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1.
Bottom surges generated from dredged material discharges in the open ocean have been observed using high frequency acoustic concentration profilers in several field studies during the past five years. the locations, water depths, bottom slopes, oceanographic conditions, and dredged material composition differed from study to study. Observed surges at three dredged material disposal sites may develop more than one surge peak for a single discharge. for water depths of the order of 10 m, surge height of the leading peak was estimated to be about one quarter of the water depth. for water of greater depth, of the order of 100 m, surge height reached 70 m, about 70% of the water depth. Surge height is established instantaneously when dredged material hits the bottom, and remains relatively constant as the surge advances horizontally. Total surge length reached 150 m for water depths of 10 m when measured from the impact point to the leading edge. for water depths of more than 100 m, the surge length reached more than 100 m. Length of the leading surge peak was as large as 45 m at this water depth.

Dimensional analysis was applied to relate the surge height of the leading surge peak to discharge parameters and oceanographic conditions. Results showed that the ratio of surge height to water depth was proportional to 1/10 power of the ratio of discharge volume to the third power of water depth.  相似文献   

2.
A field data collection project was undertaken to investigate the short-term fate of dredged material discharged in the designated Miami Ocean Dredged Material Disposal Site (ODMDS) before dredging of the Miami River and the Miami Harbor Turning Basin begins. the designated ODMDS is located in relatively deep water for discharge sites with a typical bottom depth of 150 metres and is also located in the western boundary region of the Gulf Stream current off Miami. Acoustical backscattering, current, particulate, temperature and salinity data were gathered over a three day period from April 24, 1990 through April 26, 1990. the major generic features of shallow-water discharge plumes were observed to be present: (a) the presence of a rapid convective descending plume portion; (b) impact of that plume portion with the ocean bottom and concomitant generation of a bottom surge; (c) rapid horizontal width growth of the descending plume through entrainment; and (d) retention of a residual plume portion within the water column. A well-mixed upper water column layer extending to a depth of 40 to 60 metres below the surface of the ocean permitted measurements of the plume entrainment coefficient free from bottom boundary, water column density gradient, and vertical current shear effects which are usually present in relatively shallow, e.g. less than 40 metres bottom depth, coastal ocean discharge studies. Entrainment coefficient estimates obtained in this study were between 0.5 to 0.7. the residual water plume material was tracked over one-half hour during each of eight discharge events and was transported in a north-northeast direction.  相似文献   

3.
A field data collection project was undertaken to investigate the short-term fate of dredged material discharged in the designated Miami Ocean Dredged Material Disposal Site (ODMDS) before dredging of the Miami River and the Miami Harbor Turning Basin begins. the designated ODMDS is located in relatively deep water for discharge sites with a typical bottom depth of 150 metres and is also located in the western boundary region of the Gulf Stream current off Miami. Acoustical backscattering, current, particulate, temperature and salinity data were gathered over a three day period from April 24, 1990 through April 26, 1990. the major generic features of shallow-water discharge plumes were observed to be present: (a) the presence of a rapid convective descending plume portion; (b) impact of that plume portion with the ocean bottom and concomitant generation of a bottom surge; (c) rapid horizontal width growth of the descending plume through entrainment; and (d) retention of a residual plume portion within the water column. A well-mixed upper water column layer extending to a depth of 40 to 60 metres below the surface of the ocean permitted measurements of the plume entrainment coefficient free from bottom boundary, water column density gradient, and vertical current shear effects which are usually present in relatively shallow, e.g. less than 40 metres bottom depth, coastal ocean discharge studies. Entrainment coefficient estimates obtained in this study were between 0.5 to 0.7. the residual water plume material was tracked over one-half hour during each of eight discharge events and was transported in a north-northeast direction.  相似文献   

4.
A positive surge results from a sudden change in flow that increases the depth. It is the unsteady flow analogy of the stationary hydraulic jump and a geophysical application is the tidal bore. Positive surges are commonly studied using the method of characteristics and the Saint-Venant equations. The article presents the results from new experimental investigations conducted in a large rectangular channel. Detailed unsteady velocity measurements were performed with a high temporal resolution using acoustic Doppler velocimetry and non-intrusive free-surface measurement devices. Several experiments were conducted with the same initial discharge (Q = 0.060 m3/s) and six different gate openings after closure resulting in both non-breaking undular and breaking bores. The analysis of undular surges revealed wave amplitude attenuation with increasing distance of surge propagation were in agreement with Ippen and Kulin theory. Also, undular wave period and wave length data were relatively close to the values predicted by the wave dispersion theory for gravity waves in intermediate water depths.  相似文献   

