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851.
The pathway for the flow of salt-marsh grass production into marsh food-webs is still not well defined. We compared the abilities
of three marsh macroinvertebrates [salt marsh periwinkles, Littoraria irrorata (Say) (=Littorina irrorata), salt-marsh coffee-bean snails, Melampus bidentatus (Say); and a talitrid amphipod, Uhlorchestia spartinophila Bounsfield and Heard] to access standing-dead leaves of smooth cordgrass (Spartina alterniflora Loisel). The invertebrates were incubated with naturally-decaying leaves, and the rates of removal of organic matter and
living fungal biomass (ergosterol) were measured. The impact of invertebrate activity upon fungal growth rates was measured
as rates of fungal-membrane synthesis (incorporation of radioacetate into ergosterol). The removal rates of organic leaf biomass
per mg individual biomass were highest for amphipods (700 μg mg−1 d−1) and lowest for periwinkles (90 μg mg−1 d−1), but the relatively large biomass of the snails made their removal rates per individual greater than those of amphipods.
Net removal of ergosterol by all three invertebrates was >50% for yellow-brown (early-decay) leaf blades. For fully-brown
(advanced-decay) blades, >50% removal of ergosterol was found only for periwinkles; exposure to coffee-bean snails and amphipods
resulted in a net ergosterol reduction of ≤20%. The lower net reduction of living fungal biomass by coffee-bean snails and
amphipods may have been due to fungal-growth stimulation (2.3-fold stimulation in coffee-bean snails and 1.5-fold stimulation
in amphipods). Grazing by periwinkles did not stimulate fungal growth, possibly because of its high intensity. Grazing by
these three salt-marsh shredders may affect marsh-grass shoot-decay in different ways. Periwinkles may abbreviate the period
of fungal production, and incorporate the decaying material relatively quickly into snail biomass and fecal-pellet rain to
the sediments. Coffee-bean snails and amphipods may enhance and prolong fungal production, along with the formation of fecal-pellet
rain. All three invertebrates fed preferentially on leaf blades rather than leaf sheaths, and feeding rates of gastropods
were higher during the night than during the day.
Received: 25 November 1998 / Accepted: 4 November 1999 相似文献
852.
853.
Genetic variation in 15 Holothuria (Microthele) nobilis (Selenka, 1867) populations on the Great Barrier Reef was studied at seven polymorphic allozyme loci. Although populations
were separated by distances up to 1300 km, there were no apparent restrictions to gene flow (F
ST
values were not significantly different from 0) and the maximum Nei's unbiased genetic distance was 0.003. Populations were
in Hardy–Weinberg equilibrium at all loci. The estimated maximum sexual input [this is the ratio of the number of sexually
produced individuals (N*) to the sample size (N
i
)] and the minimum sexual input [this is the ratio of the number of genotypes (N
go
) over the sample size (N
i
)] were used as estimators for the amount of asexual reproduction. Both parameters suggested that H. nobilis reproduces solely by sexual means (N*:N
i
: = 1; N
go
:N
i
= 0.74 to 1). The allozyme data indicated high gene flow between populations, but the possibility that allozyme frequencies
may not be at equilibrium means that it was not possible to distinguish whether the patterns reflect present-day dispersal
or dispersal that occurred in the past.
Received: 27 March 2000 / Accepted: 29 June 2000 相似文献
854.
The reproductive ecology of two eastern Pacific zooxanthellate coral species was examined as part of a continuing series of studies relating bleaching/mortality events caused by the El Niño–Southern Oscillation disturbance, and is described for study sites in Costa Rica, Panamá, and the Galápagos Islands (Ecuador). This study deals with the sibling agariciid species Pavona varians and Pavona sp.a over a 13?yr period (1985 to 1997). Both Pavona species are broadcast-spawners with some gonochoric, but mostly sequential hermaphroditic colonies. Minimum colony sizes (and ages) at first reproduction were 5?cm (5?yr) and 3?cm (2 to 3?yr), respectively, in P. varians and Pavona sp.a. In the Panamá and Galápagos populations, gonochoric colonies spawn eggs or sperm at least monthly. Six fecundity attributes were not significantly different in the two species, but the eggs of P. varians are white to beige and positively buoyant, and those of Pavona sp.a are dark green and neutrally to negatively buoyant. Eggs of both species lack zooxanthellae. Both species are reproductively active year-round, with maximum activity in the dry season in the nonupwelling Gulf of Chiriquí, and in the wet season in the upwelling Gulf of Panamá. Spawning is predominantly during full moon, and possibly also at new moon at most study sites. Spawning in P. varians and Pavona sp.a is 12?h out of phase, with the former species spawning ~1?h before sunrise and the latter about 1?h after sunset. The fecundity of Pavona spp. at Caño and the Galápagos Islands was much greater (19?900 to 27?900 eggs cm?2?yr?1) than at all Panamá sites (14?800 to 19?800 eggs cm?2?yr?1). Intraspecific crosses in both species resulted in swimming planula larvae after 25 to 36?h. Recruitment of P. varians was highest in Panamá, moderate in Costa Rica, and nil in the Galápagos Islands, matching, respectively, the contributions of P. varians to the pre-1982/1983 El Niño coral-population abundances in these areas. Recruitment success of P. varians at Uva Island was significantly related to maximum monthly positive sea surface-temperature (SST) anomalies that occurred in the year preceding recruitment over the period 1982 to 1996; recruitment failed when SST anomalies exceeded 1.6 to 1.9?C° during the severe ENSO events of 1982/1983 and 1997/1998. 相似文献
855.
