ESTIMATION OF SOIL MOISTURE FROM SEASAT SAR DATA1

ABSTRACT A study was made to determine if Seasat Synthetic Aperture Radar (SAR) data could be used to make practical estimates of soil moisture. Extensive ground measurements were collected at two primary sites near Guymon, Oklahoma, and Sublette, Kansas. The relative sensitivity of the SAR to differences in soil moisture, tillage roughness, and vegetation was determined. To validate the effects detected in the SAR data, an airborne scatterometer with a similar wavelength was flown repeatedly over the Guymon site. Soil moisture variations in the surface 2 cm and surface 15 cm of fields with bare soil, milo and alfalfa produce similar responses in the scattering coefficient from both systems. Roughness due to tillage in row crops produced as much as 12–15 dB increase in the scatterometer return. Most agricultural vegetation was effectively penetrated by the L-band frequencies; however, corn produced an exceptionally high radar return either standing or after combine harvesting. When corn had ripened, there was some evidence that tillage roughness could be detected through the canopy. Moderate tillage roughness produced by grain drill furrows caused over 12 dB increase in return when row directions changed from parallel to perpendicular with respect to the SAR look direction. Dramatic increases in return occurred when vegetation surfaces were wet. Increased radar returns from tillage roughness, some vegetation and wet vegetation surfaces, all dyanmic in nature, were significant and may limit the practical estimation of soil moisture from the radar data.     

Application of remote and in situ information to the management of wetlands in Poland

The protection and regeneration of wetlands has been of crucial importance as a goal in ecological research and in nature conservation for some time and is more important than ever now. Knowledge about the biophysical properties of wetlands' vegetation retrieved from satellite images enables us to improve the monitoring of these unique areas, which are otherwise very often impenetrable and therefore difficult to examine, analyze and assess by means of site visits. The Biebrza Wetlands are situated in the North-East part of Poland and are one of the largest areas made up of marshes and swamps in the entire EU. This is still one of the wildest areas and one of the least destroyed, damaged or changed by human impact. However, in the recent decades there have been attempts made to intensify and overexploit the natural resources of the region and implement new agriculture practices in the area. In this period, drainage canals have been built, and a good deal of the area has been drained. The area of this precious ecosystem covers 25 494 ha. This valuable area of peat with unique vegetation species and with very special birds is one of the most valuable areas in Europe and in 1995 was added to the list of Ramsar sites. The investigation of wetlands in the Biebrza River Valley has been carried out at ground level by taking measurements of soil moisture, evapotranspiration, Leaf Area Index, wet and dry biomass and the levels of ground water and meteorological parameters. Also examined were radiative temperature, detailed vegetation mapping, and APAR. For some years the deterioration of peat lands has been noticed due to the drying out of the area and the frequent outbreak of fires. The consequence is the succession of new vegetation and the appearance of new ecosystems. The Remote Sensing Centre in the Institute of Geodesy and Cartography has undertaken the investigation by applying ERS-2.SAR and ENVISAT ASAR of IS2 and IS4 and VV, HH, HV polarization for the purpose of modeling soil moisture and humidity changes of the area under investigation. The investigation also aimed at finding the best biophysical properties of wetlands' vegetation to characterize marshland habitats and its changes. At the same time as registering the microwave data, the optical data from Landsat ETM+, SPOT VEGETATION, ERS-2.ATSR, ENVISAT MERIS, and NOAA/AVHRR have been registered and information about the biomass and heat fluxes as sensible and latent heat has also been calculated. The vegetation indices are calculated from EO satellite data taking into account jointly the features of vegetation responsible for reflection in various bands and combining this information from several spectral bands. Also, the changes in the humidity of the area have been examined by extracting the backscattering coefficients from two SAR images that were taken at a similar period of the year but with a gap of 5 years. The information about soil moisture as retention, soil moisture changes, heat fluxes and evapotranspiration are all very important for estimates of CO(2) sequestration. The ENVISAT images have been obtained for the ESA AO-ID122 project. Also the SMOS and ALOS data will be applied for the Biebrza Wetlands in the future.     

