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271.
272.
The Chilean intertidal Lottiidae limpet fauna consists of at least nine species, grouped into the monophyletic taxon Scurria and a single taxon preliminarily designated as `Lottia' orbignyi (Dall). Within the Scurria clade, the most basal species are Scurria variabilis (Sowerby), S. viridula (Lamarck), and S.zebrina (Lesson), and among the most crown groups are S. ceciliana (Orbigny) and S. araucana (Orbigny). This phylogenetic information, obtained by the analysis of molecular characters (16S mtDNA), provides a framework
for a comparative study of behavioral characters. In this study, we analyze the escape responses of two basal limpet species
(`L.' orbignyi and S. viridula) and two derived limpet species (S. araucana and S. ceciliana) in the presence of the limpet predatory seastar Heliaster helianthus (Lamarck) and the limpet non-predatory seastars Stichaster striatus Muller and Troschel and Patiria chilensis Lutken. Neither P.chilensis nor Stichaster striatus induced escape responses such as those observed with the predatory H. helianthus. Moreover, in the presence of H. helianthus, basal and derived limpet species differed significantly in the percentage of individuals responding, reaction time, and
duration of the response. The basal species `L.' orbignyi and Scurriaviridula exhibited instantaneous and vigorous locomotor responses, whereas the derived species S. araucana and S. ceciliana displayed no locomotor responses. The same pattern was found in responses to H. helianthus homogenates. The results strongly suggest a correlation between phylogeny and anti-predator escape responses to the seastar
H. helianthus. Alternative escape responses have evolved for basal and derived species within the Chilean clade.
Received: 27 April 2000 / Accepted: 16 August 2000 相似文献
273.
Population structure and reproductive ecology of the burrowing ghost shrimp Callianassa filholi Milne-Edwards, 1878 were studied in populations along a latitudinal gradient throughout New Zealand during the breeding season.
Size-frequency distributions revealed unimodal populations, with predominantly sexually mature shrimp. All populations showed
an unbiased sex-ratio, and there appeared to be no significant difference in size (carapace length, CL) between sexes. At
the same time, CL and size at maturity differed significantly between populations; however, a general increase in sizes from
north to south was not consistent throughout the latitudinal range studied. The timing of the breeding cycle differed significantly
between populations, and breeding started earlier in southern populations. Number of embryos (fecundity) increased linearly
with female CL at each location, but rates differed significantly between populations. Embryo size was not related to number
of embryos, and the former increased significantly with latitude. With the exception of embryo size, observed differences
in body size/size at maturity and reproductive timing between C. filholi populations are thought to be determined by food availability rather than temperature. Thus, further study is suggested on
these aspects of thalassinid reproductive biology.
Received: 17 June 1999 / Accepted: 8 January 2000 相似文献
274.
We examined the mating behaviour of the New Zealand ocypodid crab Macrophthalmus hirtipes in the laboratory between February and June 1998. This species has a discrete breeding season. Mating and moulting were not
linked and only intermoult females with mobile gonopore opercula were attractive to males. Allometry and compatibility of
gonopods and gonopores of different-sized crabs was investigated. Under laboratory conditions, the opercula of intermoult
females remained mobile on average for 11.4 d, but the duration of receptivity did not appear to be under female control.
The operational sex ratio in the laboratory fluctuated greatly, but was always male-dominated. During the period of opercular
mobility, females mated many times with several different males. Matings in the absence of burrows were relatively short (mean
duration = 23 min, max. = 122 min) and the mating behaviour of M. hirtipes lacked courtship and mate-guarding. Males used a search-intercept method to acquire mates, with very low levels of intrasexual
competition. There was no evidence of mate preference in M. hirtipes, and males spent just as long mating with ovigerous females as with non-ovigerous ones. Although M. hirtipes has ventral-type spermathecae, as do several other ocypodid crabs, it is unclear whether this promotes last-male sperm precedence.
The role of burrows in modifying the mating behaviour of M. hirtipes in the field remains to be established.
Received: 7 January 2000 / Accepted: 5 June 2000 相似文献
275.
