Development of a Bird Integrity Index: Measuring Avian Response to Disturbance in the Blue Mountains of Oregon, USA

The Bird Integrity Index (BII) presented here uses bird assemblage information to assess human impacts to 28 stream reaches in the Blue Mountains of eastern Oregon. Eighty-one candidate metrics were extracted from bird survey data for testing. The metrics represented aspects of bird taxonomic richness, tolerance or intolerance to human disturbance, dietary preferences, foraging techniques, and nesting strategies that were expected to be positively or negatively affected by human activities in the region. To evaluate the responsiveness of each metric, it was plotted against an index of reach and watershed disturbance that included attributes of land use/land cover, road density, riparian cover, mining impacts, and percent area in clearcut and partial-cut logging. Nine of the 81 candidate bird metrics remained after eliminating unresponsive and highly correlated metrics. Individual metric scores ranged from 0 to 10, and BII scores varied between 0 and 100. BII scores varied from 78.6 for a minimally disturbed, reference stream reach to 30.4 for the most highly disturbed stream reach. The BII responded clearly to varying riparian conditions and to the cumulative effects of disturbances, such as logging, grazing, and mining, which are common in the mountains of eastern Oregon. This BII for eastern Oregon was compared to an earlier BII developed for the agricultural and urban disturbance regime of the Willamette Valley in western Oregon. The BII presented here was sensitive enough to distinguish differences in condition among stream riparian zones with disturbances that were not as obvious or irreversible as those in the agricultural/urban conditions of western Oregon.     

Development of a bird integrity index: using bird assemblages as indicators of riparian condition

We describe the development of a bird integrity index (BII) that uses bird assemblage information to assess human impacts on 13 stream reaches in the Willamette Valley, Oregon, USA. We used bird survey data to test 62 candidate metrics representing aspects of bird taxonomic richness, tolerance or intolerance to human disturbance, dietary preferences, foraging techniques, and nesting strategies that were affected positively or negatively by human activities. We evaluated the metric responsiveness by plotting each one against a measure of site disturbance that included aspects of land use/land cover, road density, riparian cover, and stream channel and substrate conditions. In addition, we eliminated imprecise and highly correlated (redundant) metrics, leaving 13 metrics for the final index. Individual metric scores ranged continuously from 0 to 10, and index scores were weighted to range from 0 to 100. Scores were calibrated using historical species information to set expectations for the number of species expected under minimally disturbed conditions. Site scores varied from 82 for the least disturbed stream reach to 8.5 for an urban site. We compared the bird integrity index site scores with the performance of other measures of biotic response developed during this study: a fish index of biointegrity (IBI) and two benthic macroinvertebrate metrics. The three assemblages agreed on the general level of disturbance; however, individual sites scored differently depending on specific indicator response to in-stream or riparian conditions. The bird integrity index appears to be a useful management and monitoring tool for assessing riparian integrity and communicating the results to the public. Used together with aquatic indicator response and watershed data, bird assemblage information contributes to a more complete picture of stream condition.     

Nitrate removal effectiveness of a riparian buffer along a small agricultural stream in western Oregon

The Willamette Valley of Oregon has extensive areas of poorly drained, commercial grass seed lands. Little is know about the ability of riparian areas in these settings to reduce nitrate in water draining from grass seed fields. We established two study sites with similar soils and hydrology but contrasting riparian vegetation along an intermittent stream that drains perennial ryegrass (Lolium perenne L.) fields in the Willamette Valley of western Oregon. We installed a series of nested piezometers along three transects at each site to examine NO3-N in shallow ground water in grass seed fields and riparian areas. Results showed that a noncultivated riparian zone comprised of grasses and herbaceous vegetation significantly reduced NO3-N concentrations of shallow ground water moving from grass seed fields. Darcy's law-based estimates of shallow ground water flow through riparian zone A/E horizons revealed that this water flowpath could account for only a very small percentage of the streamflow. Even though there is great potential for NO3-N to be reduced as water moves through the noncultivated riparian zone with grass-herbaceous vegetation, the potential was not fully realized because only a small proportion of the stream flow interacts with riparian zone soils. Consequently, effective NO3-N water quality management in poorly drained landscapes similar to the study watershed is primarily dependent on implementation of sound agricultural practices within grass seed fields and is less influenced by riparian zone vegetation. Wise fertilizer application rates and timing are key management tools to reduce export of NO3-N in stream waters.     

