Ecological Impacts of Revegetation and Management Practices of Ski Slopes in Northern Finland

Outdoor recreation and nature-based tourism represent an increasingly intensive form of land use that has considerable impacts on native ecosystems. The aim of this paper is to investigate how revegetation and management of ski runs influence soil nutrients, vegetation characteristics, and the possible invasion of nonnative plant species used in revegetation into native ecosystems. A soil and vegetation survey at ski runs and nearby forests, and a factorial experiment simulating ski run construction and management (factors: soil removal, fertilization, and seed sowing) were conducted at Ruka ski resort, in northern Finland, during 2003–2008. According to the survey, management practices had caused considerable changes in the vegetation structure and increased soil nutrient concentrations, pH, and conductivity on the ski runs relative to nearby forests. Seed mixture species sown during the revegetation of ski runs had not spread to adjacent forests. The experimental study showed that the germination of seed mixture species was favored by treatments simulating the management of ski runs, but none of them could eventually establish in the study forest. As nutrient leaching causes both environmental deterioration and changes in vegetation structure, it may eventually pose a greater environmental risk than the spread of seed mixture species alone. Machine grading and fertilization, which have the most drastic effects on soils and vegetation, should, therefore, be minimized when constructing and managing ski runs.     

Species composition and soil erosion on a ski area in Hokkaido,northern Japan

In 1986 and 1987, a study on factors governing revegetation on ski grounds was conducted at Teine ski ground (built in 1971) located near the city of Sapporo in northern Japan. Soil movement, slope gradient, distance from forest edge, vegetation cover, and number of species on the ski ground were examined. Although artificial sowing of exotic plants was undertaken in the whole ground surface at the time of opening, bare land occurred in ca. 50% of surveyed plots and the ski ground was mostly covered with native plants. The number of species was positively correlated to vegetation cover, which was low in the sites where intensive soil erosions occurred in winter. A principal component analysis of plant species distinguished three major groups of factors, i.e., vegetation cover (first axis, contribution rate 30.3%), soil erosion in winter and slope gradient (second, 23.1%), and distance from forest edge (third, 16.3%). The vegetation characteristics on the ski ground were not determined by a single environmental gradient but by the combination of factors described above. In particular, soil movements, which are mostly derived from snowmelt, are considered to be the initiator of vegetation changes.     

Prairie grass establishment on calcareous reclaimed mine soil

Reclaimed Appalachian surface mined lands have difficulty in sustaining native deciduous forest communities. Establishing prairie communities could increase ecosystem function; however, a native model system does not exist. We evaluated establishment of 15 North American prairie grasses as monocultures on reclaimed mine soil in southeast Ohio in four randomized complete blocks planted May 2005 and 2006. Population density was assessed 30 d after planting (30 DAP) and in October of the planting year (YR1) and second year following planting (YR2) and expressed as percentage of viable seeds sown (PVSS). Canopy cover of nonnative species reestablishing in the plots was measured in 2007. Eastern gamagrass ( L.) population was >50 PVSS in all censuses. Western wheatgrass [ (Rydb.) A. L?ve] was initially 7 PVSS at 30 DAP, but increased to 154 PVSS by YR2 from rhizomes spreading into gaps. Big bluestem ( Vitman) was 7 PVSS at 30 DAP and 4 PVSS at YR2. Blue grama [ (Willd. ex Kunth) Lag. ex Griffiths] and sideoats grama [ (Michx.) Torr.] did not survive past YR1. Gaps left from poor stand establishment were primarily recolonized by nonnative Kentucky bluegrass ( L.) in the 2005 planting and birdsfoot trefoil ( L.) in the 2006 planting, but was least in eastern gamagrass and tall dropseed [ (P. Beauv.) Kunth]. This research demonstrates the potential for increasing diversity and species richness on mine soil habitats with regionally native grasses that could increase functional quality through ecological resilience.     

