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621.
Efficient environmental regulation requires empirical estimates of the benefits and costs of environmental standards and policies. In the last 25 years, economists have made considerable progress in developing methods for measuring environmental benefits and in providing empirical estimates of costs and benefits. This paper reviews this progress. It examines by category of benefit the advancements made in providing estimates that can be used in benefit-cost analyses and identifies areas where additional research is required. 相似文献
622.
The effects of variable food supply on growth rate and feeding behaviour in two coexisting intertidal gastropods were examined
through field and laboratory experiments involving mark and recapture procedures, growth experiments under controlled food
supply and video observations of feeding. All cohorts of both Japeuthria species grew most in summer, with the highest growth rate being achieved by the youngest cohort of both sexes. In contrast,
almost all cohorts (except the youngest one) showed zero to negative growth in winter. The growth rates of Japeuthria species clearly varied depending on feeding frequencies and the initial size of individuals. Females of both species had
a significantly higher growth rate than males, though the difference became less marked with decreasing food supply. The total
time spent on feeding and the length of the first feeding bout generally increased as the duration of the interval between
meals was increased across treatments. From both the field growth experiment and the observations of feeding behaviour it
is clear that J. ferrea had a higher frequency of feeding than J. cingulata in the field; 1- to 3-d intervals of feeding frequency in the former and 3- to 6-d intervals in the latter. Most notably,
the two species showed a reversal in growth rates with changing conditions of food supply. When food was relatively abundant
J. ferrea achieved better growth than J. cingulata, but J. cingulata grew better than J. ferrea under low food supply. In view of the fact that J. ferrea seems to be competitively superior to J. cingulata under normal levels, the present observation that J. cingulata can perform better than J. ferrea when food availability is low gives an important advantage to the latter in terms of its coexistence with the former.
Received: 9 March 1999 / Accepted: 8 October 1999 相似文献
623.
Effect of temperature on statolith growth of the European squid Loligo vulgaris during early life 总被引:1,自引:0,他引:1
R. Villanueva 《Marine Biology》2000,136(3):449-460
Over the past decade, statolith interpretation has resulted in a major advance in our knowledge of squid population-dynamics,
but the way in which environmental conditions affect the statolith increment-deposition ratio remains virtually unknown. The
object of the present study was to determine the effect of temperature on this process, using tetracycline marks to validate
statolith growth in Loligo vulgaris Lamarck, 1798 under rearing conditions equivalent to severe winter (11 °C) and summer (19 °C) temperature regimes. Tetracycline
marking was performed every 10 d (at 10, 20, 30, 40, 50 and 60 d of age). The newly hatched squid paralarvae were slightly
smaller in summer than those hatched in winter. Survival rates were similar in both cultures, but growth rates (wet mass)
of summer squids were double those in winter. At hatching, statoliths were already longer in the summer squids, and growth
rates were 2% d−1 as opposed to 0.9% d−1 for winter statoliths. For the dorsal dome area of the statolith, where more increment counts were made, statolith growth
was of 3.25 μm d−1 in summer, and daily increment deposition was confirmed in 87% of the statoliths. The slow growth of statoliths at winter
temperatures yielded a mean growth of 1.1 μm d−1– insufficient to discern the increments using light microscopy. Subsequent SEM observation enabled only 21% of the winter
statoliths to be read; these also indicated a deposition rate of one increment d−1. Since the life span of L. vulgaris is ≃1 yr, squids will experience at least one winter during their life cycle, and this might be visible on the statolith.
Received: 28 June 1999 / Accepted: 20 December 1999 相似文献
624.
Arm damage is a widely reported but superficially investigated aspect of the biology of the starfish Asterias rubens L. In the present study, the incidence of arm damage was surveyed in populations of A.
