Pelagic crinoids (Roveacrinida, Crinoidea) discovered in the Neogene of Poland

Until recently, it has been assumed that pelagic crinoids, the roveacrinids (Roveacrinida, Crinoidea), became extinct during the Cretaceous-Paleogene boundary event. Recent finds of well-preserved roveacrinidal remains (brachials and radials) in the Danian (Early Paleogene) of Poland showed that they survived into the earliest Cenozoic. This group was thus characterized as a "dead clade walking". Here, we present fossil evidence that these pelagic crinoids survived in Poland until at least the Middle Miocene (Badenian, ca. 14?Myr ago)-more than 50?Myr after their supposed extinction. These Miocene roveacrinids constitute the first documented evidence of Roveacrinida in strata of Neogene age, thus prolonging the stratigraphic range of pelagic crinoids. This find characterizes the order as a "Lazarus taxon" rather than a "dead clade walking" group.