5.
淀山湖沉水植物——苦草群落恢复影响因子研究   总被引:1,自引:0,他引:1  
在淀山湖0.5、1.0、1.5、2.0 m水层进行苦草(Vallisneria natans)现场生长实验,试图找出淀山湖苦草植被恢复的适宜环境条件。试验期间每天测量光照强度和水体溶解氧等环境因子,每隔10 d监测苦草的根长、株高、鲜重等生长指标的变化情况。结果表明:水深对苦草的生长具较显著影响,光照强度、溶氧分别和苦草日相对生长率具显著相关性;苦草的株高、叶片数、根长等生物学生长指标在0.5、1.0 m水层生长良好,1.5 m与2.0 m均出现不同程度的负增长状况。光照强度与苦草日相对生长率之间具有显著相关性(r=0.905),0.5、1.0 m水层的光照强度下苦草的相对生长率分别为0.14和0.11,而1.5、2.0 m水层的光照强度下苦草出现负增长的情况。水中的溶解氧含量与苦草成活率之间具有显著相关性(r=0.935)。随着水深的增加,溶氧逐渐降低,苦草的成活率也逐渐下降。在2.0 m水层处,溶氧均值为2.76 mg.L-1,苦草的成活率为46.5%,在1.0 m水层处,溶氧为5.66 mg.L-1,苦草成活率为86.5%,因此,淀山湖苦草群落恢复宜先在1.0 m以浅的沿岸带开展。  相似文献   

6.
Development of contemporary Eastern Pacific coral reefs   总被引:4,自引:0,他引:4  
Dana  T. F. 《Marine Biology》1975,33(4):355-374
An overview of oceanographic conditions prevailing in the tropical Eastern Pacific Ocean suggests that the entire region is environmentally marginal for coral-reef development. The principal features of this environment are a strong, permanent, shallow thermocline and an annual north-south migration of the Intertropical Convergence resulting in wet and dry seasons. Along tropical Eastern Pacific continental margins structural coral reefs are best developed in the Gulf of Chiriquí off western Panamá. These reefs are relatively small, with reef formation taking place at a maximum depth of roughly 10 m. All of the reefs are judged to have formed since sea level approached its present height some 5,000 years ago. A study of the physical environment in the Gulf of Chiriquí revealed the following. Seasonal differences in surface temperatures were small but significant (P0.01), with the dry-season median (Md) of 28.9°C higher than the rainy season one of 28.0°C. At all times, surface-water temperatures were within the range considered optimal for coral growth. There were also significant (P<0.05) seasonal differences in the depths of the 25°, 20°, and 18°C isotherms. The first was shallowest (Md=18.5 m) during the rainy season due to vertical mixing, while the latter two were shallowest (Mds=31.5 and 33.0 m, respectively) during the dry season due to a generalized shoaling of the thermocline. All three isotherms are closely associated with the thermocline and showed remarkable variability in depth, most likely connected with internal waves. Salinities were reduced down to depths greater than 20 m and for distances of more than 50 km from the coast. Seasonal differences were slight (0.7% S) but statistically significant (P<0.01). Turbidity during the rainy season reduces the amount of light reaching the bottom at 10 m depth roughly by a factor of three compared to the dry season. Even the dry season amount is only about one half as much as would be expected to reach the same depth on the seaward reef of a Westen Pacific atoll. These conditions of cool water a short distance below the surface, reduced salinities, and high seasonal turbidity combine to make the region a poor one for coral-reef formation. The history of the Eastern Pacific coral fauna is traced from the Cretaceous to the Holocene. The present fauna is of Indo-Western Pacific origin, having become established following (1) the final closure of the connections between the Caribbean and Eastern Pacific (Pliocene), (2) movement of the northern Line Islands by sea-floor spreading into the path of the North Pacific Equatorial Countercurrent (Pliocene), and (3) the loss of all Eastern Pacific hermatypes during the Pleistocene.  相似文献   

