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131.
Honey bee foragers as sensory units of their colonies 总被引:5,自引:0,他引:5
Thomas D. Seeley 《Behavioral ecology and sociobiology》1994,34(1):51-62
Forager honey bees function not only as gatherers of food for their colonies, but also as sensory units shaped by natural selection to gather information regarding the location and profitability of forage sites. They transmit this information to colony members by means of waggle dances. To investigate the way bees transduce the stimulus of nectar-source profitability into the response of number of waggle runs, I performed experiments in which bees were stimulated with a sucrose solution feeder of known profitability and their dance responses were videorecorded. The results suggest that several attributes of this transduction process are adaptations to enhance a bee's effectiveness in reporting on a forage site. (1) Bees register the profitability of a nectar source not by sensing the energy gain per foraging trip or the rate of energy gain per trip, but evidently by sensing the energetic efficiency of their foraging. Perhaps this criterion of nectar-source profitability has been favored by natural selection because the foraging gains of honey bees are typically limited by energy expenditure rather than time availability. (2) There is a linear relationship between the stimulus of energetic efficiency of foraging and the response of number of waggle runs per dance. Such a simple stimulus-response function appears adequate because the range of suprathreshold stimuli (max/min ratio of about 10) is far smaller than the range of responses (max/min ratio of about 100). Although all bees show a linear stimulus-response function, there are large differences among individuals in both the response threshold and the slope of the stimulus-response function. This variation gives the colony a broader dynamic range in responding to food sources than if all bees had identical thresholds of dance response. (3) There is little or no adaptation in the dance response to a strong stimulus (tonic response). Thus each dancing bee reports on the current level of profitability of her forage site rather than the changes in its profitability. This seems appropriate since presumably it is the current profitability of a forage site, not the change in its profitability, which determines a site's attractiveness to other bees. (4) The level of forage-site quality that is the threshold for dancing is tuned by the bees in relation to forage availability. Bees operate with a lower dance threshold when forage is sparse than when it is abundant. Thus a colony utilizes input about a wide range of forage sites when food is scarce, but filters out input about low-reward sites when food is plentiful. (5) A dancing bee does not present her information in one spot within the hive but instead distributes it over much of the dance floor. Consequently, the dances for different forage sites are mixed together on the dance floor. This helps each bee following the dances to take a random sample of the dance information, which is appropriate for the foraging strategy of a honey bee colony since it is evidently designed to allocate foragers among forage sites in proportion to their profitability. 相似文献
132.
Bert?H?lldoblerEmail author E.?David Morgan Neil?J.?Oldham Jürgen?Liebig Yue? Liu 《Chemoecology》2004,14(2):101-106
Summary. Dufour gland secretions of the harvester ants
Pogonomyrmex barbatus, P. rugosus
and P. maricopa were
investigated. The glandular blends exhibit species specific
patterns, but more importantly, based on indiviual profiles of
Dufour gland contents of colony members it is possible to
separate the colonies from each other in each species. It is
possible that these collective colony specific cues serve as
longer lasting trunk route markers. Behavioral tests demonstrate
that workers of P. rugosus
prefer homing routes marked by members of their own colony to routes marked
by a foreign conspecific colony. 相似文献
133.
In many environmental studies measuring the amount of a contaminant in a sampling unit is expensive. In such cases, composite sampling is often used to reduce data collection cost. However, composite sampling is known to be beneficial for estimating the mean of a population, but not necessarily for estimating the variance or other parameters. As some applications, for example, Monte Carlo risk assessment, require an estimate of the entire distribution, and as the lognormal model is commonly used in environmental risk assessment, in this paper we investigate efficiency of composite sampling for estimating a lognormal distribution. In particular, we examine the magnitude of savings in the number of measurements over simple random sampling, and the nature of its dependence on composite size and the parameters of the distribution utilizing simulation and asymptotic calculations. 相似文献
134.
Lynn E. Fletcher 《Behavioral ecology and sociobiology》2007,61(12):1809-1821
Group living confers both benefits and costs to the individuals involved. Benefits may include enhanced defense, thermoregulation,
and increased foraging efficiency while costs often involve competition for resources such as food, shelter, and mates. Communication
provides a medium of exchange among individuals engaged in either cooperative or competitive interactions. The functional
analysis of signals within groups therefore requires testing both cooperative and competitive functions, although the latter
is infrequently done. In this paper, I study the use of two vibrational signals in a gregarious, processionary Australian
sawfly larva, Perga affinis: tapping and contractions. Tapping involves striking the substrate with the sclerotized portion of the abdominal tail and
a contraction is a fast, whole-body twitch, which is both tactile and vibrational in its transmission. For tapping, I first
demonstrate that it is a form of communication, as tapping of one larva elicits tapping in another, and that it is transmitted
through substrate vibrations. I then test whether the signal is mostly cooperative or competitive in nature by examining it
in light of two hypotheses: (1) the Group Coordination hypothesis, stating that the signal functions to maintain group cohesiveness
and (2) the Competitive Signaling hypothesis, stating that tapping serves as a competitive assessment signal between larvae
while feeding. For contractions, I test only the group coordination hypothesis that they serve to coordinate and initiate
group movement. Results support the group coordination hypothesis for each signal. While feeding, lone larvae (without potential
competitors) were significantly more likely to tap than those in groups, and this trend continued in non-feeding situations.