Monitoring Forest Carbon Sequestration with Remote Sensing and Carbon Cycle Modeling

Sources and sinks of carbon associated with forests depend strongly on the management regime and spatial patterns in potential productivity. Satellite remote sensing can provide spatially explicit information on land cover, stand-age class, and harvesting. Carbon-cycle process models coupled to regional climate databases can provide information on potential rates of production and related rates of decomposition. The integration of remote sensing and modeling thus produces spatially explicit information on carbon storage and flux. This integrated approach was employed to compare carbon flux for the period 1992–1997 over two 165-km² areas in western Oregon. The Coast Range study area was predominately private land managed for timber production, whereas the West Cascades study area was predominantly public land that was less productive but experienced little harvesting in the 1990s. In the Coast Range area, 17% of the land base was harvested between 1991 and 2000. Much of the area was in relatively young, productive-age classes that simulations indicate are a carbon sink. Mean annual harvest removals from the Coast Range were greater than mean annual net ecosystem production. On the West Cascades study area, a relatively small proportion (< 1%) of the land was harvested and the area as a whole was accumulating carbon. The spatially and temporally explicit nature of this approach permits identification of mechanisms underlying land base carbon flux. Published online     

Summer Temperature Patterns in Headwater Streams of the Oregon Coast Range1

Abstract: Cool summertime stream temperature is an important component of high quality aquatic habitat in Oregon coastal streams. Within the Oregon Coast Range, small headwater streams make up a majority of the stream network; yet, little information is available on temperature patterns and the longitudinal variability for these streams. In this paper we describe preharvest spatial and temporal patterns in summer stream temperature for small streams of the Oregon Coast Range in forests managed for timber production. We also explore relationships between stream and riparian attributes and observed stream temperature conditions and patterns. Summer stream temperature, channel, and riparian data were collected on 36 headwater streams in 2002, 2003, and 2004. Mean stream temperatures were consistent among summers and generally warmed in a downstream direction. However, longitudinal trends in maximum temperatures were more variable. At the reach scale of 0.5‐1.7 km, maximum temperatures increased in 17 streams, decreased in seven streams and did not change in three reaches. At the subreach scale (0.1‐1.5 km), maximum temperatures increased in 28 subreaches, decreased in 14, and did not change in 12 subreaches. Models of increasing temperature in a downstream direction may oversimplify fine‐scale patterns in small streams. Stream and riparian attributes that correlated with observed temperature patterns included cover, channel substrate, channel gradient, instream wood jam volume, riparian stand density, and geology type. Longitudinal patterns of stream temperature are an important consideration for background characterization of water quality. Studies attempting to evaluate stream temperature response to timber harvest or other modifications should quantify variability in longitudinal patterns of stream temperature prior to logging.     

Development and Validation of an Aquatic Fine Sediment Biotic Index

The Fine Sediment Biotic Index (FSBI) is a regional, stressor-specific biomonitoring index to assess fine sediment (<2 mm) impacts on macroinvertebrate communities in northwestern US streams. We examined previously collected data of benthic macroinvertebrate assemblages and substrate particle sizes for 1,139 streams spanning 16 western US Level III Ecoregions to determine macroinvertebrate sensitivity (mostly at species level) to fine sediment. We developed FSBI for four ecoregion groupings that include nine of the ecoregions. The grouping were: the Coast (Coast Range ecoregion) (136 streams), Northern Mountains (Cascades, N. Rockies, ID Batholith ecoregions) (428 streams), Rockies (Middle Rockies, Southern Rockies ecoregions) (199 streams), and Basin and Plains (Columbia Plateau, Snake River Basin, Northern Basin and Range ecoregions) (262 streams). We excluded rare taxa and taxa identified at coarse taxonomic levels, including Chironomidae. This reduced the 685 taxa from all data sets to 206. Of these 93 exhibited some sensitivity to fine sediment which we classified into four categories: extremely, very, moderately, and slightly sensitive; containing 11, 22, 30, and 30 taxa, respectively. Categories were weighted and a FSBI score calculated by summing the sensitive taxa found in a stream. There were no orders or families that were solely sensitive or resistant to fine sediment. Although, among the three orders commonly regarded as indicators of high water quality, the Plecoptera (5), Trichoptera (3), and Ephemeroptera (2) contained all but one of the species or species groups classified as extremely sensitive. Index validation with an independent data set of 255 streams found FSBI scores to accurately predict both high and low levels of measured fine sediment.     