7.
Contrasting conditions at-sea are likely to affect the foraging behaviour of seabirds. However, the effect of season on the dive parameters of penguins is poorly known. We report here on an extensive study of the diving behaviour of king penguins (Aptenodytes patagonicus) over the bird's complete annual cycle at the Crozet Islands. Time-depth recorders were used to record dive duration, bottom duration, post-dive interval, ascent rate and descent rate in breeding adults during different seasons in 1995 and 1996. Seasons included summer (n=6, incubation; n=6, chick brooding), autumn and winter (n=5 and n=3, respectively, chick at the crèche stage), and spring (n=4, birds at the post-moult stage). In all seasons dive duration increased with dive depth, but, for a given depth, dives were longer in winter (6.8 min when averaged over the 100-210 m depth layer) than in spring (4.6 min) and summer (4.4 min). The time spent at the bottom of the dives, which probably represents a substantial part of the feeding time, was much longer in winter (2.5 min per dive for dives over the 100-210 m layer) than during other seasons (1.0-1.4 min), i.e. there was a 2.5-fold augmentation for similar diving depths. Ascent and descent rates increased with increasing dive depth, but no difference in the relationships between rates of ascent and descent and dive depth was found among seasons. Furthermore, for all dive depths, ascent and descent rates were independent of the bottom duration. In all seasons post-dive intervals increased with dive duration and with dive depth, but they were longer in spring (2.3 min for dives over the 100-210 m layer) and summer than in autumn and winter (1.6-1.8 min). The diving efficiency decreased with increasing dive depth and was higher in autumn and winter (0.22-0.29) than in summer and spring (0.15-0.18). The large increase in bottom and dive duration from spring to winter is in agreement with the seasonal drop in prey density, with penguins spending more time searching for prey. In contrast, the consistency of the vertical velocity during contrasting conditions at-sea suggests that the transit time to depth is an important component of the foraging behaviour (scanning of the water column) that is independent of the prey availability. The time budget of the penguins during diving in a fluctuating environment appears to vary primarily during the bottom phase of the dives, with bottom duration increasing with diminishing prey supplies, while post-dive intervals shorten in the same time.  相似文献   

8.
将采用沙壤土培育的苦草幼苗(Vallisneria natans)放置于60-170 cm的水深范围内,每10 cm一个处理,使用光照计测定不同水深的光照强度并用 Skalar 水质流动分析仪测定试验水体的理化指标,观测苦草在不同的水深梯度下形态及叶片、叶绿素含量的变化,以研究水深梯度对沉水植物的个体生长发育及生物量的影响。试验结果表明:(1)试验30 d后,在60-130 cm的水深范围内,苦草株高随水深的增加而增高,在130-170 cm的水深范围内,随水深增加而降低,130 cm处苦草的平均高度最大,达67.9 cm;试验60 d,苦草的叶片长度、宽度、面积都随着水深的增加而减小,且在150-170 cm水深范围内叶片长度明显变小,苦草的生长速率随水深的增加显著下降;(2)不同试验组苦草的叶片长度、宽度、面积以及生长速率与水深呈明显的负相关(P<0.01),而叶绿素含量与水深呈明显的正相关(P<0.01),叶片面积在140-150 cm水深范围内减小最明显,且在140-160 cm水深范围内苦草各叶绿素指标较高,尤其是chl a+b在水深160 cm处高达1.876 mg·g^-1;(3)苦草叶片叶绿素含量随着水深的增加(光强降低)而升高,其变化幅度为0.369-1.876 mg·g^-1,其中叶片叶绿素a随着水深的增加呈波动递增的趋势,且在水深80 cm处较低(0.268 mg·g^-1),而叶绿素b在60-170 cm水深范围内总体较平稳,在水深160 cm处达到最高(0.505 mg·g^-1),chl a/b变化幅度较小,仅在2.49-2.84 mg·g^-1之间;(4)试验60 d后,各试验组总生物量的较大值主要集中在100-140 cm水深范围内,其中地上部分生物量平均占全株生物量的89.5%,地下部分生物量只占10.5%,生物量的分配随水深梯度变化不明显。说明水深梯度的变化对苦草的叶片生长及叶绿素含量有影响,但对生物量的分配作用不明显。研究表明在100-140 cm的水深范围内?  相似文献   