rubens at two sites in the Firth of Clyde, Scotland, and three sites in Gullmarsfjorden, Sweden. The mean (±SD) incidence across
all sites of individuals with basal arm damage (resulting from detachment at the basal autotomy plane) was 19.69 ± 8.86%,
the incidence of those with distal arm damage (resulting from amputation at more distal levels) was 7.74 ± 10.01%. The mean
incidence of arms with basal damage was 5.28 ± 4.12%, of those with distal damage 1.83 ± 2.45%. There was a significant negative
correlation between size and the incidence of basal damage at all but one site, but no significant correlation between size
and distal damage at any site. Mechanical tests on specimens of the aboral body wall from the basal region of the arm (which
included the autotomy plane) and from a more distal region revealed that with increasing body size there was a significant
increase in yield stress, ultimate stress and Young's modulus (stiffness) but no significant change in yield strain and ultimate
strain. There was no significant difference between the relationships for basal and distal specimens. It is hypothesised that
in larger individuals increased mechanical toughness replaces autotomy as an effective antipredator strategy. Using two methods
to induce autotomy, a significant positive correlation between size and the delay between the onset of stimulation and arm
detachment was found; this may represent a size-related decline in the efficiency of the autotomy mechanism through the relaxation
of selection pressure. Since size is an unreliable indicator of age in A.
rubens, the trends identified herein can be interpreted only tentatively as age-associated phenomena.
Received: 4 September 1999 / Accepted: 17 February 2000 相似文献
625.
Genetic variation in 15 Holothuria (Microthele) nobilis (Selenka, 1867) populations on the Great Barrier Reef was studied at seven polymorphic allozyme loci. Although populations
were separated by distances up to 1300 km, there were no apparent restrictions to gene flow (F
ST
values were not significantly different from 0) and the maximum Nei's unbiased genetic distance was 0.003. Populations were
in Hardy–Weinberg equilibrium at all loci. The estimated maximum sexual input [this is the ratio of the number of sexually
produced individuals (N*) to the sample size (N
i
)] and the minimum sexual input [this is the ratio of the number of genotypes (N
go
) over the sample size (N
i
)] were used as estimators for the amount of asexual reproduction. Both parameters suggested that H. nobilis reproduces solely by sexual means (N*:N
i
: = 1; N
go
:N
i
= 0.74 to 1). The allozyme data indicated high gene flow between populations, but the possibility that allozyme frequencies
may not be at equilibrium means that it was not possible to distinguish whether the patterns reflect present-day dispersal
or dispersal that occurred in the past.
Received: 27 March 2000 / Accepted: 29 June 2000 相似文献
626.
The nature of heat coma was examined in the edible periwinkle Littorina littorea (L.). Duration of acclimation did not influence heat-coma temperature at 12 °C, although other acclimation temperatures were
important in influencing thermal tolerance, with positive shifts in coma temperature observed in response to elevated temperatures.
Previous thermal history also influenced heat-coma temperatures. Individuals subjected to repeat heat-coma events on a daily
basis showed significant declines (P < 0.05) in coma-temperature; in contrast individuals exposed to repeat heat-coma events on a weekly basis showed no decline
in thermal tolerance. Size-effects occurred at selected sites, where decreased heat-coma temperatures were recorded in large
individuals.
Received: 16 August 1999 / Accepted: 13 June 2000 相似文献
627.
Acoustic telemetry was used to examine patterns of activity and space utilisation of coelacanths, nocturnal predators which
spend the day in submarine caves. Nine coelacanths (Latimeria chalumnae) were tracked, each for a period of 1 to 16 nights at Grande Comore, West Indian Ocean. Activities lasted on average 9 h,
usually starting shortly after sunset and ending before sunrise. Vertically, coelacanths moved up and down at and below cave
level by following the bottom contour, mainly between 180 and 400 m depth. The deepest record was 698 m, the shallowest 133 m.
Most time was spent between 200 and 300 m depth. Large individuals performed deep excursions to depths below 400 m, usually
once per night. The fish spent most time in water temperatures of 15 to 19 °C; they rarely ventured into waters warmer than
22 °C measured at depths shallower than 160 m depth. Horizontally, coelacanths stayed in narrow areas ranging from <1 to 10 km
of coastline. Coelacanths are extremely slow drift-hunters with an estimated average swimming speed of 3.2 m min−1, often travelling not more than 3 km per night. They probably take advantage of local upwelling and downwelling and slow
currents occurring parallel to the steep slopes. This study shows that coelacanths are inhabitants of the subphotic zone,
where they are active mainly below the depth of their daytime refuges.
Received: 7 July 1999 / Accepted: 11 February 2000 相似文献
628.
An experimental study on recolonization and succession of marine macrobenthos in defaunated sediment 总被引:3,自引:0,他引:3
Hypoxia/anoxia in coastal waters is a world wide problem which often results in mass mortality and defaunation of benthos.