A Systematic Review of Wild Burro Grazing Effects on Mojave Desert Vegetation,USA

Wild burros (Equus asinus), protected by the 1971 Wild Free-Roaming Horse and Burro Act on some federal lands but exotic animals many ecologists and resource mangers view as damaging to native ecosystems, represent one of the most contentious environmental management problems in American Southwest arid lands. This review synthesizes the scattered literature about burro effects on plant communities of the Mojave Desert, a center of burro management contentions. I classified 24 documents meeting selection criteria for this review into five categories of research: (i) diet analyses directly determining which plant species burros consume, (ii) utilization studies of individual species, (iii) control-impact comparisons, (iv) exclosure studies, and (v) forage analyses examining chemical characteristics of forage plants. Ten diet studies recorded 175 total species that burros consumed. However, these studies and two exclosure studies suggested that burros preferentially eat graminoid and forb groups over shrubs. One study in Death Valley National Park, for example, found that Achnatherum hymenoides (Indian ricegrass) was 11 times more abundant in burro diets than expected based on its availability. Utilization studies revealed that burros also exhibit preferences within the shrub group. Eighty-three percent of reviewed documents were produced in a 12-year period, from 1972 to 1983, with the most recent document produced in 1988. Because burros remain abundant on many federal lands and grazing may interact with other management concerns (e.g., desert wildfires fueled by exotic grasses), rejuvenating grazing research to better understand both past and present burro effects could help guide revegetation and grazing management scenarios.     

Establishment and Growth of Experimental Grass Species Mixtures on Coal Mine Sites Reclaimed with Municipal Biosolids

The Surface Mining Control and Reclamation Act of 1977 requires that coal mine sites in the United States be reclaimed to establish vegetative cover that is diverse, native, and capable of plant succession. However, there is a question as to whether vegetation established on coal mine sites reclaimed with biosolids is diverse and capable of plant succession. The influx of nutrients with the addition of biosolids leads to long-term dominance by early-successional species, most notably grasses, and consequently, a low establishment of woody and volunteer species. Additionally, many grass species commonly planted in reclamation have aggressive growth habits that lead to their dominance in coal mine plant communities. The establishment and growth of selected grass mixes was evaluated to determine whether alternative grass mixes would be less competitive with woody and volunteer species as compared to commonly used grass mixes. Percent vegetative cover, species richness, and the survival of direct-seeded woody species were assessed for each treatment grass mixture. It was found that Poa compressa and a mixture of P. compressa, Panicum virgatum, and Trifolium repens provided adequate coverage while still allowing the highest species richness and survival of woody species. Use of these species mixtures in coal mine reclamation with biosolids in the eastern United States would likely lead to establishment of a more species-rich plant community with a greater woody species component while still providing erosion control and site protection.     

Changes in Soil Properties After Establishment of Artemisia halodendron and Caragana microphylla on Shifting Sand Dunes in Semiarid Horqin Sandy Land, Northern China

In the semiarid Horqin sandy land of northern China, establishment of artificial sand-fixing shrubs on desertified sandy lands is an effective measure to control desertification and improve the regional environment. Caragana microphylla Lam. and Artemisia halodendron Turcz. ex Bess. are two of the dominant native shrub species, which are adapted well to windy and sandy environments, and thus, are widely used in revegetation programs to control desertification in Horqin region. To assess the effects of artificially planting these two shrub species on restoration of desertified sandy land, soil properties and plant colonization were measured 6 years after planting shrubs on shifting sand dunes. Soil samples were taken from two depths (0–5 cm and 5–20 cm) under the shrub canopy, in the mid-row location (alley) between shrub belts, and from nonvegetated shifting sand dune (as a control). Soil fine fractions, soil water holding capacity, soil organic C and total N have significantly increased, and pH and bulk density have declined at the 0–5-cm topsoil in both C. microphylla and A. halodendron. At the 5–20 cm subsurface soil, changes in soil properties are not significant, with exception of bulk density and organic C concentration under the canopy of A. halodendron and total N concentration under the canopy of C. microphylla. Soil amelioration processes are initiated under the shrub canopies, as higher C and N concentrations were found under the canopies compared with alleys. At the same time, the establishment of shrubs facilitates the colonization and development of herbaceous species. A. halodendron proved to have better effects in fixing the sand surface, improving soil properties, and restoring plant species in comparison to C. microphylla.     