Temporal and spatial development of surface soil conditions at two created riverine marshes

The amount of time it takes for created wetlands to develop soils comparable to natural wetlands is relatively unknown. Surface soil changes over time were evaluated in two created wetlands (approximately 1 ha each) at the Olentangy River Wetland Research Park in Columbus, Ohio. The two wetlands were constructed in 1993 to be identical in size and geomorphology, and maintained to have the same hydrology. The only initial difference between the wetlands was that one was planted with native macrophytes while the other was not. In May 2004, soil samples were collected (10 yr and 2 mo after the wetlands were flooded) and compared to samples collected in 1993 (after the wetlands were excavated but before flooding) and 1995 (18 mo after the wetlands were flooded). In all three years, soils were split into surface (0-8 cm) and subsurface (8-16 cm) depths and analyzed for soil organic matter, total C, total P, available P, exchangeable cations, and pH. Soils in the two wetlands have changed substantially through sedimentation and organic accretion. Between 1993 and 1995, soils were most influenced by the deposition of senescent macroalgae, the mobilization of soluble nutrients, and the precipitation of CaCO(3). Between 1995 and 2004, soil parameters were influenced more by the deposition of organic matter from colonized macrophyte communities. Mean percent organic matter at the surface increased from 5.3 +/- 0.1% in 1993, 6.1 +/- 0.2% in 1995, to 9.5 +/- 0.2% in 2004. Mean total P increased from 493 +/- 18 microg g(-1) in 1993, 600 +/- 23 microg g(-1) in 1995, to 724 +/- 20 microg g(-1) in 2004. Spatial analyses of percent organic matter (a commonly used indicator of hydric soil condition) at both wetlands in 1993, 1995, and 2004 showed that soil conditions have become increasingly more variable. High spatial structure (autocorrelation) between data points was detected in 1993 and 2004, with data in 2004 exhibiting a much higher overall variance and narrower range of spatial structure than in 1993.     

COMBINATION OF RADAR AND GAUGE DATA IN A RAINFALL ARCHIVE SYSTEM1

ABSTRACT: Near real time daily rainfall estimates for the UK are available from three sources: a sparse network of gauges, radar data, or radar data adjusted by the sparse gauges. The PARAGON rainfall archive system, which has been developed by the UK Meteorological Office, is able to produce these estimates in near real time on a 5 km grid. The ability of these estimates to reproduce the 5 km grid point field derived later from a dense network of gauges is compared using case studies. Five techniques have been used to assess the relative quality of the various estimates. There is general agreement between the results of the various techniques. For the London radar there are examples of days when the rainfall estimate was improved by incorporating radar data; conversely, there are days when the radar data make it worse. Overall little evidence was found to suggest that adjusted radar data are consistently markedly better than gauge estimates. Discriminate use of radar data is recommended.     

Selecting Sites for Converting Farmlands to Wetlands in the Sanjiang Plain, Northeast China, Based on Remote Sensing and GIS

Wetlands in the Sanjiang Plain are rich in biodiversity and natural resources in the northeast of China. However, this wetland area has decreased in size and deteriorated in quality owing to expanded agricultural activities since the 1950s. Converting farmlands to wetlands is necessary to improve these conditions. Using Remote Sensing (RS) and Geographic Information Systems (GIS) technologies, we derived farmland productivity data and hydrology data for the Sanjiang Plain. The farmland productivity data were derived from land use and net primary productivity (NPP) data of the MODIS products. We obtained three productivity farmland classes (low, medium, and high) through the NPP anomaly percentage method. We were only concerned with the low-productivity farmland. Hydrology data were modeled with a wetness index, which was derived from Digital Elevation Model (DEM) data. Based on these two data layers, we identified and prioritized sites for the conversion of farmlands to wetlands. The areas with low farmland productivity and medium or high wetness values have potential to support the conversion of farmlands to wetlands. Potential sites were prioritized in terms of patch size and proximity to natural wetlands and water bodies. We obtained three priority classes, among which the high-priority class would be used as the areas for the recent conversion of farmlands to wetlands. The area of this class was 75,888 ha and accounted for 1.3% of the total farmland area.     