9.
A positive surge is a unsteady open channel flow resulting from the rapid rise of the free-surface. The phenomenon may be observed in water supply canals and channels as well as in some estuaries during spring tidal conditions. The formation and development of positive surges can be predicted using the method of characteristics and shallow water equations. The paper is the second part of a study presenting the results from new experimental investigations conducted in a large rectangular channel. Detailed unsteady velocity measurements were performed with a high temporal resolution using acoustic Doppler velocimetry and non-intrusive free-surface measurement devices. Several experiments were conducted with the same initial discharge (Q = 0.060 m3/s) and six different gate openings after closure resulting in both non-breaking undular and breaking bores. A comparison between main features of the undular surges with literature theories demonstrated that the experimental data were mostly in agreement with Andersen’s theory. The analysis of unsteady flow field including Reynolds stresses confirmed and extended previous findings about positive surge hydrodynamics.  相似文献   

10.
Fricke  H.  Meischner  D. 《Marine Biology》1985,88(2):175-187
Depth distribution, zonation pattern and growth morphology of 17 hermatypic and 4 ahermatypic coral species were investigated at eight different locations along the Bermuda platform with the research submersible GEO and by SCUBA diving in August–September 1983. Hermatypic coral growth occurs to a depth of 50 to 70 m, with a single Montastrea cavernosa growing at 78 m. Dominant forms in shallow-water coral communities are Diploria sp. and Porites astreoides, while M. cavernosa, Agaricia fragilis and Scolymia cubensis occur in deep-water associations below 60 m. Vertical visibilities (up to 178 m) and distribution of the photosynthetically active radiation revealed good light penetration values (1% level at about 100 m depth), which should favour hermatypic coral growth to a much greater depth than it actually occurs. Nor should the prevailing temperatures limit the depth of coral growth. Most deep-water hermatypes observed grow on remnants of Pleistocene reefs down to about 60 m. The vast areas of large massed rhodolith nodules below 50 to 60 m are unsuitable bottom for coral colonisation. Macroalgae growth seems to be the strongest factor controlling coral growth in deep water. Bermuda stony corals have a low growth form diversity. Various intraspecific morphs may occur at the same as well as at different depths, with a general trend towards flatter shapes with depth. Comparison with a similar study on Red Sea corals suggests that annual distribution of radiant energy on the most northern Atlantic reefs of Bermuda may be responsible for the occurrence of flat and cuplike growth forms in relatively shallow water, and for the shallower depth limits of hermatypic growth.  相似文献   

11.
在淀山湖0.5、1.0、1.5、2.0 m水层进行苦草(Vallisneria natans)现场生长实验,试图找出淀山湖苦草植被恢复的适宜环境条件。试验期间每天测量光照强度和水体溶解氧等环境因子,每隔10 d监测苦草的根长、株高、鲜重等生长指标的变化情况。结果表明:水深对苦草的生长具较显著影响,光照强度、溶氧分别和苦草日相对生长率具显著相关性;苦草的株高、叶片数、根长等生物学生长指标在0.5、1.0 m水层生长良好,1.5 m与2.0 m均出现不同程度的负增长状况。光照强度与苦草日相对生长率之间具有显著相关性(r=0.905),0.5、1.0 m水层的光照强度下苦草的相对生长率分别为0.14和0.11,而1.5、2.0 m水层的光照强度下苦草出现负增长的情况。水中的溶解氧含量与苦草成活率之间具有显著相关性(r=0.935)。随着水深的增加,溶氧逐渐降低,苦草的成活率也逐渐下降。在2.0 m水层处,溶氧均值为2.76 mg.L-1,苦草的成活率为46.5%,在1.0 m水层处,溶氧为5.66 mg.L-1,苦草成活率为86.5%,因此,淀山湖苦草群落恢复宜先在1.0 m以浅的沿岸带开展。  相似文献   