Ski slope vegetation in central Honshu,Japan

To determine the status of the vegetation of ski slopes in northeastern-central Honshu, Japan, 94 plots (2×2 m) were set up on five ski areas (101–520 m elevation) which were established between 1945 and 1985 by forest clear-cutting, land modification, and seeding. Six vegetation types were recognized: five grasslands dominated byDigitaria adscendens, Miscanthus sinensis, Zoysia japonica, Festuca rubra, andPteridium aquilinum var.latiusculum, respectively, and bare areas of very low to no vegetation cover. Of the dominant species,F. rubra is the only introduced species; it does not, however, appear to persist. After the introduced grassland declinesM. sinensis or annual grasslands develop. Native plants, especially woody species, can establish inM. sinensis grassland but do not establish in the other grasslands. It is concluded that the introduction of exotic species is inappropriate to maintain ski slope vegetation, and the development ofM. sinensis grassland is desirable to promote natural revegetation.     

Assessing state-wide biodiversity in the Florida Gap analysis project

The Florida Gap (Fl-Gap) project provides an assessment of the degree to which native animal species and natural communities are or are not represented in existing conservation lands. Those species and communities not adequately represented in areas being managed for native species constitute 'gaps' in the existing network of conservation lands. The United States Geological Survey Gap Analysis Program is a national effort and so, eventually, all 50 states will have completed it. The objective of Fl-Gap was to provide broad geographic information on the status of terrestrial vertebrates, butterflies, skippers and ants and their respective habitats to address the loss of biological diversity. To model the distributions and potential habitat of all terrestrial species of mammals, breeding birds, reptiles, amphibians, butterflies, skippers and ants in Florida, natural land cover was mapped to the level of dominant or co-dominant plant species. Land cover was classified from Landsat Thematic Mapper (TM) satellite imagery and auxiliary data such as the national wetlands inventory (NWI), soils maps, aerial imagery, existing land use/land cover maps, and on-the-ground surveys. Wildlife distribution models were produced by identifying suitable habitat for each species within that species' range. Mammalian models also assessed a minimum critical area required for sustainability of the species' population. Wildlife species richness was summarized against land stewardship ranked by an area's mandates for conservation protection.     

Survival,reproduction, and recruitment of woody plants after 14 years on a reforested landfill

With the advent of modern sanitary landfill closure techniques, the opportunity exists for transforming municipal landfills into urban woodlands. While costs of fullscale reforestation are generally prohibitive, a modest planting of clusters of trees and shrubs could initiate or accelerate population expansions and natural plant succession from open field to diverse forest. However, among woody species that have been screened for use on landfills, these ecological potentials have not yet been investigated. We examined a 14-yr-old landfill plantation in New Jersey, USA, established to test tolerance of 19 species of trees and shrubs to landfill environments. We measured survivorship, reproduction, and recruitment within and around the experimental installation. Half of the original 190 plants were present, although survival and growth rates varied widely among species. An additional 752 trees and shrubs had colonized the plantation and its perimeter, as well as 2955 stems of vines. However, the great majority (>95%) of woody plants that had colonized were not progeny of the planted cohort, but instead belonged to 18 invading species, mostly native, bird-dispersed, and associated with intermediate stages of secondary plant succession. Based on this evidence, we recommend that several ecological criteria be applied to choices of woody species for the restoration of municipal landfills and similar degraded sites, in order to maximize rapid and economical establishment of diverse, productive woodlands.     

Investing in rangeland restoration in the Arid West,USA: Countering the effects of an invasive weed on the long-term fire cycle

In large areas of the arid western United States, much of which are federally managed, fire frequencies and associated management costs are escalating as flammable, invasive cheatgrass (Bromus tectorum) increases its stronghold. Cheatgrass invasion and the subsequent increase in fire frequency result in the loss of native vegetation, less predictable forage availability for livestock and wildlife, and increased costs and risk associated with firefighting. Revegetation following fire on land that is partially invaded by cheatgrass can reduce both the dominance of cheatgrass and its associated high fire rate. Thus restoration can be viewed as an investment in fire-prevention and, if native seed is used, an investment in maintaining native vegetation on the landscape. Here we develop and employ a Markov model of vegetation dynamics for the sagebrush steppe ecosystem to predict vegetation change and management costs under different intensities and types of post-fire revegetation. We use the results to estimate the minimum total cost curves for maintaining native vegetation on the landscape and for preventing cheatgrass dominance. Our results show that across a variety of model parameter possibilities, increased investment in post-fire revegetation reduces long-term fire management costs by more than enough to offset the costs of revegetation. These results support that a policy of intensive post-fire revegetation will reduce long-term management costs for this ecosystem, in addition to providing environmental benefits. This information may help justify costs associated with revegetation and raise the priority of restoration in federal land budgets.     