In this study, field experiments were carried out to examine recolonization and succession of macrobenthic infauna in defaunated
sediments, and the time required for recovery from complete defaunation to a stable community. Trays (33 cm length × 25.5 cm
width × 11 cm depth) of defaunated sediment were exposed at the subtidal of a pristine site in subtropical Hong Kong. Temporal
changes of macrobenthic communities in defaunated sediment were analyzed by univariate and multivariate statistics, and compared
with those in undisturbed natural sediment at the same site. Initial colonization of macrobenthos occurred rapidly. A total
of 42 species was found, with an average of 258 animals per tray and 24 species per tray recorded in the first month. Abundance
showed a small peak (496 animals per tray) after 3 months, reached a sharp peak (1154 animals per tray) after 6 months, and
declined thereafter. Species number increased gradually, reached a maximum (68 species per tray) after 9 months, and then
decreased. Recolonization was predominantly contributed by larval settlement rather than adult migration. Temporal changes
in abundance, species number and diversity of the macrobenthic community in defaunated sediment resemble the spatial changes
along a decreasing pollution gradient previously defined by other authors. Results of this experiment suggest that newly available
sediment may allow more species to colonize (or coexist) than sediment pre-occupied by an established community. This is probably
due to less interspecific competition in the former habitat. No significant difference in abundance or species richness was
observed between defaunated and natural sediments after 15 months, suggesting that a stable community had been achieved, although
minor variations in species composition were still discernible between defaunated and natural sediments.
Received: 24 May 1999 / Accepted: 20 October 1999 相似文献
629.
Early development of the Antarctic sea urchin Sterechinus neumayeri was examined under two differ-ent culture regimes: one to simulate development near-bottom (“demersal development”) and the
other to simulate the development of embryos in the water column (“pelagic development”). When embryos of both treatments
reached the hatching blastula stage at 5 d post-fertilization (−1.5 °C), the blastulae that had undergone demersal development
evidenced significant differences (by ANOVA or suitable non-parametric comparison) in the following: a thicker blastoderm
layer (12%, P < 0.001), higher ash-free dry weights (19%, P < 0.01), lower mass-specific respiration rates (50%, P < 0.001), higher incorporation rates of 35S-methionine into protein (23%, P < 0.003), and a differential pattern of protein synthesis. When embryos developed demersally, they remained in the jelly-coat
material released with the eggs at spawning. Quantitative isolation of this jelly-coat material in S. neumayeri demonstrated that it contained a significant amount of organic matter, 115 ng ash-free dry mass per egg, equivalent to 17%
of the egg's initial organic mass. Uptake of external nutrients during embryogenesis may be a significant component of the
physiological energetics of this polar invertebrate by allowing the utilization of jelly-coat material released by a female
during spawning.
Received: 21 April 1999 / Accepted: 5 June 2000 相似文献
630.
Maternal investment in rhesus macaques (Macaca mulatta): reproductive costs and consequences of raising sons 总被引:1,自引:0,他引:1
Maternal investment in offspring is expected to vary according to offspring sex when the reproductive success of the progeny
is a function of differential levels of parental expenditure. We conducted a longitudinal investigation of rhesus macaques
to determine whether variation in male progeny production, measured with both DNA fingerprinting and short tandem repeat marker
typing, could be traced back to patterns of maternal investment. Males weigh significantly more than females at birth, despite
an absence of sex differences in gestation length. Size dimorphism increases during infancy, with maternal rank associated
with son’s, but not daughter’s, weight at the end of the period of maternal investment. Son’s, but not daughter’s, weight
at 1 year of age is significantly correlated with adult weight, and male, but not female, weight accounts for a portion of
the variance in reproductive success. Variance in annual offspring output was three- to fourfold higher in males than in females.
We suggest that energetic costs of rearing sons could be buffered by fetal delivery of testosterone to the mother, which is
aromatized to estrogen and fosters fat accumulation during gestation. We conclude that maternal investment is only slightly
greater in sons than in daughters, with mothers endowing sons with extra resources because son, but not daughter, mass has
ramifications for offspring sirehood. However, male reproductive tactics supersede maternal investment patterns as fundamental
regulators of male fitness.
Received: 23 July 1999 / Received in revised form: 23 February 2000 / Accepted: 13 March 2000 相似文献