State-of-the-art of ecological modelling with emphasis on development of structural dynamic models

The paper deals with two major problems in ecological modelling today, namely how to get reliable parameters? and how to build ecosystem properties into our models? The use of new mathematical tools to answer these questions is mentioned briefly, but the main focus of the paper is on development of structural dynamic models which are models using goal functions to reflect a current change of the properties of the biological components in the models. These changes of the properties are due to the enormous adaptability of the biological components to the prevailing conditions. All species in an ecosystem attempt to obtain most biomass, i.e. to move as far away as possible from thermodynamic equilibrium which can be measured by the thermodynamic concept exergy. Consequently, exergy has been proposed as a goal function in ecological models with dynamic structure, meaning currently changed properties of the biological components and in model language currently changed parameters. An equation to compute an exergy index of a model is presented. The theoretical considerations leading to this equation are not presented here but references to literature where the basis theory can be found are given. Two case studies of structural dynamic modelling are presented: a shallow lake where the structural dynamic changes have been determined before the model was developed, and the application of biomanipulation in lake management, where the structural dynamic changes are generally known. Moreover. it is also discussed how the same idea of using exergy as a goal function in ecological modelling may be applied to facilitate the estimation of parameters.