Proximate causes of natal dispersal in female yellow-bellied marmots, Marmota flaviventris

We investigated factors influencing natal dispersal in 231 female yearling yellow-bellied marmots (Marmota flaviventris) using comprehensive analysis of 10 years (1983-1993) of radiotelemetry and 37 years (1963-1999) of capture-mark-recapture data. Only individuals whose dispersal status was verified, primarily by radiotelemetry, were considered. Univariate analyses revealed that six of the 24 variables we studied significantly influenced dispersal: dispersal was less likely when the mother was present, amicable behavior with the mother and play behavior were more frequent, and spatial overlap was greater with the mother, with matriline females, and with other yearling females. Using both univariate and multivariate analyses, we tested several hypotheses proposed as proximate causes of dispersal. We rejected inbreeding avoidance, population density, body size, social intolerance, and kin competition as factors influencing dispersal. Instead, our results indicate that kin cooperation, expressed via cohesive behaviors and with a focus on the mother, influenced dispersal by promoting philopatry. Kin cooperation may be an underappreciated factor influencing dispersal in both social and nonsocial species.     

Human preferences for facial masculinity change with relationship type and environmental harshness

In humans (Homo sapiens), sexual dimorphism in face shape has been proposed to be linked to quality in both men and women. Although preferences for high-quality mates might be expected, previous work has suggested that high quality may be associated with decreased investment in partnerships. In line with a trade-off between partner quality and investment, human females have been found to prefer higher levels of masculinity when judging under conditions where the benefits of quality would be maximised and the costs of low investment would be minimised. In this study, we examined facultative preferences for masculinity/femininity under hypothetical high and low environmental harshness in terms of resource availability in which participants were asked to imagine themselves in harsh/safe environments. We demonstrate that environmental harshness influences preferences for sexual dimorphism differently according to whether the relationship is likely to be short or long term. Women prefer less-masculine male faces and men prefer less-feminine female faces for long-term than short-term relationships under conditions of environmental harshness. Such findings are consistent with the idea that high-quality partners may be low investors and suggest that under harsh ecological conditions, both men and women favour a low-quality/high-investment partner for long-term relationships. For short-term relationships, where investment is not an important variable, preferences for sexual dimorphism were similar for the low and high environmental harshness conditions. These results provide experimental evidence that human preferences may be contingent on the environment an individual finds itself inhabiting.     

Temporal persistence of biological patch structure in an abyssal benthic community

An analysis of living and dead components of a population of deep-sea agglutinated Foraminifera reveals that both segments of the fauna are dispersed nonrandomly on a scale of centimeters. Significant positive correlations between the distribution of living and dead individuals in several species imply that small-scale patch structure in this environment may persist for more than one generation. Mechanisms maintaining such structure may include habitat selection and reproductive patterns and suggest that dispersion pattern is the result of an active interaction between amimals and a heterogeneous physical and biotic environment.     

Separability,functional structure and aggregation for a class of models in environmental economics

The purpose of this paper is to provide a theoretical bridge between the many-commodity general equilibrium models and the simpler one good-one pollutant models used in environmental economics. The interest is in determining the conditions that will permit a vector of pollutants (or goods) to be treated as if it were only one quantity, the amount of environmental pollution (or consumption). A general equilibrium model is constructed and the concepts of separability and functional structure are applied to examine the requirements for consistent aggregation. It is shown that this aggregation process may in general produce two entirely different sets of aggregate measures of pollution and consumption. This result is of theoretical and empirical significance. Alternative criteria for aggregation suggest that the aggregation depends upon very stringent features of the economy concerned.     