Surface water and groundwater nitrogen dynamics in a well drained riparian forest within a poorly drained agricultural landscape

The effectiveness of riparian zones in mitigating nutrient in ground and surface water depends on the climate, management, and hydrogeomorphology of a site. The purpose of this study was to determine the efficacy of a well drained, mixed-deciduous riparian forest to buffer a river from N originating from a poorly drained grass seed cropping system. The study site was adjacent to the Calapooia River in the Willamette Valley, Oregon. Water was found to move from the rapid drainage of swale surface water. During winter hydrological events, the riparian forest also received river water. Low nitrate (NO3-) concentrations (0.2-0.4 mg NO3- -NL(-1)) in the shallow groundwater of the cropping system were associated with low rates of mineralization and nitrification (33 kg N ha(-1) yr(-1)) and high grass seed crop uptake of N (155 kg N ha(-1) yr(-1)). The riparian forest soil had higher rates of mineralization (117 kg N ha(-1) yr(-1)) that produced quantities of soil N that were within the range of literature values for plant uptake, leading to relatively low concentrations of shallow groundwater NO3 (0.6-1.8 mg NO3- -NL(-1)). The swale that dissected the cropping system and riparian area was found to have the highest rates of denitrification and to contribute dissolved organic C to the river. Given the dynamic nature of the hydrology of the Calapooia River study site, data suggest that the riparian forest plays a role not only in reducing export of NO3- from the cropping system to the river but also in processing nutrients from river water.     

Coliforms removal in full-scale activated sludge plants in India

This paper investigates the removal of coliforms in full-scale activated sludge plants (ASP) operating in northern regions of India. Log2.2 and log2.4 removal were observed for total coliforms (TC) and fecal coliforms (FC), respectively. However, the effluent still contained a significant number of TC and FC which was greater than the permissible limit for unrestricted irrigation as prescribed by WHO. The observations also suggest that extended aeration (EA) process operating under high mixed liquor suspended solids (MLSS) and sludge retention time (SRT) is more efficient in the removal of coliforms. Further attempts have been made to establish the relationship between two key wastewater parameters, i.e. biochemical oxygen demand (BOD) and suspended solids (SS) with respect to fecal and TC. The relationships were observed to be linear with a good coefficient of correlation. The interrelationship of BOD and SS with coliforms manifest that improvement of the microbiological quality of wastewater could be linked with the removal of SS. Therefore, SS can serve as a regulatory tool in lieu of an explicit coliforms standard.     

AN ANALYSIS OF COLIFORM BACTERIA IN THE UPPER ILLINOIS WATERWAY1

ABSTRACT: During the summer of 1971 about 150 water samples were examined for total and fecal coliform bacteria in the Upper Illinois Waterway at 19 river stations. The data per station were found to be normal geometric distributed. Bacteria densities changed with sampling dates and generally decreased with water movement downstream. Several sewage treatment effluents made marked pulses along the bacterial die-off curves. The observed fecal coliform results were evaluated in terms of the Illinois Pollution Control Board's standards. The FC:TC ratio on the waterway for each station were presented. Using Chick's Law, coliforms death rates were estimated. Efforts to correlate observed total and fecal densities with temperature, flow, algal densities, dissolved oxygen, and 5-day biochemical oxygen demand were not successful. (KEY TERMS: algae; biochemical oxygen demand; coliform bacteria; dissolved oxygen; flow; stream survey; temperature; water pollution; water quality standards)     

AUTUMN CHEMISTRY OF OREGON COAST RANGE STREAMS1

ABSTRACT: During an autumn runoff event we sampled 48 streams with predominantly forested watersheds and igneous bedrock in the Oregon Coast Range. The streams had acid neutralizing capacities (ANC) > 90 μeq/L and pH > 6.4. Streamwater Na ⁺, Ca^{2 +}, and Mg^{2 +} concentrations were greater than K ⁺ concentrations. Anion concentrations generally followed the order of Cl^- > NO₃^- > SO₄^2-. Chloride and Na ⁺ concentrations were highest in samples collected in streams near the Pacific Ocean and decreased markedly as distance from the coast increased. Sea salt exerted no discernible influence on stream water acid-base status during the sampling period. Nitrate concentrations in the study streams were remarkably variable, ranging from below detection to 172 μeq/L. We hypothesize that forest vegetation is the primary control of spatial variability of the NO₃^- concentrations in Oregon Coast Range streams. We believe that symbiotic N fixation by red alder in pure or mixed stands is the primary source of N to forested watersheds in the Oregon Coast Range.     