12.
In order to determine the effects of bottom depth and water column structure on slope distributions, lanternfishes in the Cape Canyon and Cape Point Valley (eastern South Atlantic) and from the Feni Ridge and Hebridean Terrace (eastern North Atlantic) were sampled with commercial midwater nets fished 10 m above the bottom at preselected depths. Fifty-one species were taken in the eastern South Atlantic and 17 species in the eastern North Atlantic. Analyses of the samples included clustering and multi-dimensional scaling ordination of the root-root transformed densities using the Bray-Curtis similarity index. In the eastern South Atlantic (March 1988), where the horizontal and vertical temperature structuring of the water column was marked, oceanic myctophids were not taken shallower than the 300 m-depth horizon and less than one-third of the total number of species occurred inside the 500 m-depth horizon. Temperatures and bottom depths correlated with the downslope distributions of the species. Down-slope zonation was apparent. In the eastern North Atlantic (May 1983), where the temperature structure of the water column was similar at all stations, no oceanic species were taken at the 200 m-depth horizon and more than one-half of the total number of species was taken at the 300 m-depth horizon. Species distributions correlated with depth, and downslope zonation was not evident. The data sets, as well as information from the West Florida slope, suggest that both the depth and the regional temperature structure of the water column effect the upper slope distributions of oceanic myctophids. Long-slope, ribbon distribution patterns (generalized down-slope zonation) in myctophids may therefore be localized artifacts produced by the interaction of these factors.  相似文献   

13.
In the heterogeneous marine environment, predators can increase foraging success by targeting physical oceanographic features, which often aggregate prey. For northern fur seals (Callorhinus ursinus), two prevalent oceanographic features characterize foraging areas during summer in the Bering Sea: a stable thermocline and a subsurface “cold pool”. The objective of this study was to examine the influence of these features on foraging behavior by equipping fur seals from St. Paul Island (Alaska, USA) with time-depth recorders that also measured water temperature. Foraging bout variables (e.g., mean dive depth and percent time diving in a bout) were compared with respect to subsurface thermal characteristics (thermocline presence and strength and cold pool presence). Over 74% of bouts occurred in association with strong thermoclines (temperature change > 5°C). Few differences were found for dive behavior in relation to the presence of a thermocline and the cold pool, but for epipelagic bouts, a strong thermocline resulted in increased bottom times, number of dive wiggles, and percent time diving when compared to moderate thermoclines. There was also a positive relationship between mean dive depth and thermocline depth. The combination of increasing foraging effort in areas with strong thermoclines and diving to depths closely related to the thermocline indicates this feature is important foraging habitat for northern fur seals and may act to concentrate prey and increase foraging success. By recognizing the environmental features northern fur seals use to find prey, managers will be better equipped to identify and protect foraging habitat that is important to northern fur seals, and possibly other marine predators in the Bering Sea.  相似文献   

14.
Deep-water assemblages of suprabenthic peracarids were analyzed in the SW Balearic Islands (Algerian Basin, southwestern Mediterranean) between 249 and 1,622 m depth; the patterns of species composition, possible zonation, and trophic structure found in this area were compared with those exhibited by peracarids in the mainland side of the Catalan Sea slope (northwestern Mediterranean). One hundred and four peracarid species (plus one leptostracan) were identified on the Balearic Islands slope, amphipods being the most diversified taxon (45 species). On the Balearic slope, two distinct depth assemblages were distinguished: one at the upper slope (US), between 249 and 402 m depth and the second at the deep slope, between 543 and 1,620 m depth. A remarkable species substitution occurred at depths between 402 and 638 m. In the Catalan Sea, in addition to the US assemblage occupying depths between 208 and 408 m, a second boundary of faunal change was found around 1,250 m. Suprabenthos biomass increased from 242 to approximately 500 m. Suprabenthos attained the highest biomass values (100 g wet weight/10,000 m2) at intermediate depths between 504 and 1,211 m, as also occurred with the associated zooplankton collected with suprabenthos (peak biomass between 502 m and 898 m). Suprabenthos biomass did not show any significant correlation with any environmental water-column variable. In contrast, zooplankton (especially small fish and decapod crustaceans) showed a significant positive correlation with fluorometry and turbidity at different levels of the water column. The feeding guilds of species showed important differences between the two areas only on the US, with a higher abundance of deposit feeders in the Catalan Sea (20.4%) than in the Balearic Islands (4.2%). The low contribution of deposit feeders in the SW Balearic Islands may ultimately be a consequence of the lack of river discharges in this area.Communicated by S.A. Poulet, Roscoff  相似文献   