Effect of agricultural extension on cotton farmer's efficiency in arid and semi-arid areas of Burkina Faso

This study shows that farmers in arid and semi-arid land can improve their efficiency if they have access to agricultural extension services. An exhaustive survey was used to obtain data from 423 cotton producers in the arid zones and 144 of wetlands in Burkina Faso. The method of the directional distance function modeling a multi-output technology was used. The results show that cotton farmers in the arid zones are not technically more inefficient than those in wetlands. On average, arid zone producers have a technical inefficiency of 0.056 while that of wetland producers is around 0.052. The agricultural extension, defined as receiving visits and advice from technical agents, is important for producers because it helps farmers choose soil and water conservation techniques and fertilizers adapted to the Agro-climatic conditions of their respective zones. Agriculture extension also helps farmers of the arid zones to reduce their technical inefficiency by advising them to cultivate small agricultural plots. On average, in these zones, the size of an agricultural plot is around 1.03 ha compared to those of wetlands which is about 1.6 ha.     

Denitrification in alluvial wetlands in an urban landscape

Riparian wetlands have been shown to be effective "sinks" for nitrate N (NO3-), minimizing the downstream export of N to streams and coastal water bodies. However, the vast majority of riparian denitrification research has been in agricultural and forested watersheds, with relatively little work on riparian wetland function in urban watersheds. We investigated the variation and magnitude of denitrification in three constructed and two relict oxbow urban wetlands, and in two forested reference wetlands in the Baltimore metropolitan area. Denitrification rates in wetland sediments were measured with a 15N-enriched NO3- "push-pull" groundwater tracer method during the summer and winter of 2008. Mean denitrification rates did not differ among the wetland types and ranged from 147 +/- 29 microg N kg soil(-1) d(-1) in constructed stormwater wetlands to 100 +/- 11 microg N kg soil(-1) d(-1) in relict oxbows to 106 +/- 32 microg N kg soil(-1) d(-1) in forested reference wetlands. High denitrification rates were observed in both summer and winter, suggesting that these wetlands are sinks for NO3- year round. Comparison of denitrification rates with NO3- standing stocks in the wetland water column and stream NO3- loads indicated that mass removal of NO3- in urban wetland sediments by denitrification could be substantial. Our results suggest that urban wetlands have the potential to reduce NO3- in urban landscapes and should be considered as a means to manage N in urban watersheds.     

A conceptual framework for assessing cumulative impacts on the hydrology of nontidal wetlands

Wetlands occur in geologic and hydrologic settings that enhance the accumulation or retention of water. Regional slope, local relief, and permeability of the land surface are major controls on the formation of wetlands by surface-water sources. However, these landscape features also have significant control over groundwater flow systems, which commonly play a role in the formation of wetlands. Because the hydrologic system is a continuum, any modification of one component will have an effect on contiguous components. Disturbances commonly affecting the hydrologic system as it relates to wetlands include weather modification, alteration of plant communities, storage of surface water, road construction, drainage of surface water and soil water, alteration of groundwater recharge and discharge areas, and pumping of groundwater. Assessments of the cumulative effects of one or more of these disturbances on the hydrologic system as related to wetlands must take into account uncertainty in the measurements and in the assumptions that are made in hydrologic studies. For example, it may be appropriate to assume that regional groundwater flow systems are recharged in uplands and discharged in lowlands. However, a similar assumption commonly does not apply on a local scale, because of the spatial and temporal dynamics of groundwater recharge. Lack of appreciation of such hydrologic factors can lead to misunderstanding of the hydrologic function of wetlands within various parts of the landscape and mismanagement of wetland ecosystems.     

FORESTED WETLANDS IN EASTERN CONNECTICUT: THEIR TRANSITION ZONES AND DELINEATION1

ABSTRACT: The delineation of inland wetlands requires close field examination of the biological and physical gradients (transition zones) between wetlands and bordering uplands. As part of a study on the detection and delineation of inland wetlands in eastern Connecticut by remote sensing techniques, this effort was designed to investigate vegetation distribution and composition and selected physical and chemical properties of the soils of wetland to upland transition zones in deciduous wetland forests. Field research was conducted during the growing season of 1975 within a test area consisting of the 45 mi² Town of Mansfield, Connecticut. Changes in vegetation composition and structure, soil pH, and soil water content were determined along line transects extended over wetland to upland transition zones. Differences in soil pH occurred along the transects but were of such magnitude that they probably have little impact on plant distribution. There were significant changes in soil water content along the wetland to upland gradients. Discriminant analysis applied to statistical “index of abundance” data describing vegetation distribution among the various zones (wetland, transition, upland) showed which plant species best distinguish wetlands from uplands. Of the criteria studied, vegetation composition and distribution, soil water content, and relief are the most useful criteria for delineating deciduous wetland forests.     

