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961.
Phylogenetic analyses of the left domain of the mitochondrial DNA control-region sequence have been used to examine the relationships
among species of the genus Merluccius (Rafinesque, 1810), and to compare these with hypotheses based on morphological, meristic and allozyme characters. Analysis
of aligned sequences revealed that transition bias was much lower than in mammalian mtDNA, and that nucleotide composition
of control-region sequences was biased toward A and T. We have roughly calibrated a molecular clock for the genus, based on
the rise of the Isthmus of Panamá, which is believed to have created a barrier to dispersal between marine species of the
Atlantic and Pacific Oceans. Our mtDNA-based phylogeny was highly congruent with allozyme-based phylogenies, but poorly so
with a previously described phylogeny based on morphology. Specifically, our phylogeny resolved two well-supported principal
clades, one of American (west Atlantic and east Pacific) species and the other of Euro–African (east Atlantic) species. This
suggests an evolutionary history during which the ancestral lineage of Merluccius was divided between two geographic regions, with subsequent dispersal and vicariant events resulting in the evolution and
distribution of extant taxa. However, the relationships between some taxa within the American clade could not be resolved.
We suggest that this is consistent with an hypothesis of a rapid origin and radiation of these taxa.
Received: 12 December 1998 / Accepted: 15 October 1999 相似文献
962.
The pathway for the flow of salt-marsh grass production into marsh food-webs is still not well defined. We compared the abilities
of three marsh macroinvertebrates [salt marsh periwinkles, Littoraria irrorata (Say) (=Littorina irrorata), salt-marsh coffee-bean snails, Melampus bidentatus (Say); and a talitrid amphipod, Uhlorchestia spartinophila Bounsfield and Heard] to access standing-dead leaves of smooth cordgrass (Spartina alterniflora Loisel). The invertebrates were incubated with naturally-decaying leaves, and the rates of removal of organic matter and
living fungal biomass (ergosterol) were measured. The impact of invertebrate activity upon fungal growth rates was measured
as rates of fungal-membrane synthesis (incorporation of radioacetate into ergosterol). The removal rates of organic leaf biomass
per mg individual biomass were highest for amphipods (700 μg mg−1 d−1) and lowest for periwinkles (90 μg mg−1 d−1), but the relatively large biomass of the snails made their removal rates per individual greater than those of amphipods.
Net removal of ergosterol by all three invertebrates was >50% for yellow-brown (early-decay) leaf blades. For fully-brown
(advanced-decay) blades, >50% removal of ergosterol was found only for periwinkles; exposure to coffee-bean snails and amphipods
resulted in a net ergosterol reduction of ≤20%. The lower net reduction of living fungal biomass by coffee-bean snails and
amphipods may have been due to fungal-growth stimulation (2.3-fold stimulation in coffee-bean snails and 1.5-fold stimulation
in amphipods). Grazing by periwinkles did not stimulate fungal growth, possibly because of its high intensity. Grazing by
these three salt-marsh shredders may affect marsh-grass shoot-decay in different ways. Periwinkles may abbreviate the period
of fungal production, and incorporate the decaying material relatively quickly into snail biomass and fecal-pellet rain to
the sediments. Coffee-bean snails and amphipods may enhance and prolong fungal production, along with the formation of fecal-pellet
rain. All three invertebrates fed preferentially on leaf blades rather than leaf sheaths, and feeding rates of gastropods
were higher during the night than during the day.
Received: 25 November 1998 / Accepted: 4 November 1999 相似文献
963.
Cadmium accumulation in the female shore crab Carcinus maenas during the moult cycle and ovarian maturation 总被引:2,自引:0,他引:2
The influence of moulting and ovarian maturation on cadmium accumulation in the tissues of female shore crabs Carcinus maenas exposed to 1 mg Cd l−1 in the water was investigated. Cadmium accumulation in all tissues examined was markedly increased in crabs in the postmoult
stages (A and B) compared to crabs in all other moult stages. During the moult cycle, average cadmium accumulation in the
midgut gland ranged from 29 μg Cd g−1 dw at premoult stage (D2) to 589 μg Cd g−1 dw at postmoult stage (A). Average cadmium concentrations in the haemolymph ranged from 0.56 μg Cd ml−1 at intermoult stage (C4) to 4.6 μg Cd ml−1 at postmoult stage (A), while the gills accumulated from 103 μg Cd g−1 dw in intermoult stage (C3) to 352 μg Cd g−1 dw in postmoult stage (A). Cadmium concentration in gills and haemolymph was also significantly higher in crabs in late premoult
stage (D3) compared to C4-crabs, while midgut gland cadmium concentration remained elevated in C1- and C3- intermoult stages relative to C4. During ovarian maturation the cadmium accumulation in midgut gland, gills, ovaries and haemolymph decreased. Average cadmium
concentration in the midgut gland decreased from 63 μg g−1 dw in ovarian Stage I to 19 μg g−1 dw in ovarian Stage VI. The same pattern was observed for gills, haemolymph and ovaries. The present study demonstrates that
cadmium accumulation in the female shore crab strongly depends on the physiological status of the animal. A possible association
between physiological calcium requirements and cadmium accumulation during moulting is discussed.