Contractions regularly preceded periods of group movement during processions and were given with increasing frequency before
departure from preforaging clusters. The vibrational signals in this processionary species likely function cooperatively to
maintain group cohesiveness and coordinate movement. 相似文献
135.
Kevin J. Delaney J. Andrew Roberts George W. Uetz 《Behavioral ecology and sociobiology》2007,62(1):67-75
Male signaling behaviors are often studied in a single context but may serve multiple functions (e.g., in male–male competition
and female mate choice). We examined the issue of dual function male signals in a wolf spider species Schizocosa ocreata (Hentz) that displays the same species-specific signaling behaviors in both male–male and male–female contexts. These signaling
behaviors have been described as either aggression or courtship according to the context observed. We tested the possibility
of dual functions by comparing the relationship between behaviors and outcome of male–male contests (winner/loser) and male–female
mating encounters (mating success). Frequency, rate, and mean duration of signaling behaviors did not vary with outcome of
male–male contests, which appears instead to be based upon relative size and body mass. Winners of contests had significantly
greater body mass than losers, and greater mass relative to opponents was significantly associated with probability of winning.
Overall, signaling rates were much higher in male–female interactions than in male–male contests and were higher for males
that successfully mated than for those that did not mate. Mean duration of some male displays was also greater for males that
successfully mated. However, male size was not associated with probability of mating. Taken together, results suggest an intersexual
selection context for the current function of male signals in these wolf spiders and that increased display vigor is associated
with male mating success. 相似文献
136.
质量保证与质量控制是水质自动监测中十分重要的技术工作和管理工作,包括人员素质、运行与管理机制、试剂与标准溶液的配制及文件记录等.在实际工作中,监测人员可根据仪器的运行状况,定期时仪器进行校准、标准溶液的核查、比对实验验证及试剂有效性检查,用以检查仪器的基线飘移情况.如果分析中出现一个异常值,它的前后均为正常值,这种数值大多是由仪器的进样、仪器内部试剂传输等原因所致,如果发生原因不明的数据异常,就要及时检查系统的各个环节并采集实际水样进行人工分析,直至查清原因,剔除异常值,并在周报中加以说明,以确保上报数据的准确性、精密性和可比性. 相似文献
137.
138.
139.
自由飞行能够提高空域容量从而缓解空域拥挤问题,在保障自由飞行安全性的基础上尽可能减小飞机的安全间距,需要研究自由飞行条件下飞机的安全评估问题。分析了所需CNS性能、防撞系统、风的因素等自由飞行下航空器定位误差的影响因素,考虑到航空器定位误差在短时间内具有累积性,提出采用INS、GPS数据融合算法对定位误差进行周期性修正。给出修正后的航空器定位误差分布在时间上的变化规律,并将其运用到自由飞行下碰撞风险评估中。结果表明,碰撞风险随安全间距增加呈下降趋势,以1.5×10-8次/飞行小时的目标安全水平为例,对航空器定位误差修正的最小安全间距为8 600 m,不对航空器定位误差进行修正的最小安全间距为9 100 m。 相似文献
140.
This paper presents a dynamic temperature model for a proton exchange membrane fuel cell (PEMFC) system. The proposed model overcomes the complexity of conventional models using first-order expressions consisting of load current and ambient temperature. The proposed model also incorporates a PEMFC cooling system, which depends upon the temperature difference between events. A dynamic algorithm is developed to detect load changing events and calculate instantaneous PEMFC temperature variations. The parameters of the model are extracted by employing the lightning search algorithm (LSA). The temperature characteristics of the NEXA 1.2 kW PEMFC system are experimentally studied to validate model performance. The results show that the proposed model output and the temperature data obtained from experiments for linear and abrupt changes in PEMFC load current are in agreement. The root-mean-square error between the model output and experimental results is less than 0.9. Moreover, the proposed model outperforms the conventional models and provides advantages such as simplicity and adaptability for low and high sampling data rates of input variables, namely, load current and ambient temperature. The model is not only helpful for simulations but also suitable for dynamic real-time controllers and emulators. 相似文献