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561.
Characteristics that influence male reproductive success on a lek of Lethrinops c.f. parvidens (Teleostei: Cichlidae) 总被引:1,自引:1,他引:0
Sexual selection has long been proposed as a mechanism leading to the diverse cichlid (Teleostei: Cichlidae) fauna of Lake
Malawi, Africa. Many of the shallow-water, sand-dwelling, bower-building cichlid species are particularly well suited for
studies of sexual selection because they participate in leks. Since females in lekking systems appear to acquire only genetic
material from their mates, it has been suggested that leks are ideal systems to study female mate choice. The objectives of
the investigation were to examine Lethrinops c.f. parvidens male bower characteristics (i.e., bower size and location) as well as other male characteristics (i.e., length, gular color,
and duration on the lek) for their influence on male mating success as measured by the number of visits, circles, and eggs
laid by females. These measures are nested in that a visit by a female may or may not lead to circling, and circling by a
female may or may not lead to egg-laying. We found increased bower height and higher numbers of conspecific neighbors (analogous
to shallow-water, near-shore bower positions) to be positively, significantly associated with the number of visits by females.
The only significant correlate with the number of circles was visits, and similarly circles was the only significant correlate
with the number of eggs laid. The R
2 value for the egg-laying regression was quite low (19.8%) compared with visits (54.3%) and circling (78.9%), suggesting that
females may be using additional cues, that we failed to measure, when in close proximity to males or simply that a small proportion
of the females were ready to spawn. Both indirect selection and direct selection pressure due to egg predation may have influenced
female choice on the lek.
Received: 10 April 1999 / Received in revised form: 26 July 1999 / Accepted: 18 September 1999 相似文献
562.
Experimental evidence of a testosterone-induced shift from paternal to mating behaviour in a facultatively polygynous songbird 总被引:5,自引:2,他引:3
Previous studies have suggested that testosterone (T) profiles of male birds reflect a trade-off between mate attraction behaviours
(requiring high T levels) and parental care activities (requiring low T levels). In this study, we experimentally elevated
T levels of monogamous males in the facultatively polygynous European starling (Sturnus vulgaris), and compared mate attraction and paternal behaviour of T-treated males with those of controls (C-males). T-males significantly
reduced their participation in incubation and fed nestlings significantly less often than C-males. Females paired to T-treated
males did not compensate for their mate’s lower paternal effort. The observed reduction in a male’s investment in incubating
the eggs was accompanied by an increased investment in typical female-attracting behaviours: T-males spent a significantly
higher proportion of their time singing to attract additional females. They also occupied more additional nestboxes than C-males,
although the differences just failed to be significant, and carried significantly more green nesting materials into an additional
nestbox (a behaviour previously shown to serve a courtship function). T-males also behaved significantly more aggressively
than C-males. During the nestling period, the frequency of mate-attracting behaviours by T-treated and control males no longer
differed significantly. Despite the reduced paternal effort by T-males and the lack of compensation behaviour by females,
hatching and breeding success did not differ significantly between T- and C-pairs.
Received: 7 February 2000 / Revised: 10 August 2000 / Accepted: 3 September 2000 相似文献
563.
Weather-mediated natural selection on arrival time in cliff swallows (Petrochelidon pyrrhonota) 总被引:2,自引:0,他引:2
An unusually long period of cold weather in May 1996 caused extensive mortality among insectivorous cliff swallows (Petrochelidon pyrrhonota) in the northern and central Great Plains. We analyzed how viability selection affected spring arrival time in a migratory
Nebraska population by comparing capture histories of survivors with those of birds known to have died and by documenting
how arrival time changed in the year following the selection event. Surviving birds had significantly later first-capture
dates (an index of arrival time) in the years prior to selection than those that died; a significant selection differential
suggested directional selection for birds that arrived later. Colony sites were occupied significantly later following the
selection event, and the distribution of first-capture dates in the season after selection was significantly shifted toward
later arrivals. Offspring of the survivors tended to arrive later than birds of the same age prior to the selection event.
While major weather-caused mortality events of this magnitude are rare in the study area, spells of cold weather severe enough
to cause limited mortality are frequent in April and early May. At least 25 probable mortality events of varying severity
were identified in the last 50 years based on climatological data. Periodic weather-mediated selection against early arrival
constrains the cliff swallow’s breeding season and may partly prevent directional selection for earlier nesting.