Stability and structure of a fish community on a coral patch reef in Hawaii

Two complete collections of the fishes residing on an isolated coral patch reef ( 1500 m²) at Oahu, Hawaii, were made 11 years apart. Of the 112 species of fishes in both collections combined, only 40% were in common, but these made up more than 85% of the wet biomass in each collection. The two assemblages of fishes were similar in trophic structure and standing crop. Many coral reef fish communities are dominated by carnivorous forms. In the present study, planktivorous fishes were the most important trophic group in the community; this was related to abundant zooplankton resources. Following the second collection in 1977, recolonization by fishes was followed for 1 year. Recolonization proceeded rapidly and was primarily by juvenile fishes well beyond larval metamorphosis. Within 6 months of the second collection, the trophic structure had been re-established. The MacArthur-Wilson model of insular colonization described the recolonization process and predicted an equilibrium situation in less than 2 years. The recolonization data suggested that chance factors may explain the colonization process on a small scale, but a relatively deterministic pattern emerged when considering the entire reef. Thus, at the community level the fishes are a persistent and predictable entity.     

青藏高原东部原始林下地表主要苔藓斑块特征及其影响因素

引入并应用斑块概念对大渡河上游藓类-冷杉原始林下地表1hm^2面积的锦丝藓、大羽藓、赤茎藓、塔藓、锦丝-赤茎藓等五种地表苔藓斑块结构特征作了调查分析，并同时测定了斑块相关环境因子（空气温度、空气湿度、基质湿度、光照强度、乔木盖度、灌木盖度、草本盖度及生物量、凋落物盖度）。对数据进行综合分析发现，不同斑块类型在生物量、个体密度、层片厚度和物种丰富度上都表现出一定的差异性，所调查环境因素各个方面在五种斑块之间都表现出较强的差异性．且各类斑块的结构特征与环境因子间分别表现山很强的相关性。影响斑块结构特征的最主要因素是藓丛表面空气温度，空气湿度、光照强度和草本层盖度、灌木层的盖度等也对斑块特征产生重要影响，而草本层生物器、凋落物盖度及乔木层盖度仅对个别斑块的结构特征产生影响，基质湿度并未对斑块特征产生显著影响，可能是因为不同藓类生长基质不同所致。     

Linking richness, community variability, and invasion resistance with patch size

The influence of community dynamics on the success or failure of an invasion is of considerable interest. What has not been explored is the influence of patch size on the outcomes of invasions for communities with the same species pool. Here we use an empirically validated spatial model of a marine epibenthic community to examine the effects of patch size on community variability, species richness, invasion, and the relationships between these variables. We found that the qualitative form of the relationship between community variability and species richness is determined by the size of the model patch. In small patches, variability decreases with species richness, but beyond a critical patch size, variability increases with increasing richness. This occurs because in large patches large, long-lived colonies attain sufficient size to minimize mortality and dominate the community, leading to decreased species richness and community variability. This mechanism cannot operate on smaller patches where the size of colonies is limited by the patch size and mortality is high irrespective of species identity. Further, invasion resistance is strongly correlated with community variability. Thus, the relationship between species richness and invasion resistance is also determined by patch size. These patterns are generated largely by an inverse relationship between colony size and mortality, and they depend on the spatial nature and patch size of the community. Our results suggest that a continuum of possible relationships can exist between species richness, community variability, invasion resistance, and area. These relationships are emergent behaviors generated by the individual properties of the particular component species of a community.     

Group foraging, patch exploitation time and the finder's advantage

 We use a combination of the marginal value theorem (MVT) of Charnov (1976), and a group foraging model featuring information sharing to address patch residence in an environment where food occurs in discrete patches. We shall show that among equal competitors the optimal patch time for the individual that finds the food patch is shorter than that for the non-finder among equal competitors, T _E < T _N. This is the case if the patch-finder commences food harvesting in the patch earlier and manages to monopolise a fraction of the prey items (finder's advantage) before the other individuals come to take their benefit. When individuals differ in their food-searching abilities so that some of them (producers) contribute proportionally more to food-searching than others (scroungers), and differ in ability to compete for the food found, a difference emerges between producer and scrounger individuals in the optimal patch time. Within a patch we always have the finder's advantage (T _E < T _N) regardless of phenotype. Between patches a suite of optimal patch times for encountering individuals emerges depending on the performance of producers and scroungers when changing from solitary feeding to feeding in a group. The optimal patch time for individuals that are affected more severely by competition is shorter than that for individuals of the phenotype with better competitive ability. When both phenotypes are affected similarly no difference in optimal patch times emerges. Received: 13 February 1996 / Accepted after revision: 28 September 1996     