Linking Landscape Characteristics and High Stream Nitrogen in the Oregon Coast Range: Red Alder Complicates Use of Nutrient Criteria

Red alder (Alnus rubra), a nitrogen(N)‐fixing deciduous broadleaf tree, can strongly influence N concentrations in western Oregon and Washington. We compiled a database of stream N and GIS‐derived landscape characteristics in order to examine geographic variation in N across the Oregon Coast Range. Basal area of alder, expressed as a percent of watershed area, accounted for 37% and 38% of the variation in summer nitrate and total N (TN) concentrations, respectively. Relationships between alder and nitrate were strongest in winter when streamflow and landscape connections are highest. Distance to the coast and latitude, potential surrogates for sea salt inputs, and watershed area were also related to nitrate concentrations in an all‐subsets regression analysis, which accounted for 46% of the variation in summer nitrate concentrations. The model with the lowest Akaike's Information Criterion did not include developed or agricultural land cover, probably because few watersheds in our database had substantial levels of these land cover classes. Our results provide evidence, at a regional scale, that background sources and processes cause many Coast Range streams to exceed proposed nutrient criteria, and that the prevalence of a single tree species (N‐fixing red alder) exerts a dominant control over stream N concentrations across this region.     

Diuron occurrence and distribution in soil and surface and ground water associated with grass seed production

Little is known about the occurrence and distribution of the herbicide diuron [3-(3,4-dichlorophenyl)-1,1-dimethyl urea] in soil, ground water, and surface water in areas affected by grass-seed production. A field study was designed to investigate the occurrence and distribution of diuron and its transformation products at a poorly drained field site located along an intermittent tributary of Lake Creek in the southern Willamette Valley of Oregon. The experimental sites consisted of a field under commercial grass seed production with a cultivated riparian zone and a second site that was part of the same grass seed field but with a noncultivated riparian zone. Diuron and its transformation product DCPMU [3-(3,4-dichlorophenyl)-1-methylurea] were the only significant residues detected in this study. Concentrations of diuron in surface water declined from a maximum of 28 microg/L immediately following application to low levels that persisted as long as flow was present. Diuron and DCPMU concentrations in shallow ground water (15-36 cm below ground surface) were highest (2-13 microg/L) in the zone immediately adjacent (0.5 m) to Lake Creek and indicated the influence of stream water on shallow ground water near the stream. Diuron and DCPMU detected in soil prior to the second season's application indicated the persistence of diuron and DCPMU from the previous year's application. Surface runoff during the rainy season removes only a very small percentage (<1%) of the applied herbicide. In addition, no evidence was obtained for the downward transport of diuron or its transformation products to deep ground water.     

NITRATE CONTAMINATION IN OREGON WELL WATER: GEOLOGIC VARIABILITY AND THE PUBLIC'S PERCEPTION1

ABSTRACT: Approximately 4.5 million people in the United States who rely on well water are exposed to nitrate‐N concentrations exceeding the 10 mg/l standard. In this study in the Southern Willamette Valley in Oregon we reassessed nitrate‐N in rural wells sampled in 2000–2001, compared nitrate‐N concentrations among geological units, and surveyed residents about their perceptions of well water quality. Nitrate‐N concentrations were again sampled in 2002 and found to have increased significantly from the previous period. With rapid population growth in the area, the potential health risk in drinking well water that exceeds 10 mg/l nitrate‐N warrants continued public education. Nitrate‐N concentrations were found to be higher in the Holocene alluvium of the Willamette River and the Pleistocene sand and gravel post‐Missoula Flood deposits. Researchers conducting future studies may choose to stratify and monitor wells by geologic unit and by other parameters that estimate input of nutrients to the environment. Opinions differed between agricultural landowners and nonagricultural landowners with regard to the impact that agricultural fertilizers may have on water quality. Participants were supportive of a range of regulatory actions that might be used by homeowners or landowners to address ground water contamination. Given that the area is now designated a Groundwater Management Area, understanding local stakeholders’perceptions is critical and strategic and has the potential to help public agencies manage potential conflicts of opinion among stakeholders, build consensus, and help guide the approach to restoring ground water quality.     

