Constraints on group size in primates and carnivores: population density and day-range as assays of exploitation competition

Summary We hypothesise that foraging group size (FGS) and population group size (PGS) in primates and carnivores are related to quantifiable variables indexing the intensity of exploitative competition. Group size is predicted to increase with both food density and travel capabilities, as estimated by the constraint-free day-range (DR_s), i.e. the day-range of a solitary individual uninfluenced by competition from conspecifics. We test this exploitation competition hypothesis among primates and carnivores, using data on populations, species and genera. Food density was indexed by population density. Where DR_s could be estimated by regression it was found to be correlated with observed day-range (DR_m). DR_s was therefore indexed by DR_m in all species. Population density and DR_m were associated with each other, but in a multiple regression each variable contributed independently to explaining variation in group size. PGS was predicted better than FGS, but regressions involving either measure of group size were significant in all analyses. The multiple regression analyses were validated by the method of linear contrasts, which accounts for possible lack of independence among taxa. We conclude that species differences in group size are influenced by both food density and DR_s. This suggests that variation in the intensity of exploitation competition is partly responsible for differences in group size.Correspondence to: R.W. Wrangham     

Use of signature whistles during separations and reunions by wild bottlenose dolphin mothers and infants

We examine the contexts and patterns of signature whistle production by wild bottlenose dolphin mother-infant pairs (Tursiops spp.) to gain insight into the functional significance of whistles. Results are based on focal observations and simultaneous recordings of underwater vocalizations. Whistles occur primarily when mother-infant pairs are separated, and the probability of whistles increases with distance of separation. The timing of whistles during separations varies, but whistles tend to be produced in repetitive series and are generally concentrated toward the later stages of the separation, i.e., during the process of reunion. Although we focused on infants, mothers do not appear to whistle during separations as frequently as infants. Infant whistles may function to facilitate reunions by conveying information to the mother concerning the infant's motivation to reunite and/or its location. Infant whistles could induce a cooperative response from the mother including approach, slowing to allow the infant to catch up or whistling. Highly individualized signature whistles may be particularly useful in a fission-fusion society in which individuals (mothers and infants as well as adults) join and leave temporary parties in a fluid manner, yet maintain consistent, long-term associations with particular individuals. Correspondence to: R.A. Smolker     

Locomotory behaviour and swimming performance of the Norway lobster,Nephrops norvegicus,in the presence of an acoustic tag

Observations have been made on the locomotory behaviour and swimming performances of the Norway lobster, Nephrops norvegicus (L.), fitted with an acoustic transmitter ventrally under the cephalothorax. The walking behaviour of adult males (44 mm carapace length) appeared to be unaffected, but the tag caused significant reductions in certain measures of tail-flip swimming performance such as swimming speed and endurance. Flume-tank experiments in low water currents suggested that the transmitter would increase hydrodynamic drag during swimming by 9 to 32%, depending on lobster size. Given the weight and dimensions of the acoustic transmitters currently available, it is considered advisable to confine acoustic tracking studies to relatively large N. norvegicus.Correspondence to: C. J. Chapman     

Famine forecasting; Prices and peasant behaviour in Northern Ethiopia

This paper, completed in January 1984, begins by briefly outlining the contrasting views of price behaviour during the last Ethiopian famine (1972–1974) put forward by Seaman and Holt (1980) and Sen (1981). A hypothesis is developed to account for peasant behaviour and price responses under developing famine conditions. This is then tested with data recently made available. The paper concludes by summarizing the findings and their implications for famine forecasting, and argues that famine conditions in Northern Ethiopia are likely to worsen during 1984.     

Spontaneous resettlement after drought: An Ethiopian example*

Based upon two recent periods of fieldwork among a spontaneously resettled population of cattle herders in southwestern Ethiopia, this paper is an attempt to illustrate, by a particular case, the general proposition that external aid, if it is not to be counter-productive, must meet people's needs as they themselves define them. The people in question, the Mursi, have experienced, since 1970, their worst period of drought and hunger in living memory. They have responded to this experience with resilience and inventiveness and largely without external assistance. The paper concentrates on a recent, drought induced, migration of Mursi to higher land, outside their traditional territory, and shows how the migrants are adapting to a new way of life which brings them into direct conflict with some of their most cherished cultural traditions.     

A simple economic model of endangered species preservation in the United States

Concern for the acceleration in threats to species diversity in the United States led to the passage of the Endangered Species Act of 1973. In this paper, this legislation is examined in an economic context. A simple model is developed where management and constraints on economic activity are substitutes in the production of species stocks. Whether economic growth constraints (the primary policy tool of the 1973 Act) are justified for preservation purposes depends on technical substitution possibilities, relative cost ratios, and the existence of budget constraints for species management. Since the same combination of these factors will not occur for all species, it is argued that more flexibility in endangered species policy may preserve the same levels of species stocks at lower costs to society.     

POLITICAL ECONOMICS OF OFFSHORE OIL

The exploration of offshore areas for hydrocarbons has involved new technical and economic considerations and has posed new problems, not the least of which has been the changing relationship between companies and governments. In this article, the authors present an analysis of the different perspectives and policies which emanate from national governments on the one hand and from transnational oil companies on the other. The issues examined include the considerations involved in establishing exploration legislation, post-discovery problems, the impact on revenues of increased production, the interests of companies in developing fields and the overall economic environment within which decisions have to be agreed.