Ecological features and trends in the formation of the flora of liverworts under the impact of current volcanic and glacial activities: The example of Ushkovskii Volcano, Kamchatka

The flora of liverworts has been studied in the vicinity of the Bil’chenok glacier (the Ushkovskii volcano, Kamchatka), which is in a zone of volcanic ash fallout. Specific complexes of liverworts growing on young moraine and ash deposits have been identified. These complexes are dominated by thallose marchantioid species. Under the effect of periodic ash fallout (in the absence of glacial activity), communities are formed that contain increased proportions of xerophytes and species preferring sites with a disturbed ground vegetation cover.