Relative Importance of Water‐Quality Stressors in Predicting Fish Community Responses in Midwestern Streams

Fish, habitat, and water chemistry data were collected from 98 streams in the midwestern United States, an area dominated by intense cultivation of row crops, in order to identify important water‐quality stressors to fish communities. We focused on 10 stressors including riparian disturbance, riparian vegetative cover, instream fish cover, streambed sedimentation, streamflow variability, total nitrogen, total phosphorus, minimum dissolved oxygen, pesticides, and bed sediment contaminants. Fish community response variables included a measure of observed/expected taxonomic completeness; species‐specific tolerances to nitrogen, phosphorus, dissolved oxygen, and water temperature; the percent of species classified as macrohabitat generalists; and an index of pesticide toxicity to fish. Multivariate analysis indicated that total nitrogen was the most important stressor, signifying that fish communities were responding to total nitrogen despite relatively high levels common to an agricultural setting. Individually, fish taxonomic completeness decreased with increasing streambed sedimentation, whereas fish community tolerance to total phosphorus increased with increasing streambed sedimentation, riparian disturbance, and total nitrogen. These findings underscore the importance of multiple biological response metrics to better understand the effects of water‐quality stressors on fish communities and highlight the complex relations between total phosphorus and fish communities.     

Including Riparian Vegetation in the Definition of Morphologic Reference Conditions for Large Rivers: A Case Study for Europe’s Western Plains

Methods for defining and retrieving reference conditions for large rivers were explored with emphasis on hydromorphologic and biologic quality indicators. For a set of four large rivers in the European Western Plains ecoregion, i.e., the rivers Meuse, Loire, Allier, and Dordogne, reference reaches were selected based on geomorphologic characteristics. A survey of riparian land use, vegetation, and bed geometry was done for the selected reaches. Responses of the riparian landscape to hydromorphologic conditions were determined with a set of existing and newly developed measures of riparian dynamics and forest development. Strong correlations were observed at the reach and local levels between the ratios of width to depth and embankment and the developed measures of riparian dynamics and forest. Boundary conditions for riparian forest development were determined for the hydromorphologic and biologic indicators of riparian dynamics and vegetation structure. These conditions also proved useful for determining the presence of sustainable populations of Populus nigra and Salix purpurea. From this agreement between abiotic and biotic boundary conditions, a set of useful reference conditions was determined, and a framework for the definition of reference and good status conditions subsequently evolved. Finally, a proposal for assessment and monitoring the proposed indicators is discussed for its applicability.     

Bedding and within-pen location effects on feedlot pen runoff quality using a rainfall simulator

Soluble salts, nutrients, and pathogenic bacteria in feedlot-pen runoff have the potential to cause pollution of the environment. A 2-yr study (1998-1999) was conducted at a beef cattle (Bos taurus) feedlot in southern Alberta, Canada, to determine the effect of bedding material [barley (Hordeum vulgare L.) straw versus wood chips] and within-pen location on the chemical and bacterial properties of pen-floor runoff. Runoff was generated with a portable rainfall simulator and analyzed for chemical content (nitrogen [N], phosphorus [P], soluble salts, electrical conductivity [EC], sodium adsorption ratio [SAR], dissolved oxygen [DO], and pH) and populations of three groups of bacteria (Escherichia coli, total coliforms, total aerobic heterotrophs at 27 degrees C) in 1998 and 1999. Bedding had a significant (P < or = 0.05) effect on NH4-N concentration and load in 1999, SO4 load in 1998, SO4 concentration and load in 1999, and total coliforms in both years; where these three variables were higher in wood than straw pens. Location had a significant effect on EC and concentrations of total Kjeldahl nitrogen (TKN), Na, K, SO4, and Cl in 1998, and total coliforms in both years. These seven variables were higher at the bedding pack than pen floor location, indicating that bedding packs were major reservoirs of TKN, soluble salts, and total coliforms. Significantly higher dissolved reactive phosphorus (DRP), total P, and NH4-N concentrations and loads at the bedding pack location in wood pens in 1998, and a similar trend for TKN concentration in 1999, indicated that this bedding-location treatment was a greater source of nutrients to runoff than the other three bedding-location treatments. Bedding, location, and their interaction may therefore be a potential tool to manage nutrients, soluble salts, and bacteria in feedlot runoff.     