Phylogenetic analyses of the left domain of the mitochondrial DNA control-region sequence have been used to examine the relationships
among species of the genus Merluccius (Rafinesque, 1810), and to compare these with hypotheses based on morphological, meristic and allozyme characters. Analysis
of aligned sequences revealed that transition bias was much lower than in mammalian mtDNA, and that nucleotide composition
of control-region sequences was biased toward A and T. We have roughly calibrated a molecular clock for the genus, based on
the rise of the Isthmus of Panamá, which is believed to have created a barrier to dispersal between marine species of the
Atlantic and Pacific Oceans. Our mtDNA-based phylogeny was highly congruent with allozyme-based phylogenies, but poorly so
with a previously described phylogeny based on morphology. Specifically, our phylogeny resolved two well-supported principal
clades, one of American (west Atlantic and east Pacific) species and the other of Euro–African (east Atlantic) species. This
suggests an evolutionary history during which the ancestral lineage of Merluccius was divided between two geographic regions, with subsequent dispersal and vicariant events resulting in the evolution and
distribution of extant taxa. However, the relationships between some taxa within the American clade could not be resolved.
We suggest that this is consistent with an hypothesis of a rapid origin and radiation of these taxa.
Received: 12 December 1998 / Accepted: 15 October 1999 相似文献
276.
The pathway for the flow of salt-marsh grass production into marsh food-webs is still not well defined. We compared the abilities
of three marsh macroinvertebrates [salt marsh periwinkles, Littoraria irrorata (Say) (=Littorina irrorata), salt-marsh coffee-bean snails, Melampus bidentatus (Say); and a talitrid amphipod, Uhlorchestia spartinophila Bounsfield and Heard] to access standing-dead leaves of smooth cordgrass (Spartina alterniflora Loisel). The invertebrates were incubated with naturally-decaying leaves, and the rates of removal of organic matter and
living fungal biomass (ergosterol) were measured. The impact of invertebrate activity upon fungal growth rates was measured
as rates of fungal-membrane synthesis (incorporation of radioacetate into ergosterol). The removal rates of organic leaf biomass
per mg individual biomass were highest for amphipods (700 μg mg−1 d−1) and lowest for periwinkles (90 μg mg−1 d−1), but the relatively large biomass of the snails made their removal rates per individual greater than those of amphipods.
Net removal of ergosterol by all three invertebrates was >50% for yellow-brown (early-decay) leaf blades. For fully-brown
(advanced-decay) blades, >50% removal of ergosterol was found only for periwinkles; exposure to coffee-bean snails and amphipods
resulted in a net ergosterol reduction of ≤20%. The lower net reduction of living fungal biomass by coffee-bean snails and
amphipods may have been due to fungal-growth stimulation (2.3-fold stimulation in coffee-bean snails and 1.5-fold stimulation
in amphipods). Grazing by periwinkles did not stimulate fungal growth, possibly because of its high intensity. Grazing by
these three salt-marsh shredders may affect marsh-grass shoot-decay in different ways. Periwinkles may abbreviate the period
of fungal production, and incorporate the decaying material relatively quickly into snail biomass and fecal-pellet rain to
the sediments. Coffee-bean snails and amphipods may enhance and prolong fungal production, along with the formation of fecal-pellet
rain. All three invertebrates fed preferentially on leaf blades rather than leaf sheaths, and feeding rates of gastropods
were higher during the night than during the day.
Received: 25 November 1998 / Accepted: 4 November 1999 相似文献
277.
Arm damage is a widely reported but superficially investigated aspect of the biology of the starfish Asterias rubens L. In the present study, the incidence of arm damage was surveyed in populations of A.
