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991.
Numerous objects of gold displaying an impressive variety of types and manufacturing techniques are known from the Late Bronze Age (LBA) contexts of Mycenaean Greece, but very little is known about the origin and processing of gold during the second millennium B.C: . Ancient literature and recent research indicate that northern Greece is probably the richest gold-bearing region in Greece, and yet, very little evidence exists regarding the exploitation of its deposits and the production as well as use of gold in the area during prehistory. The unusual find of a group of small stone crucibles at the prehistoric settlement of Thessaloniki Toumba, one with visible traces of gold melting, proves local production and offers a rare opportunity to examine the process of on-site gold working. Furthermore, the comparison of the chemical composition of prehistoric artefacts from two settlements with those of gold deposits in their immediate areas supports the local extraction of gold and opens up the prospect for some of the Mycenaean gold to have originated in northern Greece. The scarcity of gold items in northern Greek LBA contexts may not represent the actual amount of gold produced and consumed, but could be a result of the local social attitudes towards the circulation and deposition of artefacts from precious metals. 相似文献
992.
In the honeybee, Apis mellifera, the queen monopolizes reproduction, while the sterile workers cooperate harmoniously in nest maintenance. However, under queenless (QL) conditions, cooperation collapses and reproductive competition among workers ensues. This is mediated through aggression and worker oviposition, as well as shifts in pheromones, from worker to queen-like composition. Many studies suggest a dichotomy between conflict resolution through aggression or through pheromonal signaling. In this paper, we demonstrate that both phenomena comprise essential components of reproductive competition and that pheromone signaling actually triggers the onset of aggression. We kept workers as QL groups until first aggression was observed and subsequently determined the contestants' reproductive status and content of the mandibular (MG) and Dufour's glands (DG). In groups in which aggression occurred early, the attacked bee had consistently more queen-like pheromone in both the MG and DG, although both contestants had undeveloped ovaries. In groups with late aggression, the attacked bee had consistently larger oocytes and more queen-like pheromone in the DG, but not the MG. We suggest that at early stages of competition, the MG secretion is utilized to establish dominance and that the DG provides an honest fertility signal. We further argue that it is the higher amount of DG pheromone that triggers aggression. 相似文献
993.
Cribb BW Stewart A Huang H Truss R Noller B Rasch R Zalucki MP 《Die Naturwissenschaften》2008,95(5):433-441
Previously, the presence of metals in arthropod mandibles has been linked with harder cuticle, and in termites, a 20% increase in hardness has been found for mandibles containing major quantities of zinc. The current study utilises electron microscopy and energy-dispersive X-ray microanalysis to assess incidence and abundance of metals in all extant subfamilies of the Isoptera. The basal clades contain no zinc and little to no manganese in the cutting edge of the mandible cuticle, suggesting that these states are ancestral for termites. However, experimentation with mandibles in vitro indicates the presence of some elements of the cuticular biochemistry necessary to enable uptake of zinc. The Termopsidae, Serritermitidae, Rhinotermitidae and Termitidae all contain minor quantities of manganese, while trace to minor quantities of zinc occur in all except the Serritermitidae. In contrast, all Kalotermitidae or drywood termites contain major levels of zinc in the mandible edge. Diet and life type are explored as links to metal profiles across the termites. The presence of harder mandibles in the drywood termites may be related to lack of access to free water with which to moisten wood. Scratch tests were applied to a set of mandibles. The coefficient of friction for Cryptotermes primus (Kalotermitidae) mandibles, when compared with species from other subfamilies, indicates that zinc-containing mandibles are likely to be more scratch resistant. 相似文献
994.
Sherratt TN 《Die Naturwissenschaften》2008,95(8):681-695
It is now 130 years since Fritz Müller proposed an evolutionary explanation for the close similarity of co-existing unpalatable prey species, a phenomenon now known as Müllerian mimicry. Müller's hypothesis was that unpalatable species evolve a similar appearance to reduce the mortality involved in training predators to avoid them, and he backed up his arguments with a mathematical model in which predators attack a fixed number (n) of each distinct unpalatable type in a given season before avoiding them. Here, I review what has since been discovered about Müllerian mimicry and consider in particular its relationship to other forms of mimicry. Müller's specific model of associative learning involving a "fixed n" in a given season has not been supported, and several experiments now suggest that two distinct unpalatable prey types may be just as easy to learn to avoid as one. Nevertheless, Müller's general insight that novel unpalatable forms have higher mortality than common unpalatable forms as a result of predation has been well supported by field experiments. From its inception, there has been a heated debate over the nature of the relationship between Müllerian co-mimics that differ in their level of defence. There is now a growing awareness that this relationship can be mediated by many factors, including synergistic effects between co-mimics that differ in their mode of defence, rates of generalisation among warning signals and concomitant changes in prey density as mimicry evolves. I highlight areas for future enquiry, including the possibility of Müllerian mimicry systems based on profitability rather than unprofitability and the co-evolution of defence. 相似文献
995.