FLOW CAPACITY OF CULVERTS ON OREGON COAST RANGE FOREST ROADS1

ABSTRACT: One hundred twenty-eight stream-crossing culverts in the central Oregon Coast Range were evaluated for peak flow capacity and were compared with current design guidelines. Their ability to pass the 25-year peak flow, as mandated by Oregon State Forest Practice Rules, and their maximum flow capacity were determined. Over 40 percent of the culverts were unable to pass the 25-year peak flow at a headwater to diameter ratio of 1. About 17 percent could not pass the 25-year peak flow without headwater overtopping the roadfill. Installing the next larger pipe size at an additional original installation cost of about 14 percent would have allowed nearly all these culverts to pass the 25-year peak flow. Culvert capacity varied with ownership and watershed size.     

Comparison of Hydrology of Wetlands in Pennsylvania and Oregon (USA) as an Indicator of Transferability of Hydrogeomorphic (HGM) Functional Models Between Regions

The hydrogeomorphic (HGM) approach to wetland classification and functional assessment is becoming more widespread in the United States but its use has been limited by the length of time needed to develop appropriate data sets and functional assessment models. One particularly difficult aspect is the transferability among geographic regions of specific models used to assess wetland function. Sharing of models could considerably shorten development and implementation of HGM throughout the United States and elsewhere. As hydrology is the driving force behind wetland functions, we assessed the comparability of hydrologic characteristics of three HGM subclasses (slope, headwater floodplain, mainstem floodplain) using comparable long-term hydrologic data sets from different regions of the United States (Ridge and Valley Province in Pennsylvania and the Willamette Valley in Oregon). If hydrology by HGM subclass were similar between different geographic regions, it might be possible to more readily transfer extant models between those regions. We found that slope wetlands (typically groundwater-driven) had similar hydrologic characteristics, even though absolute details (such as depth of water) differed. We did not find the floodplain subclasses to be comparable, likely due to effects of urbanization in Oregon, regional differences in soils and, perhaps, climate. Slight differences in hydrology can shift wetland functions from those mediated by aerobic processes to those dominated by anaerobic processes. Functions such as nutrient cycling can be noticeably altered as a result. Our data suggest considerable caution in the application of models outside of the region for which they were developed.     

Upland disturbance affects headwater stream nutrients and suspended sediments during baseflow and stormflow

Because catchment characteristics determine sediment and nutrient inputs to streams, upland disturbance can affect stream chemistry. Catchments at the Fort Benning Military Installation (near Columbus, Georgia) experience a range of upland disturbance intensities due to spatial variability in the intensity of military training. We used this disturbance gradient to investigate the effects of upland soil and vegetation disturbance on stream chemistry. During baseflow, mean total suspended sediment (TSS) concentration and mean inorganic suspended sediment (ISS) concentration increased with catchment disturbance intensity (TSS: R2= 0.7, p = 0.005, range = 4.0-10.1 mg L(-1); ISS: R2= 0.71, p = 0.004, range = 2.04-7.3 mg L(-1)); dissolved organic carbon (DOC) concentration (R2= 0.79, p = 0.001, range = 1.5-4.1 mg L(-1)) and soluble reactive phosphorus (SRP) concentration (R2= 0.75, p = 0.008, range = 1.9-6.2 microg L(-1)) decreased with increasing disturbance intensity; and ammonia (NH4+), nitrate (NO3-), and dissolved inorganic nitrogen (DIN) concentrations were unrelated to disturbance intensity. The increase in TSS and ISS during storms was positively correlated with disturbance (R2= 0.78 and 0.78, p = 0.01 and 0.01, respectively); mean maximum change in SRP during storms increased with disturbance (r = 0.7, p = 0.04); and mean maximum change in NO3- during storms was marginally correlated with disturbance (r = 0.58, p = 0.06). Soil characteristics were significant predictors of baseflow DOC, SRP, and Ca2+, but were not correlated with suspended sediment fractions, any nitrogen species, or pH. Despite the largely intact riparian zones of these headwater streams, upland soil and vegetation disturbances had clear effects on stream chemistry during baseflow and stormflow conditions.