Microscopic structure of soil Fe-Mn nodules: environmental implication

The internal structure, iron and manganese distributions of Fe-Mn nodules from imperfectly drained soils of central Greece were studied by scanning electron microscopy (SEM) and elemental analysis. Nodules exhibited a differentiated structure with discontinuous concentric rings as expression of their redox history. Fe and Mn have shown distinct repartition and were distributed mainly in concentric bands. The microstructure and the arrangement of iron—manganese compounds reflect the periodical development and the relative slow accretion rates of the studied nodules. These properties provide ample opportunity for adsorption and isolation of heavy metals from the soil system.     

Mate guarding and gallivanting by male hoary marmots (Marmota caligata)

Summary Seven years data on the vernal behavior of hoarty marmots, Marmota caligata, suggest that males engage in a two-part reproductive strategy, which consists of guarding their mates against possible copulation with additional males, and also gallivanting — wandering about in search of additional reproductive opportunities for themselves. Data are presented which support seven predictions derived from the assumption that mate guarding and gallivanting are parts of a reproductive strategy by male marmots.     

北仑河口海域小型硅藻群落结构及其与环境因子的关系

北仑河口地处亚热带-热带地区,气候温暖湿润,是我国北部湾沿岸与越南交界的一个重要入海河口。于2008年3、5、8和11月分别采集该海域小型浮游硅藻,同时测定水温、盐度等水体参数,以及氮、磷、硅等18项水化学指标,分析了小型硅藻群落结构变化与环境因子之间的关系。该河口区的小型硅藻主要由纤细翼鼻状藻Proboscia alata f.gracillima、拟旋链角毛藻Chaetoceros pseudocurvisetus、细弱海链藻Thalassiosira subtilis、中华齿状藻Odontella sinensis和菱形藻Nitzschiasp.等优势种类组成,并且存在明显的季节性演替。应用典范对应分析,发现影响北仑河口硅藻群落结构的主要环境因子为温度、盐度、N、Si、Hg和悬浮体质量浓度等,且不同季节的主控因素有所差异。菱形藻对温度和Cu、As、Zn和Hg等重金属的耐受性较强。纤细翼鼻状藻适宜于生长在水体透明度高的海域,悬浮体质量浓度以6.27 mg.L-1以下为宜。拟旋链角毛藻受Cr的影响较大,高含量的Cr会限制其生长。而细弱海链藻和中华齿状藻对Hg的耐受性较强。     

Genetic relatedness and space use in a behaviorally flexible species of marmot, the woodchuck (Marmota monax)

Solitary species show several patterns of space use and relatedness. Individuals may associate randomly or may live near female or male kin, often as a result of natal philopatry or dispersal patterns. Although usually described as solitary or asocial, woodchucks (Marmota monax) are behaviorally flexible marmots that exhibit greater sociality in some populations than others. I examined relationships between kinship, geographic distance, and home range overlap, as well as dispersal and philopatry, to determine the extent to which kin associated spatially. I used a combination of microsatellite DNA analysis, long-term behavioral observations, and radiotelemetry to test predictions that females, but not males, would associate with kin. Indeed, woodchucks lived closer and shared a greater proportion of their home range with more closely related animals. Overlap of females' and males' home ranges was positively correlated with kinship, and male–female dyads shared more area with closer kin. Most juveniles delayed dispersal beyond their first summer. Females often remained philopatric and settled near their natal range. Although males often dispersed as yearlings, some males also established territories within or immediately adjacent to their natal home ranges. A combination of factors can explain these spatial patterns, including high population density associated with the study site's location within a suburban environment, high dispersal costs, and abundant food. Thus, despite their asocial and solitary reputation, woodchucks displayed spatial patterns seen in other, more social species of ground-dwelling sciurids.     