15.
Miniature depth gauges were attached in December 1987 and January 1988 to Adélie penguins,Pygoscelis adeliae, breeding at Esperanza on the Antarctic Peninsula. Results from 34 birds showed that foraging penguins with eggs and with brooded and crèching chicks spent mean periods away from the nest of 96, 36 and 21 h, respectively, during which time means of 29.0 h (30%), 11.2 h (31%) and 2.7 h (13%), respectively, were spent under water at depths > 5 m. Time under water was positively correlated with time absent from nest. Maximum depth reached was 170 m but overall birds spent most time at shallower depths. Birds foraging for brooded chicks dived deeper than birds foraging for crèching chicks. Stomach-pumping indicated that the principal prey caught at this time was krill,Euphausia superba. Mean mass changes of adults during single foraging trips indicated that krill were caught at a mean rate of 7.2 g min–1 spent under water.  相似文献   

16.
The distribution of three important dissolved forms of nitrogen, viz. nitrate, nitrite and urea in the surface and bottom water samples collected from 27 selected hydrographic profiles, in the Arabian Sea, along the west coast of India is described. Of the three forms, nitrate concentrations were the highest and comparatively higher concentrations were observed in the bottom water. Decomposition of organic matter resulting in the release of the thermodynamically stable nitrogen species, i.e. nitrate, may be the major factor resulting in higher nitrate concentrations at these depths, where the water is also characterized by low values of dissolved oxygen and temperature. The significant positive correlation between A.O.U. and nitrate of the bottom water samples emphasizes the role of oxidative decomposition of organic matter which plays an active role in reducing the oxygen concentrations below the theoretical values since at this depth ( ≈200 m) the net production is taken to be zero. This is also evidenced by the negative correlation of nitrate with dissolved oxygen and temperature, for the bottom samples.  相似文献   

17.
铅锌冶炼厂渣堆场周边土壤铅污染特征   总被引:3,自引:0,他引:3  
金属冶炼过程留下的废渣经过雨水冲刷及渗滤液等的作用使残留在废渣中的重金属发生迁移转化,导致渣堆场下及周边土壤受到重金属污染。了解冶炼厂渣堆场下及周边土壤重金属污染状况对场地修复及土地利用规划均有重大意义。本研究调查了湖南某铅锌冶炼厂渣堆场0~4 km内三个采样区0~20 cm表层土壤及0~100 cm深度土壤中铅的污染状况,采用单项污染指数法进行铅污染评价,并分析了铅纵向迁移随深度变化和横向迁移随距离变化的分布特征。结果表明,铅锌冶炼厂渣堆场下及周边0~1 km范围内土壤受到了铅污染,渣堆场下、距渣堆场10 m处及1 km处表层土壤中重金属铅的质量分数分别可达775.25、645.33和309.80 mg·kg^-1,超过了当地土壤中铅的背景值,也超过了土壤二级质量标准甚至三级质量标准。三个采样区的铅污染指数分别为2.6、2.1及1.03,污染等级均为Ⅱ级,污染程度为轻度污染。三个采样区土壤中铅污染主要集中于0~20 cm土壤层中,铅的质量分数分别达775.25,645.33和309.80 mg·kg^-1,20~100 cm土壤层中铅的质量分数低于0~20 cm的,分别在88.48~120.96 mg·kg-1、235.01~380.16 mg·kg^-1及309.80~59.32 mg·kg^-1之间。渣堆场下土壤中的铅从0~20cm土壤层往下至20~100cm土壤层迁移量远小于距渣场10 m处及1 km处的。三个采样区表层0~20 cm土壤层中铅的变化规律为距渣堆场0 m(渣堆场下)〉距渣堆场10 m〉距渣堆场1 km〉距渣堆场4 km,质量分数随距离增加而降低。20~40 cm及40~60 cm土壤层中铅的变化规律为距渣堆场10 m〉距渣堆场1 km〉距渣堆场0 m处(渣堆场下)〉距渣堆场4 km,60~80 cm及80~100 cm土壤层的变化规律为距渣堆场10 m〉距渣堆场0 m处(渣堆场下)〉距渣堆场1 km〉距渣堆场4 km,铅的质量分数随距离的增加先升高后降低。  相似文献   