Received: 19 October 1999 / Received in revised form: 15 January 2000 / Accepted: 24 January 2000 相似文献
564.
Ectoparasites of house sparrows (Passer domesticus): an experimental test of the immunocompetence handicap hypothesis and a new model 总被引:2,自引:0,他引:2
Conspicuous secondary sexual traits may have evolved as handicap-revealing signals or as badges of status. We present results
of an experiment using males of the sexually dimorphic house sparrow (Passer domesticus), that support the idea that the male-specific bib can be both a handicap-revealing signal and a reliable badge indicating
the physical condition of the bird. In a test of the immunocompetence handicap hypothesis, wild-caught adult male house sparrows
were studied in captivity. Birds implanted with elevated doses of testosterone were more dominant, had higher circulating
levels of both testosterone and corticosterone and they also harboured relatively larger ectoparasite loads. Higher parasite
loads were also associated with individuals showing lower immunocompetence and larger changes in bib size. A new model for
immunocompetence effects in sexual selection is introduced, integrating actions that the hypothalamopituitary axis exerts
on gonads, adrenals and the thyroid gland. The ”integrated immunocompetence model” synthesizes both the ”handicap” (i.e. survival-decreasing)
and ”badge of status” (i.e. survival- enhancing) models for evolution of secondary sexual traits.
Received: 15 May 1999 / Received in revised form: 2 November 1999 / Accepted: 6 November 1999 相似文献
565.
Social and mating system of cooperatively breeding laughing kookaburras (Dacelo novaeguineae) 总被引:4,自引:0,他引:4
DNA fingerprinting was combined with field observations over four breeding seasons to investigate the social structure and
mating system of the laughing kookaburra (Dacelo novaeguineae). Groups comprised a socially dominant pair and up to six helpers of either sex. Helpers were always recruited from young
hatched in the group. Territorial inheritance, which is a feature of other cooperative breeders and an oft-cited benefit of
philopatry, did not occur. Helpers only attained dominant status in an established group by dispersing into a vacant dominant
position in that group. However, helpers could also form new groups by excising a new territory, often through a ”budding”
process. The mating system was overwhelmingly monogamous. There were no cases of extra-group parentage in a sample of 140
nestlings; within groups of three or more birds, dominance predicted parentage almost perfectly (99.2% of 129 nestlings),
irrespective of whether helpers in the group were related to one or both dominant birds. This is contrary to predictions from
models of reproductive skew, possibly because they currently fail to incorporate the willingness of females to share reproduction
among males.
Received: 15 May 1999 / Received in revised form: 2 November 1999 / Accepted: 6 November 1999 相似文献
566.
Aggression and resource sharing among foundresses in the social wasp Polistes dominulus: testing transactional theories of conflict 总被引:3,自引:0,他引:3
Tug-of-war models of within-group conflict predict that the frequency of conflict will be positively related to the degree
of reproductive sharing within the group; in contrast, a negative relationship supports transactional models, in which reproductive
payments among group members limit the degree of within-group selfishness. We tested predictions of the tug-of-war and transactional
models by examining cofoundress interactions during the founding (preworker) phase of colony development in 30 naturally nesting
colonies of the paper wasp Polistes dominulus. We found that the mean rate of foundress aggression and the mean probability of food sharing were significantly negatively
associated, which supports the prediction of the transactional, not the tug-of-war model. Further, cofoundress aggression
significantly increased over the founding phase (independently of temperature), while the fraction of aggression initiated
by the dominant (alpha) foundress significantly decreased over this period. We show that both of these results are predicted
by the transactional model of within-group conflict. Interestingly, the alpha’s rate of aggression was significantly positively
temperature dependent, while the beta’s was not. This indicates that the alpha’s aggression level may often be near her physiological
maximum, while the beta’s aggression is limi- ted by other factors, contradicting the prediction of the tug-of-war model.
Moreover, the alpha’s aggression was significantly positively temperature dependent only in the second half of the founding
period, as predicted by the transactional model since this is when there is least reproductive sharing. Finally, our results
indicate that the alpha’s level of aggression depends on the resources controlled by the beta.