Tan K Yang MX Radloff SE Hepburn HR Zhang ZY Luo LJ Li H 《Die Naturwissenschaften》2008,95(12):1165-1168
Although the structure of the dance language is very similar among species of honeybees, communication of the distance component
of the message varies both intraspecifically and interspecifically. However, it is not known whether different honeybee species
would attend interspecific waggle dances and, if so, whether they can decipher such dances. Using mixed-species colonies of
Apis cerana and Apis mellifera, we show that, despite internal differences in the structure of the waggle dances of foragers, both species attend, and act
on the information encoded in each other’s waggle dances but with limited accuracy. These observations indicate that direction
and distance communication pre-date speciation in honeybees. 相似文献
996.
A new assemblage of basal dichobunoid artiodactyls from the middle-Eocene Shanghuang fissure fillings includes the diacodexeid
Jiangsudon shanghuangensis gen. and sp. nov., a new species of the lantianine dichobunoid Elaschitotherium, Elaschitotherium crepaturus sp. nov., and an indeterminate suoid which is presently the earliest record of this clade. Diacodexeids are also represented
by two forms provisionally referred to cf. Diacodexis sp. and to an indeterminate Diacodexeidae, respectively. The occurrence of diacodexeids in Shanghuang contrasts with the
early and earliest middle-Eocene chronological range of the family in Europe and North America and suggests that the stratigraphic
range of the family in Asia extends up to the middle Eocene. This may reflect particular habitats in coastal China that may
have been relatively stable during the early and middle Eocene, thus preserving forest-dwelling artiodactyls that became extinct
in the other Holarctic regions. Compared to other supposedly coeval North American, European, and Asian faunas, the Shanghuang
mammalian assemblage is most similar to early Uintan faunas of North America but is also remarkable in recording forms close
to taxa that are characteristic of the Wasatchian and Bridgerian North American Land Mammal Ages. The Irdinmanhan age of the
Shanghuang fauna is supported by the mammalian assemblage recovered from the fissure D, but an Arshantan age cannot be completely
ruled out at this point. Although the Shanghuang assemblage is biased towards preservation of small components of the mammalian
fauna, the Shanghuang fauna provide an important and unique window into the Eocene diversity and early evolution of cetartiodactyls
in eastern Asia. 相似文献
997.
Sperm number is often a good predictor of success in sperm competition; however, it has become increasingly clear that, for
some species, variation in probability of paternity cannot be explained by sperm number alone. Intraspecific variation in
ejaculate characteristics, such as the number of viable sperm and sperm longevity, may play an equally important role in determining
fertilization success. Here, we assess variation among ejaculates in three factors that may contribute to fertilization success
(number of sperm per ejaculate, viability, and longevity), in a population of Peron’s tree frog (Litoria peronii). We detected large variation among males in the number of sperm per ejaculate and the proportion of viable sperm within
ejaculates, which could not be explained by variation in either male size or body condition. However, the proportion of viable
sperm released by males increased over the season. Finally, we assessed sperm longevity (proportion viable sperm determined
using a dual-fluorochrome vital dye) at two different temperatures. At 23°C, on average, 75% of sperm remained viable after
2 h, but there were significant differences amongst males with the percentage of viable sperm ranging from 43% to 95%. For
sperm incubated at 4°C, ejaculates varied fivefold in sperm longevity with some males having 50% viable sperm after 5 days.
Our data suggest that ejaculate characteristics (sperm number, viability, and longevity) vary widely in Peron’s tree frog
and may therefore play an important role in determining siring success both in the presence and absence of sperm competition.
We discuss the results in relation to selection on ejaculate traits via natural and sexual selection in this and other amphibians. 相似文献
998.
Flower colour is an important signal used by flowering plants to attract pollinators. Many anthophilous insects have an innate colour preference that is displayed during their first foraging bouts and which could help them locate their first nectar reward. Nevertheless, learning capabilities allow insects to switch their colour preferences with experience and thus, to track variation in floral nectar availability. Manduca sexta, a crepuscular hawkmoth widely studied as a model system for sensory physiology and behaviour, visits mostly white, night-blooming flowers lacking UV reflectance throughout its range in the Americas. Nevertheless, the spectral sensitivity of the feeding behaviour of naïve moths shows a narrow peak around 450 nm wavelengths, suggesting an innate preference for the colour blue. Under more natural conditions (i.e. broader wavelength reflectance) than in previous studies, we used dual choice experiments with blue- and white-coloured feeders to investigate the innate preference of naïve moths and trained different groups to each colour to evaluate their learning capabilities. We confirmed the innate preference of M. sexta for blue and found that these moths were able to switch colour preferences after training experience. These results unequivocally demonstrate that M. sexta moths innately prefer blue when presented against white flower models and offer novel experimental evidence supporting the hypothesis that learning capabilities could be involved in their foraging preferences, including their widely observed attraction to white flowers in nature. 相似文献
999.