Received: 20 January 2000 / Accepted: 20 July 2000 相似文献
964.
Efficient environmental regulation requires empirical estimates of the benefits and costs of environmental standards and policies. In the last 25 years, economists have made considerable progress in developing methods for measuring environmental benefits and in providing empirical estimates of costs and benefits. This paper reviews this progress. It examines by category of benefit the advancements made in providing estimates that can be used in benefit-cost analyses and identifies areas where additional research is required. 相似文献
965.
The effects of variable food supply on growth rate and feeding behaviour in two coexisting intertidal gastropods were examined
through field and laboratory experiments involving mark and recapture procedures, growth experiments under controlled food
supply and video observations of feeding. All cohorts of both Japeuthria species grew most in summer, with the highest growth rate being achieved by the youngest cohort of both sexes. In contrast,
almost all cohorts (except the youngest one) showed zero to negative growth in winter. The growth rates of Japeuthria species clearly varied depending on feeding frequencies and the initial size of individuals. Females of both species had
a significantly higher growth rate than males, though the difference became less marked with decreasing food supply. The total
time spent on feeding and the length of the first feeding bout generally increased as the duration of the interval between
meals was increased across treatments. From both the field growth experiment and the observations of feeding behaviour it
is clear that J. ferrea had a higher frequency of feeding than J. cingulata in the field; 1- to 3-d intervals of feeding frequency in the former and 3- to 6-d intervals in the latter. Most notably,
the two species showed a reversal in growth rates with changing conditions of food supply. When food was relatively abundant
J. ferrea achieved better growth than J. cingulata, but J. cingulata grew better than J. ferrea under low food supply. In view of the fact that J. ferrea seems to be competitively superior to J. cingulata under normal levels, the present observation that J. cingulata can perform better than J. ferrea when food availability is low gives an important advantage to the latter in terms of its coexistence with the former.
Received: 9 March 1999 / Accepted: 8 October 1999 相似文献
966.
Effect of temperature on statolith growth of the European squid Loligo vulgaris during early life 总被引:1,自引:0,他引:1
R. Villanueva 《Marine Biology》2000,136(3):449-460
Over the past decade, statolith interpretation has resulted in a major advance in our knowledge of squid population-dynamics,
but the way in which environmental conditions affect the statolith increment-deposition ratio remains virtually unknown. The
object of the present study was to determine the effect of temperature on this process, using tetracycline marks to validate
statolith growth in Loligo vulgaris Lamarck, 1798 under rearing conditions equivalent to severe winter (11 °C) and summer (19 °C) temperature regimes. Tetracycline
marking was performed every 10 d (at 10, 20, 30, 40, 50 and 60 d of age). The newly hatched squid paralarvae were slightly
smaller in summer than those hatched in winter. Survival rates were similar in both cultures, but growth rates (wet mass)
of summer squids were double those in winter. At hatching, statoliths were already longer in the summer squids, and growth
rates were 2% d−1 as opposed to 0.9% d−1 for winter statoliths. For the dorsal dome area of the statolith, where more increment counts were made, statolith growth
was of 3.25 μm d−1 in summer, and daily increment deposition was confirmed in 87% of the statoliths. The slow growth of statoliths at winter
temperatures yielded a mean growth of 1.1 μm d−1– insufficient to discern the increments using light microscopy. Subsequent SEM observation enabled only 21% of the winter
statoliths to be read; these also indicated a deposition rate of one increment d−1. Since the life span of L. vulgaris is ≃1 yr, squids will experience at least one winter during their life cycle, and this might be visible on the statolith.
Received: 28 June 1999 / Accepted: 20 December 1999 相似文献
967.