An experimental study on recolonization and succession of marine macrobenthos in defaunated sediment 总被引:3,自引:0,他引:3
Hypoxia/anoxia in coastal waters is a world wide problem which often results in mass mortality and defaunation of benthos.
In this study, field experiments were carried out to examine recolonization and succession of macrobenthic infauna in defaunated
sediments, and the time required for recovery from complete defaunation to a stable community. Trays (33 cm length × 25.5 cm
width × 11 cm depth) of defaunated sediment were exposed at the subtidal of a pristine site in subtropical Hong Kong. Temporal
changes of macrobenthic communities in defaunated sediment were analyzed by univariate and multivariate statistics, and compared
with those in undisturbed natural sediment at the same site. Initial colonization of macrobenthos occurred rapidly. A total
of 42 species was found, with an average of 258 animals per tray and 24 species per tray recorded in the first month. Abundance
showed a small peak (496 animals per tray) after 3 months, reached a sharp peak (1154 animals per tray) after 6 months, and
declined thereafter. Species number increased gradually, reached a maximum (68 species per tray) after 9 months, and then
decreased. Recolonization was predominantly contributed by larval settlement rather than adult migration. Temporal changes
in abundance, species number and diversity of the macrobenthic community in defaunated sediment resemble the spatial changes
along a decreasing pollution gradient previously defined by other authors. Results of this experiment suggest that newly available
sediment may allow more species to colonize (or coexist) than sediment pre-occupied by an established community. This is probably
due to less interspecific competition in the former habitat. No significant difference in abundance or species richness was
observed between defaunated and natural sediments after 15 months, suggesting that a stable community had been achieved, although
minor variations in species composition were still discernible between defaunated and natural sediments.
Received: 24 May 1999 / Accepted: 20 October 1999 相似文献
856.
Spatial and temporal patterns of predation on seeds of the seagrass Phyllospadix torreyi S. Watson were quantified at four sites near Santa Barbara, California, USA. Over a period of four flowering seasons during
1995 to 1998, monthly patterns of seed fall and intensity of seed predation were similar among sites, but were temporally
quite variable. Abundance of dispersed seeds varied greatly both among seasons and years. Within any one year, seeds were
present in the environment every month, but they peaked in abundance during the fall months following the annual flowering
period. Seeds were more abundant during the earlier years of the study. The intensity of predation steadily increased throughout
the study period, from a low of ≤10% seeds consumed during 1995 to ∼50% consumption by 1997, and it was not correlated with
abundance of seeds in the environment. Pre-dispersal seed loss also was estimated in two flowering seasons by counting the
numbers of seeds consumed prior to release from the plant, and was relatively low (<15%). No differences were detected among
the four study sites in patterns of pre-dispersal loss. Field surveys were done at two sites to identify potential seed predators.
Three of the most abundant species identified in these surveys were tested in the laboratory to determine size-specific patterns
of feeding activity and to assess which early life stages of P. torreyi (fruits within spadices, dehisced fruits, seedlings) were most vulnerable to predation. All three of the early life stages
were consumed. The crabs Pugettiaproducta (Randall) and Pachygrapsus crassipes Randall were much more voracious predators than the isopod Idotea resecata Stimpson.
Received: 8 June 1999 / Accepted: 2 February 2000 相似文献
857.