rubens at two sites in the Firth of Clyde, Scotland, and three sites in Gullmarsfjorden, Sweden. The mean (±SD) incidence across
all sites of individuals with basal arm damage (resulting from detachment at the basal autotomy plane) was 19.69 ± 8.86%,
the incidence of those with distal arm damage (resulting from amputation at more distal levels) was 7.74 ± 10.01%. The mean
incidence of arms with basal damage was 5.28 ± 4.12%, of those with distal damage 1.83 ± 2.45%. There was a significant negative
correlation between size and the incidence of basal damage at all but one site, but no significant correlation between size
and distal damage at any site. Mechanical tests on specimens of the aboral body wall from the basal region of the arm (which
included the autotomy plane) and from a more distal region revealed that with increasing body size there was a significant
increase in yield stress, ultimate stress and Young's modulus (stiffness) but no significant change in yield strain and ultimate
strain. There was no significant difference between the relationships for basal and distal specimens. It is hypothesised that
in larger individuals increased mechanical toughness replaces autotomy as an effective antipredator strategy. Using two methods
to induce autotomy, a significant positive correlation between size and the delay between the onset of stimulation and arm
detachment was found; this may represent a size-related decline in the efficiency of the autotomy mechanism through the relaxation
of selection pressure. Since size is an unreliable indicator of age in A.
rubens, the trends identified herein can be interpreted only tentatively as age-associated phenomena.
Received: 4 September 1999 / Accepted: 17 February 2000 相似文献
278.
Genetic variation in 15 Holothuria (Microthele) nobilis (Selenka, 1867) populations on the Great Barrier Reef was studied at seven polymorphic allozyme loci. Although populations
were separated by distances up to 1300 km, there were no apparent restrictions to gene flow (F
ST
values were not significantly different from 0) and the maximum Nei's unbiased genetic distance was 0.003. Populations were
in Hardy–Weinberg equilibrium at all loci. The estimated maximum sexual input [this is the ratio of the number of sexually
produced individuals (N*) to the sample size (N
i
)] and the minimum sexual input [this is the ratio of the number of genotypes (N
go
) over the sample size (N
i
)] were used as estimators for the amount of asexual reproduction. Both parameters suggested that H. nobilis reproduces solely by sexual means (N*:N
i
: = 1; N
go
:N
i
= 0.74 to 1). The allozyme data indicated high gene flow between populations, but the possibility that allozyme frequencies
may not be at equilibrium means that it was not possible to distinguish whether the patterns reflect present-day dispersal
or dispersal that occurred in the past.
Received: 27 March 2000 / Accepted: 29 June 2000 相似文献
279.
The nature of heat coma was examined in the edible periwinkle Littorina littorea (L.). Duration of acclimation did not influence heat-coma temperature at 12 °C, although other acclimation temperatures were
important in influencing thermal tolerance, with positive shifts in coma temperature observed in response to elevated temperatures.
Previous thermal history also influenced heat-coma temperatures. Individuals subjected to repeat heat-coma events on a daily
basis showed significant declines (P < 0.05) in coma-temperature; in contrast individuals exposed to repeat heat-coma events on a weekly basis showed no decline
in thermal tolerance. Size-effects occurred at selected sites, where decreased heat-coma temperatures were recorded in large
individuals.
Received: 16 August 1999 / Accepted: 13 June 2000 相似文献
280.
Acoustic telemetry was used to examine patterns of activity and space utilisation of coelacanths, nocturnal predators which
spend the day in submarine caves. Nine coelacanths (Latimeria chalumnae) were tracked, each for a period of 1 to 16 nights at Grande Comore, West Indian Ocean. Activities lasted on average 9 h,
usually starting shortly after sunset and ending before sunrise. Vertically, coelacanths moved up and down at and below cave
level by following the bottom contour, mainly between 180 and 400 m depth. The deepest record was 698 m, the shallowest 133 m.
Most time was spent between 200 and 300 m depth. Large individuals performed deep excursions to depths below 400 m, usually
once per night. The fish spent most time in water temperatures of 15 to 19 °C; they rarely ventured into waters warmer than
22 °C measured at depths shallower than 160 m depth. Horizontally, coelacanths stayed in narrow areas ranging from <1 to 10 km
of coastline. Coelacanths are extremely slow drift-hunters with an estimated average swimming speed of 3.2 m min−1, often travelling not more than 3 km per night. They probably take advantage of local upwelling and downwelling and slow
currents occurring parallel to the steep slopes. This study shows that coelacanths are inhabitants of the subphotic zone,
where they are active mainly below the depth of their daytime refuges.
Received: 7 July 1999 / Accepted: 11 February 2000 相似文献