Vulnerability assessment and adaptation to the impacts of sea level rise on the Kingdom of Bahrain 总被引:2,自引:0,他引:2
S. Al-Jeneid M. Bahnassy S. Nasr M. El Raey 《Mitigation and Adaptation Strategies for Global Change》2008,13(1):87-104
This paper assesses quantitatively the impact of sea level rise (SLR) at the global and regional scale as a result of climate
change (CC) on the coastal areas of the Kingdom of Bahrain’s islands (36 Islands). The standard Intergovernmental Panel on
Climate Change (IPCC) guidelines was modified as appropriate for the situation of the study area. Geographic Information Systems
(GIS) coupled with Remote Sensing (RS) were used as the main techniques of collecting, analyzing, modeling simulating and
disseminating information to build SLR scenarios in a geographically referenced context. Also, these tools were used to assess
vulnerability and risk of the coastal area of the islands with the expectation that coastal planner and government authorities
will profit from integrating these knowledge into a broad based environmental decision making. Three SLR scenarios: low, moderate
and high were developed to examine the impacts from SLR on all islands. The low SLR scenario (Optimistic) assumes a 0.5-m
rise above current sea level, the moderate scenario (Intermediate) assumes a one meter rise, and the high scenario (Pessimistic)
assumes a 1.5 m rise in sea level. Two more SLR scenarios were assumed to perform risk analysis, a 2 and 5 meter rise above
current sea level. The simulation of SLR are quite straightforward, emphasizing on the uses of both of the data that are incorporated
from the satellite images and the created Digital Elevation Model (DEM) to estimate SLR scenarios that are adapted in the
study. These data were used to predict consequences of the possibility of the rise in sea level at different scenarios which
may alter the landuse and patterns of human communities. Results indicate that low-lying coastal areas of Bahrain islands
are at risk from the effects of any SLR resulting from CC. These islands are vulnerable under different SLR Scenarios. More
than 17% of the country total area may be inundated under 1.5 m SLR in 2100. The total area that might be lost under different
sea level scenarios will vary from more than 77 km2 if SLR reaches 0.5 m, to about 100 km2 under 1.0 m SLR and may reach 124 km2 under 1.5 m SLR scenario. The total inundated areas due to risk scenarios will reach 133 km2, if the SLR rises to 2.0 m, and it is estimated to be more than (22%) of the main island total area. Under the second scenario,
if the SLR reaches 5.0 m, the main islands will lose approximately half of its area (47%) equal to 280 km2. Hawar islands group will lose about (30%) of its total area under 2.0 m SLR, which is about 15.5 km2.A SLR adaptation policy framework (APF) and adaptation policy initiatives (APIs) are suggested for planners to build upon
for reducing the likely effects of SLR in the Kingdom of Bahrain. The framework is composed of four steps namely, acquisition
of information, planning and design, implementation and monitoring and evaluation. A general policy framework for a national
response to SLR is suggested. Additionally, a range of policy adaptation options/initiatives to sustain coastal developments
under the likely effects of SLR are recommended. 相似文献
1000.
Kenneth T. Gillingham Steven J. Smith Ronald D. Sands 《Mitigation and Adaptation Strategies for Global Change》2008,13(7):675-701
In the coming century, modern bioenergy crops have the potential to play a crucial role in the global energy mix, especially
under policies to reduce carbon dioxide emissions as proposed by many in the international community. Previous studies have
not fully addressed many of the dynamic interactions and effects of a policy-induced expansion of bioenergy crop production,
particularly on crop yields and human food demand. This study combines an updated agriculture and land use (AgLU) model with
a well-developed energy-economic model to provide an analysis of the effects of bioenergy crops on energy, agricultural and
land use systems. The results indicate that carbon dioxide mitigation policies can stimulate a large production of bioenergy
crops, dependent on the level of the policy. This production of bioenergy crops can lead to several impacts on the agriculture
and land use system: decreases in forestland and unmanaged land, decreases in the average yield of food crops, increases in
the prices of food crops, and decreases in the level of human demand of calories.
相似文献
Steven J. Smith (Corresponding author)Email: |