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1.
The modern theory of biological evolution: an expanded synthesis 总被引:12,自引:2,他引:12
In 1858, two naturalists, Charles Darwin and Alfred Russel Wallace, independently proposed natural selection as the basic mechanism responsible for the origin of new phenotypic variants and, ultimately, new species. A large body of evidence for this hypothesis was published in Darwins Origin of Species one year later, the appearance of which provoked other leading scientists like August Weismann to adopt and amplify Darwins perspective. Weismanns neo-Darwinian theory of evolution was further elaborated, most notably in a series of books by Theodosius Dobzhansky, Ernst Mayr, Julian Huxley and others. In this article we first summarize the history of life on Earth and provide recent evidence demonstrating that Darwins dilemma (the apparent missing Precambrian record of life) has been resolved. Next, the historical development and structure of the modern synthesis is described within the context of the following topics: paleobiology and rates of evolution, mass extinctions and species selection, macroevolution and punctuated equilibrium, sexual reproduction and recombination, sexual selection and altruism, endosymbiosis and eukaryotic cell evolution, evolutionary developmental biology, phenotypic plasticity, epigenetic inheritance and molecular evolution, experimental bacterial evolution, and computer simulations (in silico evolution of digital organisms). In addition, we discuss the expansion of the modern synthesis, embracing all branches of scientific disciplines. It is concluded that the basic tenets of the synthetic theory have survived, but in modified form. These sub-theories require continued elaboration, particularly in light of molecular biology, to answer open-ended questions concerning the mechanisms of evolution in all five kingdoms of life.Dedicated to Prof. Dr. Dr. hc mult. Ernst Mayr on the occasion of his 100th birthdayThis revised version was published online in March 2004, with corrections to the caption of Figure 6. 相似文献
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Hopanoids are pentacyclic triterpenoid lipids occurring in bacteria. They are synthesized from isopentenyl units which are
formed in a new biosynthetic route leading to isopentenyl diphosphate. Six C5 units are joined to form squalene, the immediate precursor in hopanoid synthesis. In a highly complex cyclization reaction
that shares considerable similarities with that of oxidosqualene to sterols, the hopane skeleton is formed from squalene by
the squalene-hopene cyclase. Recent elucidation of the X-ray structure of this membrane-bound cyclase has shed some light
on the properties of this unusual enzyme. The active site is located in a cavity within the enzyme. The squalene substrate
diffuses through a channel structure from the membrane into this cavity and is there transformed into hopene. Polar side chains
are attached to hopene resulting in the amphiphilic molecular structure of many hopanoids. These hopanoids are membrane components
involved in regulating membrane fluidity and stability. However, the many structural variants of hopanoids indicate that they
may have other interesting but as yet unknown functions. 相似文献
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K. Bruce Jones Daniel T. Heggem Timothy G. Wade Anne C. Neale Donald W. Ebert Maliha S. Nash Megan H. Mehaffey Karl A. Hermann Anthony R. Selle Scott Augustine Iris A. Goodman Joel Pedersen David Bolgrien J. Max Viger Dean Chiang Cindy J. Lin Yehong Zhong Joan Baker Rick D. Van Remortel 《Environmental monitoring and assessment》2000,64(1):227-245
The Environmental Monitoring and Assessment Program (EMAP) is proposing an ambitious agenda to assess the status of streams and estuaries in a 12-State area of the western United States by the end of 2003. Additionally, EMAP is proposing to access landscape conditions as they relate to stream and estuary conditions across the west. The goal of this landscape project is to develop a landscape model that can be used to identify the relative risks of streams and estuaries to potential declines due to watershed-scale, landscape conditions across the west. To do so, requires an understanding of quantitative relationships between landscape composition and pattern metrics and parameters of stream and estuary conditions. This paper describes a strategic approach for evaluating the degree to which landscape composition and pattern influence stream and estuary condition, and the development and implementation of a spatially-distributed, landscape analysis approach. 相似文献
9.
Gunga HC Suthau T Bellmann A Friedrich A Schwanebeck T Stoinski S Trippel T Kirsch K Hellwich O 《Die Naturwissenschaften》2007,94(8):623-630
Both body mass and surface area are factors determining the essence of any living organism. This should also hold true for
an extinct organism such as a dinosaur. The present report discusses the use of a new 3D laser scanner method to establish
body masses and surface areas of an Asian elephant (Zoological Museum of Copenhagen, Denmark) and of Plateosaurus engelhardti, a prosauropod from the Upper Triassic, exhibited at the Paleontological Museum in Tübingen (Germany). This method was used
to study the effect that slight changes in body shape had on body mass for P. engelhardti. It was established that body volumes varied between 0.79 m3 (slim version) and 1.14 m3 (robust version), resulting in a presumable body mass of 630 and 912 kg, respectively. The total body surface areas ranged
between 8.8 and 10.2 m2, of which, in both reconstructions of P. engelhardti, ∼33% account for the thorax area alone. The main difference between the two models is in the tail and hind limb reconstruction.
The tail of the slim version has a surface area of 1.98 m2, whereas that of the robust version has a surface area of 2.73 m2. The body volumes calculated for the slim version were as follows: head 0.006 m3, neck 0.016 m3, fore limbs 0.020 m3, hind limbs 0.08 m3, thoracic cavity 0.533 m3, and tail 0.136 m3. For the robust model, the following volumes were established: 0.01 m3 head, neck 0.026 m3, fore limbs 0.025 m3, hind limbs 0.18 m3, thoracic cavity 0.616 m3, and finally, tail 0.28 m3. Based on these body volumes, scaling equations were used to assess the size that the organs of this extinct dinosaur have. 相似文献
10.
To ensure science better informs the decision-making process, researchers and policy/program managers need to understand and respect each other's way of working, culture and operational timelines. However, there is little practical guidance on how this should be done and even less documented experience with specific mechanisms that better link these two groups. The published literature on information transfer has largely emphasized the dissemination of standard packages of information to ill-defined constituencies whose needs for scientific information are not well understood. Environment Canada's National Water Research Institute, on behalf of the Canadian Council of Ministers of the Environment, led a series of “Linking Water Science to Policy Workshops” as one such mechanism by which recent science could be delivered to practitioners, and practitioners could identify their research needs to scientists and research managers. There is a pressing need to explore and share experiences using creative mechanisms for sustained dialogue and networking between scientists and policy and program managers. The lessons learned from the workshop series and the need for science to continually inform the decision-making process has particular relevance for Canada's Ecosystem Initiatives given their integrated, place-based focus on long-term restoration and protection, and the challenge of continually changing ecosystems.The Canadian Crown reserves the right to retain a non-exclusive, royalty free licence in and to any copyright. 相似文献