A spatially explicit investigation of phosphorus sorption and related soil properties in two riparian wetlands

Soils of riparian wetlands are highly effective at phosphorus (P) sorption. However, these soils exhibit extreme spatial variability across riparian zones. We used a spatially explicit sampling design in two riparian wetlands in North Carolina to better understand the relationships among P sorption, soil properties, and spatial variability. Our objectives were to quantify patterns of spatial variability of P sorption and related soil properties, and to determine which soil properties best explained the variability in P sorption after accounting for the effects of spatial autocorrelation. We measured bulk density, moisture, pH, soil organic matter (SOM), texture (percent clay, silt, and sand), oxalate-extractable aluminum (Al(ox)), iron (Fe(ox)), and the phosphorus sorption index (PSI). Due to differences in texture, Al(ox), and Fe(ox), the two sites had substantially different mean PSIs. At each site, we found considerable differences in the spatial variability of soil properties. For example, semivariance analysis and kriging illustrated that soil properties at Site 1 varied at smaller scales than those at Site 2. At both sites, after accounting for the effects of spatial autocorrelation and all other soil properties, we determined that Al(ox) had the highest Mantel correlation with PSI. We believe this geostatistic and Mantel approach is robust and could serve as a model for research on other biogeochemical processes such as denitrification.     

A Method for Improving the Management of Controversial Wetland

Valley bottom wetlands in agricultural landscapes often are neglected in national and regional wetland inventories. Although these areas are small, located in the bottomlands of the headwater catchments, and scattered in the rural landscape, they strongly influence hydrology, water quality, and biodiversity over the whole catchment area. Valley bottom wetlands often are considered as controversial wetlands. Awareness of the functional role of wetlands is increasing, in parallel with their progressive disappearance in intensive farming landscapes. The need to improve tools for controlling wetland management is a primary consideration for decision makers and land users. This article proposes a method for the inventory of valley bottom wetlands. The method is based on the functional analysis of potential, existing, and efficient valley bottom wetlands (the PEEW approach). Several indicators are proposed for checking the validity of such an approach. Potential wetlands are delineated by means of a topographic index using topographic and pedoclimatic criteria computed from a Digital Elevation Model and easily accessible databases. Existing wetlands are identified from observed surface moisture, the presence of specific wetland vegetation, or soil feature criteria. Efficient wetlands are defined through a given function, such as flow or pollutant regulation or biodiversity control. An analysis of areas at the limits between potential, existing, and efficient wetlands highlights land cultivated or drained in the past, which currently represents negotiating areas in which rehabilitation and other intended management actions can be implemented.     

Effects of prolonged irrigation with treated municipal effluent on runoff rate

The use of domestic effluents for the irrigation of crops has been widespread in Israel for the past 30 years. The sodium adsorption ratio (SAR) of the standardized domestic effluents ranges between 4 and 6. According to the literature, when soils with SAR levels of 4 to 6 are exposed to direct raindrop impact they are subjected to enhanced aggregate disintegration, leading to sealing processes of the soil surface and subsequent increased runoff and soil erosion. However, these phenomena were not observed in the laboratory and field experiments of this study. On the other hand, a rapid decrease of the soil SAR to its initial values was observed, in laboratory and fieldwork, once the soil was subjected to a simulated rainstorm of distilled water (laboratory) or natural rainstorms (field plots). We can conclude that the process of SAR increase during irrigation with standardized effluent water is reversible. Further investigation in this direction can lead to recommendations regarding the necessary levels of domestic sewage water purification in correlation with soil types, climatic conditions, and hazards to tap water aquifers.     