The Norway lobster Nephrops norvegicus (L.) inhabits burrows in muddy clay sediments (e.g. on the Swedish west coast), where an autumnal oxygen deficiency in the
bottom water can occur. Our experiments investigated whether the irrigation of the burrows would reflect a behavioural adaptation
to hypoxia, and whether any gender differences of such behaviour exist. Irrigation is performed by the pleopods which may
compensate for a decreasing oxygen tension. Pleopod activity (total number of strokes per sampling time), associated with
oxygen concentration and gender, was studied in N. norvegicus kept in artificial burrows resembling their natural habitat. Male and female lobsters were separately exposed to either normoxia
(70% oxygen saturation) or hypoxia (30% oxygen saturation). A sexual difference in behaviour was found, where females irrigated
the burrow less than males during normoxia. Females showed a significant increase of pleopod activity in hypoxia compared
with normoxic conditions, which was not displayed by the males probably due to the degree of individual variation found. However,
when only males were studied during progressive hypoxia (from 60 to 5% oxygen saturation), following any changes of irrigational
behaviour, a significant increase of accumulated pleopod activity occurred. A major increase of pleopod activity appeared
between 60 and 50% oxygen saturation, below which the activity remained high until a critical point (<10% saturation, 11 °C,
33 psu) where irrigation dropped to a level close to that of normoxic values. Activity sessions during hypoxia were longer
and had a higher stroke rate than during normoxia.
Received: 22 October 1997 / Accepted: 26 February 1998 相似文献
858.
The sibling species, Pseudocalanus moultoni (Frost, 1989) and P. newmani (Frost, 1989), occur sympatrically on Georges Bank. Taxonomic discrimination of the species relies on subtle morphological
characteristics, making routine identification of the species very difficult. DNA sequence variation of two mitochondrial
genes, 16S rRNA and cytochrome oxidase I (COI), reliably discriminated P. moultoni and P.␣newmani. Levels of DNA sequence variation for both genes were consistent with those between species of calanoid copepods. A molecular
systematic protocol (based on allele-specific PCR amplification) was designed from the COI sequences and used to discriminate
females of the two species. The distributions and relative abundances of the two species were mapped for April 1996 based
on samples of 15 to 30 females from 12 collections across Georges Bank. The results of this study indicated that P. moultoni females predominated along the northern flank of Georges Bank, while P. newmani females were common on the southern flank, deeper than the 60-m isobath.
Received: 16 May 1997 / Accepted: 26 April 1998 相似文献
859.
Recovery of benthic communities at Macquarie Island (sub-Antarctic) following a small oil spill 总被引:5,自引:0,他引:5
Intertidal communities at Macquarie Island (sub-Antarctic) were re-evaluated seven years after a small oil spill resulting
from the grounding of the “Nella Dan”, and six years after the first assessment of biological impact. Sampling was conducted
to evaluate community structure in three zones of the exposed rocky shore (upper red, kelp, and lower red) and in samples
of Durvillaea antarctica holdfasts. There were no significant differences between the community structure in oiled and control locations in any of
the three shore zones, but holdfast macrofaunal communities at oiled sites still showed evidence of impact. Holdfast community-structure
in samples from heavily oiled sites showed moderate levels of recovery, with increased abundances of species which were considered
sensitive to the oiling in the first post-impact studies and decreased abundances of opportunistic polychaete and oligochaete
worms. In contrast, samples collected from the moderately oiled location at Secluded Bay showed little evidence of recovery.
Holdfasts at this site were filled with sediment containing traces of diesel oil, and the macrofaunal community was dominated
by opportunistic worms. This study clearly indicates that even small incidents of anthropogenic perturbation can have long-lasting
consequences for marine communities at Macquarie Island.
Received: 23 July 1997 / Accepted: 5 February 1998 相似文献
860.
Through two-choice gustatory experiments, a␣preference hierarchy was established␣for␣the␣herbivorous gastropod Norrisianorrisi Sowerby, with laminarialean kelps preferred over all other seaweeds. Among the kelps, laminae of Macrocystispyrifera were slightly preferred over Egregiamenziesii, and both were strongly preferred over sporophylls of Eiseniaarborea. E.arborea, the least preferred kelp, was consistently chosen over other algae common in the snail's habitat (Halidrysdioica, Dictyotaflabellata, and Pterocladiacapillacea) and over seaweeds believed to be edible and palatable based on their morphology, structure, and secondary chemistry (Endarachnebinghamiae, Mazzaellaflaccida, and Ulvalobata). The morphologies and structural toughness of tested seaweeds varied significantly as did their nutritional (% carbon, %
nitrogen, C:N ratio, and % ash) contents and phlorotannin concentrations; however, snails preferred to feed on kelps regardless
of nutritional content, toughness or phlorotannin concentration; and among kelps preferred to feed on the least tough species
(based on penetrometer measurements), which also were those containing the lowest phlorotannin concentrations. Preference
for kelp was not upheld in experiments using agarose thalli to which freeze-dried powder, of either the kelp Eiseniaarborea or non-kelp Endarachnebinghamiae was added, suggesting the destruction of attractant chemicals during the making of the artificial foods. Our data suggest
that the preference of N. norrisi for kelps over other potentially edible and palatable seaweeds may not be related to nutritional content, but instead may
have evolved in response to factors such as availability, habitat provision, or refuge from predation.
Received: 27 September 1996 / Accepted: 7 October 1997 相似文献