Received: 18 January 2000 / Revised: 19 June 2000 / Accepted: 24 June 2000 相似文献
567.
Male willow warblers have song repertoires which vary in complexity along several dimensions. We examined whether female choice,
as measured by date of pairing, was based on these song characteristics in 4 different years. Pairing date was negatively
correlated with song repertoire size in 1 year, and with song versatility in another year, but there was no consistent effect
of any song characteristic on pairing throughout the years or in the pooled sample. The variable that best explained how soon
a male pairs is male arrival date (only males that had settled territories before the first female arrived were considered
in the analysis). This correlation is consistently significant in all years. This is most parsimoniously interpreted as females
choosing some habitat characteristic in the same way that males do. A small percentage of males (8.3%) attracted and paired
with a second female. The likelihood of becoming polygynous was not explained by any measured song characteristic, but it
was related to arrival date: early males were more likely to pair with two females. Males with large repertoires fledged more
young in their primary nests, and there was a trend for the offspring of these males to have a greater probability of being
recruited into the population. In conclusion, the results show that in most years there is no sexual selection by female preference
on song characteristics, although the data on reproductive success is consistent with the idea of repertoire size being an
indicator of male quality.
Received: 4 June 1999 / Received in revised form: 1 December 1999 / Accepted: 31 December 1999 相似文献
568.
In the meerkat (Suricata suricatta), a cooperative mongoose, pups follow potential feeders while the group is foraging and emit incessant calls when soliciting
food from them. In contrast to a ’stationary’ brood of chicks, in which nestlings are fed at a fixed location, meerkat pups
are ’mobile’ and become spread out. The question arises whether meerkat pups that experience different constraints to those
facing chicks have evolved similar begging strategies. This paper describes the vocalisations that meerkat pups emit in the
context of begging and investigates the influence of these calls on food allocation by older group members and on the behaviour
of littermates. Meerkat pups use two types of calls when soliciting food from a potential feeder. The most common is a ’repeat’
call, which pups emit continuously when following an older forager over several hours a day. In addition, when a potential
feeder finds a prey item, the pups next to it emit a bout of calls with increased calling rate, amplitude and fundamental
frequency, termed ’high-pitched’ calls. Observations, together with playback experiments, showed that more prey was allocated
to pups that called longer and more intensely. The pup closest to a feeder was almost always fed. The probability of emitting
high-pitched calls did not depend on the time since a pup had received food, and the change from repeat to high-pitched calls
occurred suddenly. The main function of the high-pitched call, therefore, does not appear to be to signal a pup’s hunger state.
More likely, the two calls, in the context of begging, may be an adaptation to energetic constraints in a mobile feeding system.
Pups, which are dispersed during foraging, may emit repeat calls over long periods to prevent potential feeders from eating
all the prey themselves. At the moment a potential feeder finds prey, pups may give the more intense high-pitched calls to
direct feeders to bring the food item to them and not to a littermate. Therefore, unlike the stationary feeding system where
chicks emit one type of begging call when the feeder approaches the nest, meerkats, with a mobile feeding system, have evolved
two discrete types of vocalisations in the context of begging.
Received: 22 November 1999 / Revised: 1 July 2000 / Accepted: 17 July 2000 相似文献
569.
T. Lubjuhn Wolfgang Winkel Jörg Thomas Epplen Jörg Brün 《Behavioral ecology and sociobiology》2000,48(1):12-17
One of the most interesting aspects of the mating system of pied flycatchers is the regular occurrence of polygyny. Here we
present data on the reproductive success of polygynously paired pied flycatcher males compared to monogamous males based on
paternity analyses through DNA fingerprinting. Males paired with two females suffered a higher loss in reproductive output
per female compared to monogamous males due to (1) a greater proportion of unhatched eggs in their broods, (2) greater nestling
mortality and (3) a greater probability of being cuckolded. Nevertheless, the number of fledglings was significantly greater
for polygynous males. Based on the number of nestlings that returned for subsequent breeding seasons, however, the reproductive
success of monogamous and polygynous males did not differ significantly. These data raise the question as to why males attempt
polygyny.
Received: 16 August 1999 / Received in revised form: 15 March 2000 / Accepted: 18 March 2000 相似文献
570.
Fuzzy Sets and Threatened Species Classification 总被引:1,自引:0,他引:1