Control of Tamarix in the Western United States: Implications for Water Salvage, Wildlife Use, and Riparian Restoration

Non-native shrub species in the genus Tamarix (saltcedar, tamarisk) have colonized hundreds of thousands of hectares of floodplains, reservoir margins, and other wetlands in western North America. Many resource managers seek to reduce saltcedar abundance and control its spread to increase the flow of water in streams that might otherwise be lost to evapotranspiration, to restore native riparian (streamside) vegetation, and to improve wildlife habitat. However, increased water yield might not always occur and has been substantially lower than expected in water salvage experiments, the potential for successful revegetation is variable, and not all wildlife taxa clearly prefer native plant habitats over saltcedar. As a result, there is considerable debate surrounding saltcedar control efforts. We review the literature on saltcedar control, water use, wildlife use, and riparian restoration to provide resource managers, researchers, and policy-makers with a balanced summary of the state of the science. To best ensure that the desired outcomes of removal programs are met, scientists and resource managers should use existing information and methodologies to carefully select and prioritize sites for removal, apply the most appropriate and cost-effective control methods, and then rigorously monitor control efficacy, revegetation success, water yield changes, and wildlife use.     

Ecological restoration as a strategy for conserving biological diversity

Though the restoration of disturbed ecosystems has so far played a relatively modest role in the effort to conserve biological diversity, there are reasons to suspect that its role will increase and that its contribution to the maintenance of diversity will ultimately prove crucial as techniques are further refined and as pristine areas for preservation become scarcer and more expensive.It is now possible to restore a number of North American communities with some confidence. However, it should be noted that many current efforts to return degraded lands to productive use, like attempts to reclaim land disturbed by mining, try only for rehabilitation to a socially acceptable condition and fall considerably short of actually restoring a native ecological community.Possible uses for restoration in the conservation of biodiversity include not only the creation of habitat on derelict sites, but also techniques for enlarging and redesigning existing reserves. Restoration may even make it possible to move reserves entirely in response to long-term events, such as changes in climate. Restoration in the form of reintroduction of single species to preexisting or restored habitat is also a critical link in programs to conserve species ex situ in the expectation of eventually returning them to the wild. And restoration provides opportunities to increase diversity through activities as diverse as management of utility corridors, transportation rights-of-way, and parks.     

Using native plants as problem-solvers

The Federal Highway Administration encourages state highway agencies to use native plants in erosion control, revegetation, and landscaping solutions. This paper explains both policy reasons and technical reasons for the use of native plants. How native species can be used is shown through a roadside case study. Other applications of native plant use will be explained through a plant community approach.     

Revegetation of an abandoned uranium millsite on the Colorado Plateau, Arizona

We attempted to restore native plants on disturbed sites at a former uranium mill on the Colorado Plateau near Tuba City, AZ. Four-wing saltbush [Atriplex canescens (Pursh) Nutt.] was successfully established in compacted caliche soil and in unconsolidated dune soil when transplants were irrigated through the first summer with 20 L/plant/wk. The caliche soil was ripped before planting to improve water-holding capacity. The diploid saltbush variety, angustifolia, had higher survival and growth than the common tetraploid variety, occidentalis, especially on dune soil. The angustifolia variety grew to 0.3 to 0.4 m3 per plant over 3 yr even though irrigation was provided only during the establishment year. By contrast, direct seeding of a variety of native forbs, grasses, and shrubs yielded poor results, despite supplemental irrigation throughout the first summer. In this arid environment (precipitation = 100 to 200 mm/yr), the most effective revegetation strategy is to establish keystone native shrubs, such as four-wing saltbush, using transplants and irrigation during the establishment year, rather than attempting to establish a diverse plant community all at once.     