Regulator reputation, enforcement, and environmental compliance

This paper explores empirically the impact of enforcement efforts on environmental compliance, focusing on the role of regulator reputation spillover effects. We find that, on the margin, the impact of a fine for water pollutant violations is about a two-thirds reduction in the statewide violation rate in the year following a fine. This large result obtains through the regulator's enhanced reputation; the deterrence impact on other plants in a state is almost as strong as the impact on the sanctioned plant. Focusing only on the response of the sanctioned plant, as in previous studies, may therefore seriously underestimate the efficacy of fines and other sanctions. This paper also examines the relative effectiveness of monitoring and enforcement instruments. Non-monetary sanctions contribute no detected impact on compliance, and the marginal fine induces substantially greater compliance than the marginal inspection.     

北仑河口海域小型硅藻群落结构及其与环境因子的关系

北仑河口地处亚热带-热带地区,气候温暖湿润,是我国北部湾沿岸与越南交界的一个重要入海河口。于2008年3、5、8和11月分别采集该海域小型浮游硅藻,同时测定水温、盐度等水体参数,以及氮、磷、硅等18项水化学指标,分析了小型硅藻群落结构变化与环境因子之间的关系。该河口区的小型硅藻主要由纤细翼鼻状藻Proboscia alata f.gracillima、拟旋链角毛藻Chaetoceros pseudocurvisetus、细弱海链藻Thalassiosira subtilis、中华齿状藻Odontella sinensis和菱形藻Nitzschiasp.等优势种类组成,并且存在明显的季节性演替。应用典范对应分析,发现影响北仑河口硅藻群落结构的主要环境因子为温度、盐度、N、Si、Hg和悬浮体质量浓度等,且不同季节的主控因素有所差异。菱形藻对温度和Cu、As、Zn和Hg等重金属的耐受性较强。纤细翼鼻状藻适宜于生长在水体透明度高的海域,悬浮体质量浓度以6.27 mg.L-1以下为宜。拟旋链角毛藻受Cr的影响较大,高含量的Cr会限制其生长。而细弱海链藻和中华齿状藻对Hg的耐受性较强。     

Plant vintages, grandfathering, and environmental policy

Environmental regulations that grandfather existing plants, by not holding them to the same strict standards as new plants, may have the unintended consequence of retarding new investment. If new plants are cleaner, then this effect may increase pollution in the short run. I develop a dynamic model of a facility’s decisions over scrapping and abatement, which depend on capital depreciation, profitability shocks, and environmental policy. Using data from fossil fuel fired boilers at electric power plants, I estimate the structural parameters of the model and assess the impact of grandfathering in the Clean Air Act on sulfur dioxide emissions. Counterfactual policy simulations show that an increase in the stringency of performance standards would have led to a decrease in investment in new boilers. However, this does not lead to increased emissions, since there is less investment in dirtier coal boilers as compared to relatively cleaner oil or natural gas boilers.     

Epiphyte metapopulation dynamics are explained by species traits, connectivity, and patch dynamics

The colonization-extinction dynamics of many species are affected by the dynamics of their patches. For increasing our understanding of the metapopulation dynamics of sessile species confined to dynamic patches, we fitted a Bayesian incidence function model extended for dynamic landscapes to snapshot data on five epiphytic lichens among 2083 mapped oaks (dynamic patches). We estimate the age at which trees become suitable patches for different species, which defines their niche breadth (number of suitable trees). We show that the colonization rates were generally low, but increased with increasing connectivity in accordance with metapopulation theory. The rates were related to species traits, and we show, for the first time, that they are higher for species with wide niches and small dispersal propagules than for species with narrow niches or large propagules. We also show frequent long-distance dispersal in epiphytes by quantifying the relative importance of local dispersal and background deposition of dispersal propagules. Local stochastic extinctions from intact trees were negligible in all study species, and thus, the extinction rate is set by the rate of patch destruction (tree fall). These findings mean that epiphyte metapopulations may have slow colonization-extinction dynamics that are explained by connectivity, species traits, and patch dynamics.     