Use of Synthetic Aperture Radar for Selecting Alaskan Lakes for Winter Water Use1

Abstract: Water resources are limited in many areas of the North Slope, Alaska, particularly during winter. Water is used by the oil industry for ice road construction and maintenance, drilling and facility operations, and potable water supplies. The coastal plain between Teshekpuk Lake, in the National Petroleum Reserve‐Alaska (NPR‐A) and the Colville River has numerous shallow lakes, but further south in the northern foothills of the Brooks Range, and east to the Canning River, lakes are fewer. While many oil and gas lease sales have been conducted, or are proposed, access to the leases may be limited because of the lack of available water for ice road construction. Ice roads are the main means by which exploration is conducted in the Arctic, putting a stress on freshwater bodies that do not freeze to the lakebed in winter. Lakes that do not freeze to the lakebed also serve as overwintering habitat for fish. The purpose of this paper is to report on the potential distribution of water bodies that may provide overwinter water in selected areas from Teshekpuk Lake to the Canning River. The project used synthetic aperture radar (SAR) imagery to search for the presence of water in lakes in March 2006. In the Kuparuk and Canning SAR images, 52 and 61% of lakes were frozen to their beds by March 2006, accounting for 49 and 57% of the lake area in these study regions. Conversely, only 2% of the lakes in the Teshekpuk region were frozen to the bottom by March 2006. Unfrozen water was more available because of deeper and more numerous lakes in the Teshekpuk Lake region (west) than in the Canning River area (east). While only specific SAR tiles were analyzed herein, the method will be a useful tool for land managers who seek to evaluate the potential for ice road construction across the Arctic.     

Soil biogeochemical characteristics influenced by alum application in a municipal wastewater treatment wetland

Constructed treatment wetlands are a relatively low-cost alternative used for tertiary treatment of wastewater. Phosphorus (P) removal capacity of these wetlands may decline, however, as P is released from the accrued organic soils. Little research has been done on methods to restore the treatment capacity of aging constructed wetlands. One possibility is the seasonal addition of alum during periods of low productivity and nutrient removal. Our 3-mo mesocosm study investigated the effectiveness of alum in immobilizing P during periods of reduced treatment efficiency, as well as the effects on soil biogeochemistry. Eighteen mesocosms were established, triplicate experimental and control units for Typha sp., Schoenoplectus californicus, and submerged aquatic vegetation (SAV) (Najas guadalupensis dominated). Alum was slowly dripped to the water column of the experimental units at a rate of 0.91 g Al m(-2) d(-1) and water quality parameters were monitored. Soil cores were collected at experiment initiation and completion and sectioned into 0- to 5- and 5- to 10-cm intervals for characterization. The alum floc remained in the 0- to 5-cm surface soil, however, soil pH and microbial parameters were impacted throughout the upper 10 cm with the lowest pH found in the Typha treatment. Plant type did not impact most biogeochemical parameters; however, data were more variable in the SAV mesocosms. Amorphous Al was greater in the surface soil of alum-treated mesocosms, inversely correlated with soil pH and microbial biomass P in both soil layers. Microbial activity was also suppressed in the surface soil of alum-treated mesocosms. This research suggests alum may significantly affect the biogeochemistry of treatment wetlands and needs further investigation.     

Evaluating stream water quality through land use analysis in two grassland catchments: impact of wetlands on stream nitrogen concentration

We evaluated the impacts of natural wetlands and various land uses on stream nitrogen concentration in two grassland-dominated catchments in eastern Hokkaido, Japan. Analyzing land use types in drainage basins, measuring denitrification potential of its soil, and water sampling in all seasons of 2003 were performed. Results showed a highly significant positive correlation between the concentration of stream NO3-N and the proportion of upland area in drainage basins in both catchments. The regression slope, which we assumed to reflect the impact on water quality, was 24% lower for the Akkeshi catchment (0.012 +/- 0.001) than for the Shibetsu catchment (0.016 +/- 0.001). In the Akkeshi catchment, there was a significant negative correlation between the proportion of wetlands in the drainage basins and stream NO3-N concentration. Stream dissolved organic nitrogen (DON) and carbon (DOC) concentrations were significantly higher in the Akkeshi catchment. Upland and urban land uses were strongly linked to increases in in-stream N concentrations in both catchments, whereas wetlands and forests tended to mitigate water quality degradation. The denitrification potential of the soils was highest in wetlands, medium in riparian forests, and lowest in grasslands; and was significant in wetlands and riparian forests in the Akkeshi catchment. The solubility of soil organic carbon (SOC) and soil moisture tended to determine the denitrification potential. These results indicate that the water environment within the catchments, which influences denitrification potential and soil organic matter content, could have caused the difference in stream water quality between the two catchments.     