Arm damage is a widely reported but superficially investigated aspect of the biology of the starfish Asterias rubens L. In the present study, the incidence of arm damage was surveyed in populations of A.
rubens at two sites in the Firth of Clyde, Scotland, and three sites in Gullmarsfjorden, Sweden. The mean (±SD) incidence across
all sites of individuals with basal arm damage (resulting from detachment at the basal autotomy plane) was 19.69 ± 8.86%,
the incidence of those with distal arm damage (resulting from amputation at more distal levels) was 7.74 ± 10.01%. The mean
incidence of arms with basal damage was 5.28 ± 4.12%, of those with distal damage 1.83 ± 2.45%. There was a significant negative
correlation between size and the incidence of basal damage at all but one site, but no significant correlation between size
and distal damage at any site. Mechanical tests on specimens of the aboral body wall from the basal region of the arm (which
included the autotomy plane) and from a more distal region revealed that with increasing body size there was a significant
increase in yield stress, ultimate stress and Young's modulus (stiffness) but no significant change in yield strain and ultimate
strain. There was no significant difference between the relationships for basal and distal specimens. It is hypothesised that
in larger individuals increased mechanical toughness replaces autotomy as an effective antipredator strategy. Using two methods
to induce autotomy, a significant positive correlation between size and the delay between the onset of stimulation and arm
detachment was found; this may represent a size-related decline in the efficiency of the autotomy mechanism through the relaxation
of selection pressure. Since size is an unreliable indicator of age in A.
rubens, the trends identified herein can be interpreted only tentatively as age-associated phenomena.
Received: 4 September 1999 / Accepted: 17 February 2000 相似文献
968.
Genetic variation in 15 Holothuria (Microthele) nobilis (Selenka, 1867) populations on the Great Barrier Reef was studied at seven polymorphic allozyme loci. Although populations
were separated by distances up to 1300 km, there were no apparent restrictions to gene flow (F
ST
values were not significantly different from 0) and the maximum Nei's unbiased genetic distance was 0.003. Populations were
in Hardy–Weinberg equilibrium at all loci. The estimated maximum sexual input [this is the ratio of the number of sexually
produced individuals (N*) to the sample size (N
i
)] and the minimum sexual input [this is the ratio of the number of genotypes (N
go
) over the sample size (N
i
)] were used as estimators for the amount of asexual reproduction. Both parameters suggested that H. nobilis reproduces solely by sexual means (N*:N
i
: = 1; N
go
:N
i
= 0.74 to 1). The allozyme data indicated high gene flow between populations, but the possibility that allozyme frequencies
may not be at equilibrium means that it was not possible to distinguish whether the patterns reflect present-day dispersal
or dispersal that occurred in the past.
Received: 27 March 2000 / Accepted: 29 June 2000 相似文献
969.
The nature of heat coma was examined in the edible periwinkle Littorina littorea (L.). Duration of acclimation did not influence heat-coma temperature at 12 °C, although other acclimation temperatures were
important in influencing thermal tolerance, with positive shifts in coma temperature observed in response to elevated temperatures.
Previous thermal history also influenced heat-coma temperatures. Individuals subjected to repeat heat-coma events on a daily
basis showed significant declines (P < 0.05) in coma-temperature; in contrast individuals exposed to repeat heat-coma events on a weekly basis showed no decline
in thermal tolerance. Size-effects occurred at selected sites, where decreased heat-coma temperatures were recorded in large
individuals.
Received: 16 August 1999 / Accepted: 13 June 2000 相似文献
970.
Acoustic telemetry was used to examine patterns of activity and space utilisation of coelacanths, nocturnal predators which
spend the day in submarine caves. Nine coelacanths (Latimeria chalumnae) were tracked, each for a period of 1 to 16 nights at Grande Comore, West Indian Ocean. Activities lasted on average 9 h,
usually starting shortly after sunset and ending before sunrise. Vertically, coelacanths moved up and down at and below cave
level by following the bottom contour, mainly between 180 and 400 m depth. The deepest record was 698 m, the shallowest 133 m.
Most time was spent between 200 and 300 m depth. Large individuals performed deep excursions to depths below 400 m, usually
once per night. The fish spent most time in water temperatures of 15 to 19 °C; they rarely ventured into waters warmer than
22 °C measured at depths shallower than 160 m depth. Horizontally, coelacanths stayed in narrow areas ranging from <1 to 10 km
of coastline. Coelacanths are extremely slow drift-hunters with an estimated average swimming speed of 3.2 m min−1, often travelling not more than 3 km per night. They probably take advantage of local upwelling and downwelling and slow
currents occurring parallel to the steep slopes. This study shows that coelacanths are inhabitants of the subphotic zone,
where they are active mainly below the depth of their daytime refuges.
Received: 7 July 1999 / Accepted: 11 February 2000 相似文献