Genetic structure of two species of Montipora on a patch reef: conflicting results from electrophoresis and histocompatibility

Spatial patterns of graft acceptance and rejection were different for two species of the coral Montipora from Kaneohe Bay, Hawaii. Electrophoretically distinct tissues were capable of fusing, indicating that clonal identity is not necessarily inferred by acceptance of grafts. For M. dilatata, 18 of 40 cases of fusion were between dissimilar genotypes and for M. verrucosa, 3 of 7 cases. In one case, fusion occurred between genotypes which shared no alleles. A greater understanding of the genetics of invertebrate immunology is required before histocompatibility criteria can be used independently to assess population genetic structure with confidence.     

Development of environmental policy,objectives, and targets

This paper describes a process to develop and establish an environmental policy and its associated environmental targets, objectives, and management programmes. The environmental policy is defined in a statement which addresses the company expectations for programmes that are important to assure adequate environmental protection and define its future commitment towards preventing pollution and continuously improving the effectiveness of its environmental protection programmes. A set of environmental objectives, derived from the policy, define the overall goals which will ensure that the company's environmental performance will remain consistent with the commitment identified in its policy when its environmental management system (EMS) is implemented, Environmental targets reflect the requirements and standards that the company must strive to achieve each objective and minimize any harmful environmental impacts from its production and manufacturing processes or other activities. In addition to defining specific targets, environmental management programmes are also used to guide EMS implementation. These programmes, however, must provide the company with the necessary flexibility to select the most appropriate technologies to enhance its own processes and prudently allocate its available financial resources in areas that return the greatest overall environmental benefit.Though they are expected to allow some discretion in the allocation of resources targeted towards environmental protection, EMS programmes must also provide assurance that the corporation environmental policy, objectives, targets, and management programmes do not result in a facility failure to adhere to all of its certification requirements. The principle and factors described in this article should be considered to the extent that they are necessary and practical.     

Feedbacks between bivalve density, flow, and suspended sediment concentration on patch stable states

We explore the role of biophysical feedbacks occurring at the patch scale (spatial scale of tens of meters) that influence bivalve physiological condition and affect patch stability by developing a numerical model for the pinnid bivalve, Atrina zelandica, in cohesive sediments. Simulated feedbacks involve bivalve density, flow conditions (assumed to be primarily influenced by local water depth and peak current speed), suspended sediment concentration (evaluated through a balance between background concentration, deposition, and erosion), and changes in the physiology of Atrina derived from empirical study. The model demonstrates that high bivalve density can lead to skimming flow and to a concomitant decrease in resuspension that will affect suspended sediment concentration over the patch directly feeding back on bivalve physiology. Consequently, for a given flow and background suspended sediment load, the stability of a patch directly depends on the size and density of bivalves in the patch. Although under a range of conditions patch stability is ensured independently of bivalve density, simulations clearly indicate that sudden changes in bivalve density or suspended sediment concentration can substantially affect patch structure and lead to different stable states. The model highlights the role of interactions between organisms, flow, and broader scale environmental conditions in providing a mechanistic explanation for the patchy occurrence of benthic suspension feeders.     

Corruption, political competition and environmental policy

This paper examines the interplay between corruption, political competition, environmental policy and environmental outcomes. An important feature of the model, which has been neglected in the existing literature, is that corruption may occur at different levels of government, such as the payment of bribes to politicians who determine policies, or bureaucrats who administer environmental regulations. We analyse the relationship between political competition and environmental outcomes in a model of stratified corruption and identify the benefits and limits of political competition. Our results suggest that while political competition may yield policy improvements, it cannot eliminate corruption at all levels of government.