Global monitoring of wetlands--the value of ENVISAT ASAR Global mode

This paper elaborates on recent advances in the use of ScanSAR technologies for wetland-related research. Applications of active satellite radar systems include the monitoring of inundation dynamics as well as time series analyses of surface soil wetness. For management purposes many wetlands, especially those in dry regions, need to be monitored for short and long-term changes. Another application of these technologies is monitoring the impact of climate change in permafrost transition zones where peatlands form one of the major land cover types. Therefore, examples from boreal and subtropical environments are presented using the analysed ENVISAT ASAR Global mode (GM, 1 km resolution) data acquired in 2005 and 2006. In the case of the ENVISAT ASAR instrument, data availability of the rather coarse Global Mode depends on request priorities of other competing modes, but acquisition frequency may still be on average fortnightly to monthly depending on latitude. Peatland types covering varying permafrost regimes of the West Siberian Lowlands can be distinguished from each other and other land cover by multi-temporal analyses. Up to 75% of oligotrophic bogs can be identified in the seasonal permafrost zone in both years. The high seasonal and inter-annual dynamics of the subtropic Okavango Delta can also be captured by GM time series. Response to increased precipitation in 2006 differs from flood propagation patterns. In addition, relative soil moisture maps may provide a valuable data source in order to account for external hydrological factors of such complex wetland ecosystems.     

Modeling the Effects of Tile Drain Placement on the Hydrologic Function of Farmed Prairie Wetlands

The early 2000s saw large increases in agricultural tile drainage in the eastern Dakotas of North America. Agricultural practices that drain wetlands directly are sometimes limited by wetland protection programs. Little is known about the impacts of tile drainage beyond the delineated boundaries of wetlands in upland catchments that may be in agricultural production. A series of experiments were conducted using the well‐published model WETLANDSCAPE that revealed the potential for wetlands to have significantly shortened surface water inundation periods and lower mean depths when tile is placed in certain locations beyond the wetland boundary. Under the soil conditions found in agricultural areas of South Dakota in North America, wetland hydroperiod was found to be more sensitive to the depth that drain tile is installed relative to the bottom of the wetland basin than to distance‐based setbacks. Because tile drainage can change the hydrologic conditions of wetlands, even when deployed in upland catchments, tile drainage plans should be evaluated more closely for the potential impacts they might have on the ecological services that these wetlands currently provide. Future research should investigate further how drainage impacts are affected by climate variability and change.     

Capturing LiDAR‐Derived Hydrologic Spatial Parameters to Evaluate Playa Wetlands

The digital elevation model data from traditional stereo photogrammetric methods are inadequate in providing accurate vertical parameters to feed hydrologic models for low‐lying, extremely flat areas. High‐resolution light detection and ranging (LiDAR) data provide the robust capability of capturing small variations in low‐relief playa wetlands. The Rainwater Basin in south‐central Nebraska includes a complex of seasonally shallow playa wetlands that attract millions of migratory waterfowl every spring and fall. This research focuses on the development of a procedure with applicable protocols to produce LiDAR‐derived three‐dimensional wetland maps and to extract the critical surface parameters (i.e., watershed boundaries, flow direction, flow accumulation, and drainage lines) for playa wetlands. The topo‐hydrologic conditions of playa wetlands were evaluated at the watershed level. The results show that in the Rainwater Basin, 70.7% of the historic hydric soil footprints identified in the Soil Survey Geographic (SSURGO) database were not functioning as topographically depressional wetlands. This finding was confirmed by a recent five‐year Annual Habit Survey showing that 69.8% of the historic hydric soil footprints did not function during the spring migratory bird seasons between 2004 and 2009. The majority of playa wetlands' topographic conditions have been substantially changed and the SSURGO data cannot fully reflect current topographic reality in the Rainwater Basin.     