Private land management for biodiversity conservation

As human influences fragment native communities and ecosystems, remaining land must be better managed to conserve many elements of biodiversity. Much of this land is privately held, yet traditional private land-use management practices often further diminish biodiversity by promoting favored or edge-adapted species. Today, private land stewards are increasingly aware of and concerned about biodiversity, but little guidance exists for them to make land-use decisions incorporating principles and knowledge from conservation biology. Consequently, most management strategies are highly subjective. This article addresses that problem by introducing current conservation wisdom to management and use of private lands. The result is a model program for developing land management plans, with the goal of maintaining viable populations and natural distributions of native species and communities from a landscape perspective. The program establishes a protocol for classifying sites according to the importance of their species, communities, and other elements to global and regional biodiversity. These site classifications are based on the management objectives necessary to maintain important elements. Once managers classify a site, the program provides management standards, general stewardship principles, examples of land management strategies, and basic monitoring and evaluation procedures.     

US army land condition-trend analysis (LCTA) program

The US Army Land Condition-Trend Analysis (LCTA) program is a standardized method of data collection, analysis, and reporting designed to meet multiple goals and objectives. The method utilizes vascular plant inventories, permanent field plot data, and wildlife inventories. Vascular plant inventories are used for environmental documentation, training of personnel, species identification during LCTA implementation, and as a survey for state and federal endangered or threatened species. The permanent field plot data documents the vegetational, edaphic, topographic, and disturbance characteristics of the installation. Inventory plots are allocated in a stratified random fashion across the installation utilizing a geographic information system that integrates satellite imagery and soil survey information. Ground cover, canopy cover, woody plant density, slope length, slope gradient, soil information, and disturbance data are collected at each plot. Plot data are used to: (1) describe plant communities, (2) characterize wildlife and threatened and endangered species habitat, (3) document amount and kind of military and nonmilitary disturbance, (4) determine the impact of military training on vegetation and soil resources, (5) estimate soil erosion potential, (6) classify land as to the kind and amount of use it can support, (7) determine allowable use estimates for tracked vehicle training, (8) document concealment resources, (9) identify lands that require restoration and evaluate the effectiveness of restorative techniques, and (10) evaluate potential acquisition property. Wildlife inventories survey small and midsize mammals, birds, bats, amphibians, and reptiles. Data from these surveys can be used for environmental documentation, to identify state and federal endangered and threatened species, and to evaluate the impact of military activities on wildlife populations. Short- and long-term monitoring of permanent field plots is used to evaluate and adjust land management decisions.     

Assessing conservation opportunity on private land: socio-economic, behavioral, and spatial dimensions

This study presents a method for assessing conservation opportunity on private land based on landholders' socio-economic, behavioral, and farm characteristics. These characteristics include age, gender, education, level of off-farm income, farm size, proportion of remnant native vegetation on-farm, and ecological value of native vegetation on-farm. A sample of landholders who own greater than 2 ha of land in the South Australian Murray-Darling Basin region were sent a mail-based survey about their values and preferences for environmental management (N = 659, 52% response). Cross-tabulations and ANOVA statistical analysis techniques were used to compare the socio-economic attributes across three landholder classes: disengaged, moderately engaged, and highly engaged in native vegetation planting. Results indicate that highly engaged landholders were more likely to be female, formally educated, hobby farmers who managed small parcels of land and have high off-farm incomes, whereas disengaged landholders held significantly stronger farming connections (more farming experience, family have lived on the farm for more generations). Spatial analysis revealed area-specific differences in conservation opportunity and conservation priority. In some areas, properties of high ecological value were managed by highly engaged landholders, but nearby properties of high value were managed by moderately engaged or disengaged landholders. Environmental managers therefore cannot assume areas of high conservation priority will be areas of high conservation opportunity. At the regional scale, the potential for revegetation seems most promising within the moderately engaged landholder group considering the vast amount of land managed by this group in areas of high ecological value, particularly within the less represented Mallee and Coorong and Rangelands sub-regions. We suggest that incentive schemes which purchase conservation need to be targeted at disengaged landholders; mentoring schemes led by commercial farmers highly engaged in native vegetation planting should be directed at moderately engaged landholders, and; awards programs which acknowledge conservation successes should be targeted at highly engaged landholders.     

