velocity spirals, but was unaffected by curvature. Pre- and post-tracking cover of annual species, bunchgrasses, and shrubs was < or =3%. The most severe vegetation damage caused by operation of wheeled LAVs on grasslands is associated with sharp, high-velocity turns.     

Short-Term Influence of Tank Tracks on Vegetation and Microphytic Crusts in Shrubsteppe Habitat

Artemisia tridentata Nutt.) habitat within the Idaho Army National Guard Orchard Training Area in southwestern Idaho. The purpose of this study was to determine the short-term (1–2 years) influence of tank tracks on vegetation and microphytic crusts in shrubsteppe habitat. The two types of tank tracks studied were divots (area where one track has been stopped or slowed to make a sharp turn) and straight-line tracks. Divots generally had a stronger influence on vegetation and microphytic crusts than did straight-line tracks. Tank tracks increased cover of bare ground, litter, and exotic annuals, and reduced cover of vegetation, perennial native grasses, sagebrush, and microphytic crusts. Increased bare ground and reduced cover of vegetation and microphytic crusts caused by tank tracks increase the potential for soil erosion and may reduce ecosystem productivity. Reduced sagebrush cover caused by tank tracks may reduce habitat quality for rodents. Tank tracks may also facilitate the invasion of exotic annuals into sagebrush habitat, increasing the potential for wildfire and subsequent habitat degradation. Thus, creation of divots and movement through sagebrush habitat by tanks should be minimized.