Testing the Basic Assumption of the Hydrogeomorphic Approach to AssessingWetland Functions

The hydrogeomorphic (HGM) approach for developing "rapid" wetland function assessment methods stipulates that the variables used are to be scaled based on data collected at sites judged to be the best at performing the wetland functions (reference standard sites). A critical step in the process is to choose the least altered wetlands in a hydrogeomorphic subclass to use as a reference standard against which other wetlands are compared. The basic assumption made in this approach is that wetlands judged to have had the least human impact have the highest level of sustainable performance for all functions. The levels at which functions are performed in these least altered wetlands are assumed to be "characteristic" for the subclass and "sustainable." Results from data collected in wetlands in the lowlands of western Washington suggest that the assumption may not be appropriate for this region. Teams developing methods for assessing wetland functions did not find that the least altered wetlands in a subclass had a range of performance levels that could be identified as "characteristic" or "sustainable." Forty-four wetlands in four hydrogeomorphic subclasses (two depressional subclasses and two riverine subclasses) were rated by teams of experts on the severity of their human alterations and on the level of performance of 15 wetland functions. An ordinal scale of 1-5 was used to quantify alterations in water regime, soils, vegetation, buffers, and contributing basin. Performance of functions was judged on an ordinal scale of 1-7. Relatively unaltered wetlands were judged to perform individual functions at levels that spanned all of the seven possible ratings in all four subclasses. The basic assumption of the HGM approach, that the least altered wetlands represent "characteristic" and "sustainable" levels of functioning that are different from those found in altered wetlands, was not confirmed. Although the intent of the HGM approach is to use level of functioning as a metric to assess the ecological integrity or "health" of the wetland ecosystem, the metric does not seem to work in western Washington for that purpose.     

Variation in Assemblages of Small Fishes and Microcrustaceans After Inundation of Rarely Flooded Wetlands of the Lower Okavango Delta, Botswana

Water extraction from floodplain river systems may alter patterns of inundation of adjacent wetlands and lead to loss of aquatic biodiversity. Water reaching the Okavango Delta (Delta), Botswana, may decrease due to excessive water extraction and climate change. However, due to poor understanding of the link between inundation of wetlands and biological responses, it is difficult to assess the impacts of these future water developments on aquatic biota. Large floods from 2009 to 2011 inundated both rarely and frequently flooded wetlands in the Delta, creating an opportunity to examine the ecological significance of flooding of wetlands with widely differing hydrological characteristics. We studied the assemblages of small fishes and microcrustaceans, together with their trophic relationships, in temporary wetlands of the lower Delta. Densities of microcrustaceans in temporary wetlands were generally lower than previously recorded in these habitats. Microcrustacean density varied with wetland types and hydrological phase of inundation. High densities of microcrustaceans were recorded in the 2009 to 2010 flooding season after inundation of rarely flooded sites. Large numbers of small fishes were observed during this study. Community structure of small fishes differed significantly across the studied wetlands, with poeciliids predominant in frequently flooded wetlands and juvenile cichlids most abundant in rarely flooded wetlands (analysis of similarity, P < 0.05). Small fishes of <20 mm fed largely on microcrustaceans and may have led to low microcrustacean densities within the wetlands. This result matched our prediction that rarely flooded wetlands would be more productive; hence, they supported greater populations of microcrustaceans and cichlids, which are aggressive feeders. However, the predominance of microcrustaceans in the guts of small fishes (<20 mm) suggests that predation by fishes may also be an important regulatory mechanism of microcrustacean assemblages during large floods when inundated terrestrial patches of wetlands are highly accessible by fish. We predict that a decline in the amount of water reaching the Delta will negatively affect fish recruitment, particularly the cichlids that heavily exploited the rarely flooded wetlands. Cichlids are an important human food source, and their decline in fish catches will negatively affect livelihoods. Hence, priority in the management of the Delta’s ecological functioning should be centred on minimising natural water-flow modifications because any changes may be detrimental to fish-recruitment processes of the system.