Performance Criteria, Compliance Success, and Vegetation Development in Compensatory Mitigation Wetlands

The US Army Corps of Engineers often requires wetland creation or restoration as compensation for wetlands damaged during development. These wetlands are typically monitored postconstruction to determine the level of compliance with respect to site-specific performance standards. However, defining appropriate goals and measuring success of restorations has proven difficult. We reviewed monitoring information for 76 wetlands constructed between 1992 and 2002 to summarize the performance criteria used to measure progress, assess compliance with those criteria, and, finally, to evaluate the appropriateness of those criteria. Goals were overwhelmingly focused on plant communities. Attributes used to assess the quality of restored plant communities, including percent native species and the Floristic Quality Index, increased over time but were apparently unrelated to the number of species planted. Compliance frequencies varied depending on site goals; sites often failed to comply with criteria related to survival of planted vegetation or requirements that dominant plant species should not be exotic or weedy, whereas criteria related to the establishment of cover by vegetation or by wetland-dependent plants were often met. Judgment of a site’s success or failure was largely a function of the goals set for the site. Some performance criteria were too lenient to be of value in distinguishing failed from successful sites, whereas other criteria were unachievable without more intensive site management. More appropriate goals could be devised for restored wetlands by basing performance standards on past performance of similar restorations, identifying consistent temporal trends in attributes of restored sites, and using natural wetlands as references.     

Maximizing Benefits from Riparian Revegetation Efforts: Local- and Landscape-Level Determinants of Avian Response

With limited financial resources available for habitat restoration, information that ensures and/or accelerates success is needed to economize effort and maximize benefit. In the Central Valley of California USA, riparian habitat has been lost or degraded, contributing to the decline of riparian-associated birds and other wildlife. Active restoration of riparian plant communities in this region has been demonstrated to increase local population sizes and species diversity of landbirds. To evaluate factors related to variation in the rate at which bird abundance increased after restoration, we examined bird abundance as a function of local (restoration design elements) and landscape (proportion of riparian vegetation in the landscape and riparian patch density) metrics at 17 restoration projects within five project areas along the Sacramento River. We developed a priori model sets for seven species of birds and used an information theoretic approach to identify factors associated with the rate at which bird abundance increased after restoration. For six of seven species investigated, the model with the most support contained a variable for the amount of riparian forest in the surrounding landscape. Three of seven bird species were positively correlated with the number of tree species planted and three of seven were positively correlated with the planting densities of particular tree species. Our results indicate that restoration success can be enhanced by selecting sites near existing riparian habitat and planting multiple tree species. Hence, given limited resources, efforts to restore riparian habitat for birds should focus on landscape-scale site selection in areas with high proportions of existing riparian vegetation.     

Effects of Alien Plants on Ecosystem Structure and Functioning and Implications for Restoration: Insights from Three Degraded Sites in South African Fynbos

We investigated the type and extent of degradation at three sites on the Agulhas Plain, South Africa: an old field dominated by the alien grass Pennisetum clandestinum Pers. (kikuyu), an abandoned Eucalyptus plantation, and a natural fynbos community invaded by nitrogen fixing—Australian Acacia species. These forms of degradation are representative of many areas in the region. By identifying the nature and degree of ecosystem degradation we aimed to determine appropriate strategies for restoration in this biodiversity hotspot. Vegetation surveys were conducted at degraded sites and carefully selected reference sites. Soil-stored propagule seed banks and macro- and micro-soil nutrients were determined. Species richness, diversity and native cover under Eucalyptus were extremely low compared to the reference site and alterations of the soil nutrients were the most severe. The cover of indigenous species under Acacia did not differ significantly from that in reference sites, but species richness was lower under Acacia and soils were considerably enriched. Native species richness was much lower in the kikuyu site, but soil nutrient status was similar to the reference site. Removal of the alien species alone may be sufficient to re-initiate ecosystem recovery at the kikuyu site, whereas active restoration is required to restore functioning ecosystems dominated by native species in the Acacia thicket and the Eucalyptus plantation. To restore native plant communities we suggest burning, mulching with sawdust and sowing of native species.