18.
Acoustic telemetry was used to examine patterns of activity and space utilisation of coelacanths, nocturnal predators which spend the day in submarine caves. Nine coelacanths (Latimeria chalumnae) were tracked, each for a period of 1 to 16 nights at Grande Comore, West Indian Ocean. Activities lasted on average 9 h, usually starting shortly after sunset and ending before sunrise. Vertically, coelacanths moved up and down at and below cave level by following the bottom contour, mainly between 180 and 400 m depth. The deepest record was 698 m, the shallowest 133 m. Most time was spent between 200 and 300 m depth. Large individuals performed deep excursions to depths below 400 m, usually once per night. The fish spent most time in water temperatures of 15 to 19 °C; they rarely ventured into waters warmer than 22 °C measured at depths shallower than 160 m depth. Horizontally, coelacanths stayed in narrow areas ranging from <1 to 10 km of coastline. Coelacanths are extremely slow drift-hunters with an estimated average swimming speed of 3.2 m min−1, often travelling not more than 3 km per night. They probably take advantage of local upwelling and downwelling and slow currents occurring parallel to the steep slopes. This study shows that coelacanths are inhabitants of the subphotic zone, where they are active mainly below the depth of their daytime refuges. Received: 7 July 1999 / Accepted: 11 February 2000  相似文献   

19.
K. F. Wishner 《Marine Biology》1980,60(2-3):179-187
Benthopelagic plankton was collected 10 to 100 m above the bottom of the deep sea (1100 to 3200 m) with an opening-closing net. Five samples were taken from the San Diego Trough, 2 from the eastern tropical Pacific and 2 from the northeast Atlantic. Roughly 80% of the individual organisms collected were copepods; the next most abundant groups were usually isopods, ostracods, and chaetognaths. Within the depth range studied, absolute abundances of most animal groups decreased with depth, but relative abundances (%) did not change. Of over 100 species of adult gymnopleid copepods collected, a few species were abundant, but most were rare. Copepod exoskeletons were more abundant than living copepods. For the gymnopleids, similarities among the 3 geographical regions were strong at the family and genus level but weak at the species level. Species that were widely distributed horizontally tended to occupy different positions (in terms of rank order of abundance) in each community. Gut content analysis indicated that the most abundant gymnopleids were generalized particle feeders. However, size-frequency distributions provided indirect evidence for niche separation. The deep-sea benthopelagic plankton was divisible into a truly planktonic component and a bottom-associated component. Abundances of some copepods changed relative to distance off the bottom. Several copepods and other animals, previously described as epibenthic or bottom-associated, were collected more that 10 m above the bottom and thus appeared to move freely between the bottom and lower water column.  相似文献   

20.
The origin of the deep-sea benthic fauna is poorly understood and represents an enormous gap in our understanding of basic evolutionary phenomena. One obstacle to studying evolutionary patterns in the deep sea has been the technical difficulty of measuring genetic variation in species that are typically minute, rare, and must be recovered from extreme depths. We used molecular genetic techniques to quantify variation in the 16S rRNA mitochondrial gene within and among populations of the common protobranch bivalve Deminucula atacellana (Schenck, 1939). We analyzed 89 individuals from nine samples collected in the 1960s along a depth gradient from 1100 to 3800 m in the western North Atlantic. Genetic variability within populations is much lower than between populations, and peak haplotype numbers occur near the center of its depth distribution. Continental slope (<2500 m) and rise (>2500 m) populations were genetically distinct despite the lack of any obvious topographic or oceanographic features that would impede gene flow. These findings indicate that the deep-sea macrofauna can have strong population structure over small (134 km) spatial scales, similar to that observed in shallow-water and terrestrial organisms. This surprisingly high biodiversity at the genetic level affords the potential for adaptation and evolutionary diversification, the ultimate historical causes of high species diversity in the deep-sea benthos. Received: 24 July 1997 / Accepted: 26 January 1998  相似文献   

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